Hörnes 1848 : 16 Conus clavatus Lam. Conus clavatus Lamarck, 1810 Conus ventricosus Bronn—Hörnes 1851: 32 Lautoconus ventricosus ( Gmelin, 1791 ) Auinger 1879 : 48 Conus ventricosus in Hörnes 1851 Auinger 1879 : 51 Conus mediterraneus var. pupoidemiocenica Sacc. Sacco 1893b : 104 Conus mediterraneus Hoernes & Auinger, 1879 Conus ( Chelyconus ) mediterraneus var. mioexclavata Sacc. Conus clavatus Hörnes, 1851 Conus mediterraneus var. permiocenica Sacc. Sacco 1893b : 104 Conus mediterraneus Hoernes & Auinger, 1879 Conus ( Chelyconus ) vindobonensis Partsch—Strausz 1966: 458 Conus ( Chelyconus ) vindobonensis Partsch in Hoernes et Auinger, 1879 Conus ( Chelyconus ) vindobonensis Partsch in Hörnes, 1848 ( 1856 ) Conus mediterraneus Hwass—Krach 1981: 77 Conus vindobonensis vindobonensis (Partsch) Ionesi & Nicorici 1994 : 62 Conus ( Chelyconus ) pyrula Brocchi, 1814 Bałuk 1997 : 63 Lautoconus pyrula ( Brocchi, 1814 ) Conus ( Chelyconus ) vindobonensis Partsch in Hörnes, 1856 Bałuk 1997 : 65 Voia 2001 : 156 Conus ( Chelyconus ) vindobonensis Kojumdgieva & Strachimirov 1960 : 213 Conus mucronatolaevis Sacco, 1893 Chelyconus vindobonensis (Partsch in Hörnes, 1856 ) Kovács & Vicián 2013 : 62 A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea Harzhauser, Mathias Landau, Bernard Zootaxa 2016 4210 1 1 178 9CHYM [151,229,1561,1587] Gastropoda Conidae Conus Animalia Neogastropoda 150 151 Mollusca genus lato   [  Conus]  vindobonensisP.[artsch]—  Hörnes 1848: 16(nomen nudum).    Conus clavatusLam.—Hörnes 1851: 25, pl. 2, figs 4a–c [non  Conus clavatus Lamarck, 1810].   Conus ventricosusBronn—Hörnes 1851: 32(partim), pl. 3, figs 5a–c, 7a–c [non  Lautoconus ventricosus( Gmelin, 1791)].    Conus( Chelyconus) VindobonensisPartsch—Hoernes &  Auinger 1879: 48[nov. nom. pro  Conus ventricosusin Hörnes 1851, pl. 3, figs 5, 7].    Conus( Chelyconus) mediterraneusHwass—Hoernes &  Auinger 1879: 51, pl. 6, figs 9–11.  [  Conus] C.[ helyconus]  mediterraneusvar. pupoidemiocenicaSacc.—  Sacco 1893b: 104[nov. nom. pro  Conus mediterraneus Hoernes & Auinger, 1879, pl. 6, fig. 10]. [  Conus( Chelyconus) mediterraneus] var. mioexclavataSacc.—Sacco 1893: 104 [nov. nom. pro  Conus clavatusHörnes, 1851, pl. 2, fig. 4].  [  Conus] C.[ helyconus]  mediterraneusvar. permiocenicaSacc.—  Sacco 1893b: 104[nov. nom. pro  Conus mediterraneus Hoernes & Auinger, 1879, pl. 6, fig. 11].   Conus( Chelyconus) vindobonensisPartsch—Strausz 1966: 458, pl. 69, figs 3–4.   Conus( Chelyconus) vindobonensisPartsch in Hoernes et Auinger, 1879— Hinculov 1968: 149, pl. 37, fig. 16.  Conus( Chelyconus) vindobonensisPartsch in Hörnes, 1848( 1856)— Nicorici & Sagatovici 1973: 176, pl. 27, figs 2–3.  Conus mediterraneusHwass—Krach 1981: 77, pl. 20, figs 13–16, pl. 21, figs 1–2, 5.  ?  Conus vindobonensis vindobonensis(Partsch)—  Ionesi & Nicorici 1994: 62, pl. 5, figs 11–12.    Conus( Chelyconus) pyrula Brocchi, 1814—  Bałuk 1997: 63, pl. 21, figs 5–6 [non  Lautoconus pyrula( Brocchi, 1814)].  Conus( Chelyconus) vindobonensisPartsch in Hörnes, 1856—  Bałuk 1997: 65, pl. 23, figs 1–6.  ?  Conus( Chelyconus) vindobonensisPartsch in Hörnes—Chira &  Voia 2001: 156, pl. 2, figs 5a–b.  non  Conus( Chelyconus) vindobonensis(Partsch in Hoernes und Auinger 1879)—Kojumdgieva in  Kojumdgieva & Strachimirov 1960: 213, pl. 50, figs 4–5 [=  Conus s.l.  mucronatolaevisSacco, 1893].  