Oxypoda ( Baeoglena ) nova BERNHAUER, 1902: 180 Revision of theBaeoglena species of the West Palaearctic Region (Coleoptera: Staphylinidae: Aleocharinae) Assing, Volker Beiträge Zur Entomologie = Contributions to Entomology 2019 2019-06-24 69 1 1 32 4BHBY BERNHAUER, 1902 Bernhauer 1902 [212,729,1258,1286] Insecta Staphylinidae Oxypoda Animalia Coleoptera 25 26 Arthropoda species caucasica Baeoglena  ( Figs 55–58, Maps 1, 7)      Oxypoda( Baeoglena) nova BERNHAUER, 1902: 180.   Type material examined: Lectotype, present designation: “Caucasus, Helenendorf, Reitter. / fusina Rey/ vid. Bernh. / caucasica Bh./ ex coll. Skalitzky / Lectotypus   Oxypoda caucasicaBernhauer, desig. V. Assing 2018 /  Oxypoda caucasicaBernhauer, det. V. Assing 2018” ( NHMW).   Comment: The original description is based on an unspecified number of syntypesfrom “Caucasus: Helenendorf, Martkopi, Meskisches Gebige [sic], Daghestan, Lenkoran” ( BERNHAUER 1902). A male syntypefrom “Helenendorf ” is designated as the lectotype.   Material examined:   Ukraine: 1, 1 ex., Odessa obl., Dobroolexandrivka[“Alexanderhilf ”; 46°20'N, 30°29'E], leg. Leder( NHMW).    Turkey:  Bolu: 1, 1, road Bolu– Yığılca, 40°51'N, 31°39'N,  1060 m,  29.IV.2012, leg. Brachat& Meybohm(cAss); 5, 5, 23 kmS Bolu, 40°38'N, 31°37'N,  1540 m,  30.IV.2012, leg. Brachat& Meybohm(cAss); 1, Karabük, pass W Eskipazar, 40°58'N, 33°23'E,  1400 m, beech forest, litter sifted,  26.III.2010, leg. Assing(cAss).    Kastamonu: 1, 5 exs.[found together with  O. nova],  5 kmS Küre,  1200 m,  18.V.1976, leg. Besuchet& Löbl( MHNG, cAss).   Sinop: 1, 2, Çangal Dağı,  7–15. VI.1960, leg. Schubert(cAss).   Trabzon: 1, ca.  50 kmS Trabzon, 20 kmS Maçka, Altındere Milli Park, 40°40'N, 39°40'E,  1560 m, spruce forest with Rhododendron, sifted,  26.VII.2006, leg. Schülke( MNB).   Rize: 1, ca. 50 kmS Ardeşen, Çat, 40°52'N, 40°56'E,  1240 m,  Alnusforest, sifted,  3.VIII.2006, leg. Schülke( MNB); 1, ca.  30 kmSW Hopa, river valley of ÇağlayanD., ca. 41°15'N, 41°13'E,  500 m,  29.VI.1998, leg. Solodovnikov(cAss).   Artvin: 6 exs., Borçka,  1–3.VI.1960, leg. Schubert(cAss); 5 exs., pass between Borçka and Hopa,  700 m,  8.VI.1986, leg. Besuchet, Löbl& Burckhardt( MHNG, cAss); 1 ex., same locality,  14.V.1967, leg. Besuchet( MHNG); 68 exs., Karkal Dağı, Pirnalli,  1600 m,  11.VI.1986, leg. Besuchet, Löbl& Burckhardt( MHNG, cAss); 14 exs., Karkal Dağı, below Pirnalli,  1250 m,  11.VI.1986, leg. Besuchet, Löbl& Burckhardt( MHNG, cAss); 3 exs., S  Artvin,  800 m,  7.VI.1986, leg. Besuchet, Löbl& Burckhardt( MHNG); 19 exs., S  Artvin,  1500 m,  9.VI.1986, leg. Besuchet, Löbl& Burckhardt( MHNG, cAss); 41 exs., SW  Artvin,  1900 m,  9.VI.1986, leg. Besuchet, Löbl& Burckhardt( MHNG, cAss).   Kars: 1, Kars,  1750 m,  17.VI.1986, leg. Besuchet, Löbl& Burckhardt( MHNG).   Russia: 1 ex., Krasnodar, 35 kmNNE Sochi, Babuk-Aul, 43°53'N, 39°49'E, 560 m,  Fagus orientalisand  Castanea sativaforest, litter and bark sifted, 11.VII.2011, leg. Assing (cAss); 2 exs., Krasnodar, Temnolesskaia, near Mezmay, 800 m, 8.VI.1999, leg. Smetana (cAss); 25 exs.[partly teneral], Karachayevo-Cherkesskaya Respublika, 13 kmSW Teberda, 43°20'N, 41°40'E, 1450 m, moist spruce forest with scattered beech, litter, moss, and dead wood sifted, 22.VII.2011, leg. Assing (cAss); 10 exs.[partly teneral], Karachayevo-Cherkesskaya Respublika, 20 kmSW Teberda, above Dombai, 43°17'N, 41°38'E, 1950 m, mixed forest (fir, spruce, beech), leaf litter sifted, 25.VII.2011, leg. Assing (cAss); 1 ex., Karachayevo-Cherkesskaya Respublika, 9 kmSW Teberda, Teberdinski range, Baduk river valley, 43°23'N, 41°40'E, 2000 m, subalpine forest (maple, birch, with rhododendron) near lakeshore, litter sifted, 26.VII.2011, leg. Assing (cAss); 11 exs., Karachayevo-Cherkesskaya Respublika, 9 kmSW Teberda, Teberdinski range, Baduk river valley, 43°23'N, 41°40'E, 2000 m, spruce forest, bark of spruce and maple sifted, 27.VII.2011, leg. Assing (cAss); 1 ex., Krasnaja Poljana, Mt. Atchichkho, 800 m, 26.VI.1989, leg. Arndt ( SDEI); 2 exs., Krasnaja Poljana, leg. Roubal ( NHMW).    Georgia: 6 exs., Algeti National Park, W Manglisi, 41°42'N, 44°21'E,  1120 m,  11.VII.2015, leg. Brachat& Meybohm(cAss);  6 exs., Algeti National Park, W Manglisi, 41°42'N, 44°19'E,  1110 m,  11.VII.2015, leg. Brachat& Meybohm(cAss);  1 ex., Algeti National Park, W Manglisi, 41°42'N, 44°18'E,  1210 m,  11.VII.2015, leg. Brachat& Meybohm(cAss);  1 ex., Algeti National Park, Manglisi, 41°42'N, 44°23'E,  1190 m,  12.VII.2015, leg. Brachat& Meybohm(cAss);  10 exs., same data, but  13.VII.2015(cAss);  1 ex., Central Caucasus, Schatili–Mutso, 42°40'N, 45°11'E,  1390 m,  15.VII.2015, leg. Brachat& Meybohm(cAss);  5 exs., Central Caucasus, Gudani, 42°32'N, 44°59'E,  1710 m,  18.VII.2015, leg. Brachat& Meybohm(cAss);  4 exs., Gudani–Zhinvali, 42°27'N, 44°56'E,  1200 m,  19.VII.2015, leg. Brachat& Meybohm(cAss);  8 exs., Gudani–Zhinvali, 42°27'N, 44°56'E,  1200 m,  19.VII.2015, leg. Brachat& Meybohm(cAss);  1 ex., Shida Kartli, Kvishkheti, 41°58'N, 43°30'E,  810 m,  12.V.2016, leg. Brachat& Meybohm(cAss);  6 exs., Shida Kartli, Kvishkheti, 41°57'N, 43°30'E,  790 m,  12.V.2016, leg. Brachat& Meybohm(cAss);  6 exs., Samtskhe-Javakheti, Timotesubani, 41°49'N, 43°31'E,  1144 m,  13.V.2016, leg. Brachat& Meybohm(cAss);  3 exs., Samtskhe-Javakheti, Bakuriani, 41°44'N, 43°43'E,  1770 m,  13.V.2016, leg. Brachat& Meybohm(cAss);  2 exs., Samtskhe-Javakheti, N Abastumani, 41°46'N, 42°50'E,  1370 m,  15.V.2016, leg. Brachat& Meybohm(cAss);  2 exs., Samtskhe-Javakheti, W Abastumani, 41°45'N, 42°49'E,  1345 m,  15.V.2016, leg. Brachat& Meybohm(cAss);  1 ex., Abastumani, leg. Leder( NHMW);  3 exs., Racha,  4 kmN Nakerala, 42°24'N, 43°02'E,  1150 m,  18.V.2016, leg. Brachat& Meybohm(cAss);  8 exs., Shida Kartii, Kvishkheti, 41°57'N, 43°29'E,  1300 m,  24.VII.2016, leg. Meybohm(cAss);  1 ex., Zemo Svaneti,  4 kmN Mazeri, 43°06'N, 42°36'E,  1690 m,  28.VII.2016, leg. Meybohm(cAss);  2 exs., Zemo Svaneti,  2 kmN Ipari, 43°01'N, 42°50'E,  1670 m,  29.VII.2016, leg. Meybohm(cAss);  1 ex., Adjara, Skhalta valley, 