non  Chelyconus vindobonensis(Partsch in Hörnes, 1856)—  Kovács & Vicián 2013: 62, figs 28–29.     Typematerial. Syntypes: 3 spec.  NHMW1846/0037/0043, Gainfarn( Austria), including specimen illustrated in Hörnes(1851, pl. 3, fig. 5); 5 spec.   NHMW1846/0037/0049, Enzesfeld( Austria), including specimen illustrated in Hörnes(1851, pl. 3, fig. 7).   Studied material.3 spec.  NHMW1846/0037/0043, Gainfarn( Austria), including specimen illustrated in Hörnes(1851, pl. 3, fig. 5); 5 spec.   NHMW1846/0037/0049, Enzesfeld( Austria), including specimen illustrated in Hörnes(1851, pl. 3, fig. 7); 10 spec. NHMW 1855/0045/0359, 9 spec. NHMW 1853/0003/0005, 9 spec.   NHMW1856/0050/0115, all Gainfarn( Austria); 4 spec.   NHMW1846/0037/0052, Gainfarnand Steinebrunn( Austria); 7 spec.   NHMW1846/00027/0028, Steinebrunn( Austria); 16 spec.   NHMW1860/0001/0067, Mikulov-Kienberk( Czech Republic); 10 spec.   NHMW1849/0023/0002, Bad Vöslau, including specimen illustrated in  Hoernes& Auinger(1879, pl. 6, fig. 9); 1 spec.   NHMW1849/0023/0002a, Bad Vöslau, specimen illustrated in  Hoernes& Auinger(1879, pl. 6, fig. 10); 3 spec.   NHMW1846/0037/0048, Enzesfeld( Austria), including specimen illustrated in Hörnes(1851, pl. 2, fig. 4.  Illustrated material. Figs 35C1–C2: Gainfarn ( Austria): SL: 53.4 mm, MD: 26.5 mm, NHMW1856/0050/ 0115; Figs 35D1–D3: syntype, Enzesfeld ( Austria): SL: 48.8 mm, MD: 26.7 mm, NHMW1846/0037/0049, illustrated in Hörnes (1851, pl. 3, fig. 7); Figs 35E1–E3: Mikulov-Kienberk ( Czech Republic): SL: 47.5 mm, MD: 23.8 mm, NHMW1860/0001/0067; Figs 35F: Gainfarn ( Austria): SL: 58.4 mm, MD: 29.2 mm, NHMW1856/ 0050/0115; Figs 35G1–G2: Gainfarn ( Austria): SL: 30.6 mm, MD: 14.3 mm, NHMW1846/0037/0043a; Fig. 30R: Gainfarn ( Austria): SL: 30.1 mm, MD: 15.2 mm, NHMW1846/0037/0043a.  Revised description.Medium-sized moderately slender shells; spire coeloconoid to conical and of variable height. Spire whorls channelled, moderately convex to nearly flat; early spire whorls scalariform, strongly tuberculate and striate (but rarely preserved); later whorls with more or less prominent striae, which may fade out completely on last whorls. Subsutural flexure deep, moderately curved, moderately asymmetrical. Last whorl elongate, slightly ventricose, very weakly constricted with weak spiral grooves on abapical third; shoulder high, more or less prominent, rounded to subangular; position of maximum diameter slightly below shoulder. Siphonal canal short, weakly recurved; siphonal fasciole indistinct. Aperture straight, moderately narrow. Colour pattern under UV light only poorly preserved, consisting of a dense pattern of thin, roughly spirally arranged short dashes and slightly spirally extended dots.   Shell measurements and ratios.n = 20 adultspecimens (only large morphs): largest specimen: SL: 58.4 mm, MD: 29.2 mm, mean SL: 51.9 mm(σ = 3.3), mean MD: 27.2 mm(σ = 1.5), spire angle: µ = 98.3° (σ = 10.4°), last whorl angle: µ = 35.1° (σ = 1.9°), LW: µ = 1.91 (σ = 0.1), RD: µ = 0.63 (σ = 0.03), PMD: µ = 0.89 (σ = 0.03), RSH: µ = 0.17 (σ = 0.03).   Discussion. Hall (1966: 137)discussed a close relation of  Conus vindobonensiswith  Conus ventricosus Gmelin, 1791, which would result in a placement in  Lautoconus. The tuberculate, depressed scalariform early spire, however, contradicts a relation with  Lautoconus. In any case, Hall (1966)was correct to interpret this species as a very polymorphic species comparable to the extant  Lautoconus ventricosus. Especially the height of the spire and the convexity of the spire whorls are very variable. Similarly, the outline of the last whorl ranges from slightly ventricose to nearly straight-sided. Like the recent  L. ventricosus, the spire of the Miocene  C. vindobonensistends to be more or less gradate in some specimens. Consequently, the high spired, gradate specimens illustrated by Hoernes & Auinger (1879, pl. 6, figs 10–11)as  Conus mediterraneusare most probably aberrant specimens of  Conilithes vindobonensis. Therefore,  Conus mediterraneusvar. pupoidemiocenicaand permiocenica, which were introduced by Sacco (1893b)as new names for these specimens, are subjective junior synonyms of  Conus vindobonensis. The same holds true for the aberrant specimen illustrated by Hörnes (1851, pl. 2, fig. 4) as  Conus clavatus, for which Sacco (1893b)introduced the superfluous varietal name mioexclavata, although already Hoernes & Auinger (1879)identified this specimen as their  Conus vindobonensis.  Hall (1966: 137)considered this species to be a subjective junior synonym of  Conus argillicola Eichwald, 1830, from the Badenian of Ukraine. The specimen illustrated by Eichwald (1852, pl. 9, fig. 2)as  C. argillicoladiffers from  C. vindobonensisin its constricted base and the long and reflected siphonal canal. Moreover, the spiral striae on the spire whorls of  C. argillicolaare much stronger. Therefore, we doubt that both taxa represent the same species. Unfortunately, we were not able to find the type specimen(s) of  C. argillicolato solve this question definitely.  Paleoenvironment.Typically found in shallow marine nearshore environments; at Gainfarn, where it is among the most frequent cones, it was associated with seagrass.   Distribution in Paratethys.Badenian (middle Miocene):  ViennaBasin:Gainfarn, Enzesfeld, Steinebrunn, Baden, Baden-Sooß, Bad Vöslau, Möllersdorf, Pötzleinsdorf, Grinzing, ( Austria), Mikulov-Muschelberg, Mikulov-Kienberk, Hrušovany ( Czech Republic), Eisenstadt-Sopron Basin:Marz, Forchtenau ( Austria); Styrian Basin:Pöls; Alpine-Carpathian Foredeep:Grund, Guntersdorf ( Austria) ( Hoernes & Auinger 1879; Sieber 1958a, b), Korytnica, Węglinek, Łychów ( Poland) ( Krach 1981; Bałuk 1997); Pannonian Basin:Hidas ( Hungary) ( Strausz 1966); Transylvanian Basin: Lăpugiu de Sus, Coşteiu de Sus ( Romania) ( Boettger 1902);  BuzăuBasin: Crivineni, Valea Muscel ( Romania); Zârand Basin: Minişul de Sus ( Romania) ( Nicorici & Sagatovici 1973); Caransebeş-Mehadia Basin:Valea Satului ( Romania) ( Hinculov 1968). 1425219270 NHMW Austria Gainfarn Hornes 151 152 NHMW 1846 2 syntype 1425219280 NHMW Austria Enzesfeld Hornes 151 152 NHMW 1846 1 syntype 1425219284 NHMW Austria Gainfarn Hornes 151 152 NHMW 1846 1 1425219271 NHMW Austria Enzesfeld Hornes 151 152 NHMW 1846 1 1425219276 NHMW Austria Gainfarn 151 152 NHMW 1856 1 1425219290 NHMW Austria Gainfarn Steinebrunn 151 152 NHMW 1846 1 1425219278 [220,903,807,832] NHMW Austria Steinebrunn 151 152 NHMW 1846 1 1425219288 NHMW Czech Republic Mikulov-Kienberk 151 152 NHMW 1860 1 1425219281 NHMW Czech Republic Auinger Bad Voslau Hoernes 151 152 NHMW 1849 1 1425219289 NHMW Czech Republic Auinger Bad Voslau Hoernes 151 152 NHMW 1849 1 1425219292 NHMW Austria Enzesfeld Hornes 151 152 NHMW 1846 1