41°35'N, 42°22'E,  800 m,  22.VI.2017, leg. Brachat& Meybohm(cAss);  4 exs., Adjara, Skhalta valley, 41°34'N, 42°25'E,  970 m,  22.VI.2017, leg. Brachat& Meybohm(cAss);  3 exs., Svaneti, Mazeri, 43°06'N, 42°36'E,  1660 m,  26.VI.2017, leg. Brachat& Meybohm(cAss);  1 ex., Svaneti, Mestia–Hatsvali, 43°01'N, 42°45'E,  2350 m,  27.VI.2017, leg. Brachat& Meybohm(cAss);  4 exs., Svaneti, Mestia–Ushguli,  3 kmN Ipari, 43°01'N, 42°50'E,  1750 m,  28.VI.2017, leg. Brachat& Meybohm(cAss);  1 ex., Svaneti, Mestia–Ushguli, Kala, 42°57'N, 42°53'E,  1680 m,  28.VI.2017, leg. Brachat& Meybohm(cAss);  1 ex., Imereti, Baghdati, 42°01'N, 42°49'E,  290 m,  18.V.2018, leg. Brachat& Meybohm(cAss);  3 exs., Imereti, S Sairme, 41°51'N, 42°47'E,  1930 m,  19.V.2018, leg. Brachat& Meybohm(cAss);  1 ex., Imereti, S Sairme, 41°52'N, 42°47'E,  1890 m,  19.V.2018, leg. Brachat& Meybohm(cAss);  1 ex., Imereti, S Sairme, 41°53'N, 42°46'E,  1420 m,  20.V.2018, leg. Brachat& Meybohm(cAss);  1 ex., Imereti, S Sairme, 41°53'N, 42°46'E,  1360 m,  20.V.2018, leg. Brachat& Meybohm(cAss);  6 exs., Racha, E Shovi, 42°41'N, 43°42'E,  1720 m,  28.V.2018, leg. Brachat& Meybohm(cAss);  3 exs., Racha, E Shovi, 42°41'N, 43°42'E,  1700 m,  28.V.2018, leg. Brachat& Meybohm(cAss);  2 exs., Racha, W Glola, 42°42'N, 43°36'E,  1170 m,  28.V.2018, leg. Brachat& Meybohm(cAss);  1 ex., Ananuri Forest,  8.V.2006, leg. Chalandze(cAss);  1 ex., Trialetskiy Khrebet, Bakuriani,  1500–1600 m,  8.VII.1986, leg. Wrase& Schülke( SDEI);  1 ex., Tbilisienv., Zchneti,  800 m,  VI.1987, leg. Wrase& Schülke( SDEI);  2 exs.,  60 kmN Tbilisi, Dusheti,  1500 m,  25.IX.1987, leg. Oehlke( SDEI);  3 exs., Khashuri[ 41°59'N, 43°35'E] ( NHMW);  1 ex., “ Swanetien”, leg. Leder( NHMW);  6 exs., Likhskiy Khrebet[“Meskisches Geb.”], leg. Leder( NHMW).   Armenia: see ASSING & SCHÜLKE (2019).  Nagorno-Karabakh: see ASSING & SCHÜLKE (2019).    Azerbaijan: 2 exs., Lenkoran, leg. Leder( NHMW);  1 ex., Lerik, leg. Leder( NHMW);  4 exs., Talyshrange ( MHNG, NHMW).    Iran:  Mazandaran: 1 ex., Elburz Mts., Ramsar Co., Eshkatechal, 36°51'N, 50°33'E,  1050 m, small stream, sifted,  6.VI.2008, leg. Pütz(cAss);  1 ex., Elburz Mts., Babol Co.,  2 kmSW Firuz Jah, 36°11'N, 52°39'E,  840 m, small stream,  31.V.2008, leg. Pütz(cAss);  2 exs., Elburz Mts., Tonekabon Co., 4.5 km SW Khanian, Sehezar forest, 36°33'N, 50°50'E,  940 m, small stream, litter and debris sifted,  5.VI.2008, leg. Pütz(cAss);  4 exs., Elburz Mts., Sari Co., Mohammabad, NE Sangdeh, 36°04'N, 53°10'E,  1530 m, beech forest, litter sifted,  29.V.2008, leg. Pütz(cAss).  1 ex., Elburz Mts., Sari Co., Mohammabad, E Qolqol, 36°10'N, 53°10'E,  920 m, litter sifted,  29.V.2008, leg. Pütz(cAss);  1 ex., Elburz Mts., Chalus Co., SE Abbasabad, 36°39'N, 51°10'E,  280 m, leaf litter and debris sifted,  4.VI.2008, leg. Pütz( MNB);  3 exs., Taquiabad,  24.VII.1977, leg. Huber( MHNG).   Gilan: 1 ex., Elburz Mts., Rasht Co., 29 kmS Rasht, 37°01'N, 49°37'E, 140 m, sifted, 7.VI.2008, leg. Pütz (cAss).  Locality ambiguous, not identified, or not specified:  1 ex., “Armen. Geb.”, leg. Leder( NHMW);  6 exs., “Kaukas”, “Caucasus”, etc., leg. Leder, Reitter, etc. ( NHMW);  1 ex., “Kaukas, ... Urwald bei Tcharsach”,  1.V.1875, leg. Leder( MHNG).   Diagnosis: Winged species of mostly dark and more rarely reddish coloration. : median lobe of aedeagus ( Figs 55–56) approximately 0.35 mm long; ventral process weakly curved in lateral view, apically truncate in lateral view and convex in ventral view; crista apicalis long and broad; internal sac with pair of fork-shaped apical structures (lateral view) and with a weakly curved and weakly sclerotized flagellum; apical portion of paramere ( Fig. 57) relatively short, barely half as long as basal portion, rather weakly dilated basally. : spermatheca ( Fig. 58) with minute apical cuticular invagination.   Distribution and natural history: The distribution of  O. caucasicaranges from South Ukraineand Northwest Anatoliaacross North Anatolia, the Greater Caucasus, and the Lesser Caucasus (including adjacent regions) to North Iran( Maps 1, 7). The record from Germanyin SCHÜLKE & SMETANA (2015)is based on LOHSE (1989: 231)“nach Angaben von L. Zerche”, who reports a male from the environs of Gotha collected in 1867. This specimen was clearly either mislabelled or misidentified, as can be inferred from the revised distribution of  O. caucasica. The examined material was sifted from leaf litter, roots, moss, and bark in various forest (deciduous, coniferous, mixed) habitats and near streams. The altitudes range from 140 to 2350 m. The specimens were found in March (one record), April (2), May (26), June (27), July (37), August (1), and September (1). NHMW Leder Ukraine 46.333332 Dobroolexandrivka 1124 30.483334 25 26 1 Odesa 2010-03-26 2012-04-30 2010-03-26 Brachat & Meybohm & W Eskipazar & Assing Turkey 1060 40.966667 Yigilca 1159 33.383335 25 26 1 Bolu 1976-05-18 MHNG Besuchet & Lobl 1200 5 km S Kure 25 26 5 Kastamonu 1960-06-07 1960-06-15 1960-06-07 Schubert Cangal Dagi 25 26 1 Sinop 2006-07-26 MNB Schulke Macka 1560 40.666668 Rhododendron 1161 39.666668 Altindere Milli Park 25 26 1 Trabzon 1998-06-29 2006-08-03 1998-06-29 MNB Schulke & Solodovnikov Ardesen 1240 41.25 Caglayan 1157 41.216667 Cat 25 26 1 Rize 1960-06-01 1986-06-11 1960-06-01 MHNG Schubert & Besuchet & Lobl & Burckhardt Borcka 700 41.25 Pirnalli 1157 41.216667 Karkal Dagi 25 26 97 Artvin 1986-06-07 MHNG Besuchet & Lobl & Burckhardt 800 Artvin 25 26 19 Artvin 1986-06-09 MHNG Besuchet & Lobl & Burckhardt 1500 Artvin 25 26 41 Artvin 1986-06-09 MHNG Besuchet & Lobl & Burckhardt 1900 Artvin 25 26 1 Artvin 1986-06-17 MHNG Besuchet & Lobl & Burckhardt 1750 Kars 25 26 1 Kars 2015-07-11 Brachat & Meybohm Georgia 1120 41.7 Manglisi 1155 44.35 Algeti National Park 26 27 6 2015-07-11 Brachat & Meybohm Georgia 1110 41.7 Manglisi 1155 44.316666 Algeti National Park 26 27 6 2015-07-11 Brachat & Meybohm Georgia 1210 41.7 Manglisi 1155 44.3 Algeti National Park 26 27 1 2015-07-12 Brachat & Meybohm Georgia 1190 41.7 Manglisi 1155 44.383335 Algeti National Park 26 27 1 2015-07-13 Brachat & Meybohm Georgia 1190 41.7 Manglisi 1155 44.383335 Algeti National Park 26 27 10 2015-07-15 Brachat & Meybohm Georgia 1390 42.666668 Schatili 1148 45.183334 Central Caucasus 26 27 1 2015-07-18 Brachat & Meybohm Georgia 1710 42.533333 Gudani 1149 44.983334 Central Caucasus 26 27 5 2015-07-19 Brachat & Meybohm Georgia 1200 42.45 Gudani - Zhinvali 1150 44.933334 26 27 4 2015-07-19 Brachat & Meybohm Georgia 1200 42.45 Gudani - Zhinvali 1150 44.933334 26 27 8 2016-05-12 Brachat & Meybohm Georgia 810 41.966667 Kvishkheti 1153 43.5 26 27 1 Shida Kartli 2016-05-12 Brachat & Meybohm Georgia 790 41.95 Kvishkheti 1153 43.5 26 27 6 Shida Kartli 2016-05-13 Brachat & Meybohm Georgia 1144 41.816666 Timotesubani 1153 43.516666 26 27 6 Samtskhe-Javakheti 2016-05-13 Brachat & Meybohm Georgia 1770 41.733334 Bakuriani 1154 43.716667 26 27 3 Samtskhe-Javakheti 2016-05-15 Brachat & Meybohm Georgia 1370 41.766666 Abastumani 1154 42.833332 26 27 2 Samtskhe-Javakheti 2016-05-15 Brachat & Meybohm Georgia 1345 41.75 Abastumani 1154 42.816666 26 27 2 Samtskhe-Javakheti NHMW Leder Georgia Abastumani 26 27 1 2016-05-18 Brachat & Meybohm Georgia 1150 42.4 Racha 1150 43.033333 26 27 3 2016-07-24 Meybohm Georgia 1300 41.95 Kvishkheti 1153 43.483334 Shida Kartii 26 27 8 2016-07-28 Meybohm Georgia 1690 43.1 Zemo Svaneti 1145 42.6 26 27 1 2016-07-29 Meybohm Georgia 1670 43.016666 Zemo Svaneti 1146 42.833332 26 27 2 2017-06-22 Brachat & Meybohm Georgia 800 41.583332 Skhalta valley 1155 42.366665 26 27 1 Ajaria 2017-06-22 Brachat & Meybohm Georgia 970 41.566666 Skhalta valley 1155 42.416668 26 27 4 Ajaria 2017-06-26 Brachat & Meybohm Georgia 1660 43.1 Mazeri 1145 42.6 Svaneti 26 27 3 2017-06-27 Brachat & Meybohm Georgia 2350 43.016666 Mestia - Hatsvali 1146 42.75 Svaneti 26 27 1 2017-06-28 Brachat & Meybohm Georgia 1750 43.016666 Mestia - Ushguli 1146 42.833332 Svaneti 26 27 4 2017-06-28 Brachat & Meybohm Georgia Svaneti 1680 42.95 Kala 1146 42.883335 Mestia - Ushguli 26 27 1 2018-05-18 Brachat & Meybohm Georgia 290 42.016666 Baghdati 1152 42.816666 26 27 1 Imereti 2018-05-19 Brachat & Meybohm Georgia 1930 41.85 Sairme 1153 42.783333 26 27 3 Imereti 2018-05-19 Brachat & Meybohm Georgia 1890 41.866665 Sairme 1153 42.783333 26 27 1 Imereti 2018-05-20 Brachat & Meybohm Georgia 1420 41.883335 Sairme 1153 42.766666 26 27 1 Imereti 2018-05-20 Brachat & Meybohm Georgia 1360 41.883335 Sairme 1153 42.766666 26 27 1 Imereti 2018-05-28 Brachat & Meybohm Georgia 1720 42.683334 Shovi 1148 43.7 Racha 26 27 6 2018-05-28 Brachat & Meybohm Georgia 1700 42.683334 Shovi 1148 43.7 Racha 26 27 3 2018-05-28 Brachat & Meybohm Georgia 1170 42.7 Glola 1148 43.6 Racha 26 27 2 2006-05-08 Chalandze Georgia Ananuri Forest 26 27 1 1986-07-08 SDEI Wrase & Schulke Georgia 1550 Bakuriani Trialetskiy Khrebet 26 27 1 1987-06 SDEI Wrase & Schulke Georgia 800 Zchneti 26 27 1 Tbilisi 1987-09-25 SDEI Oehlke Georgia 1500 Dusheti 26 27 2 Tbilisi NHMW Georgia 41.983334 Khashuri 1153 43.583332 26 27 3 NHMW Leder Georgia Swanetien 26 27 1 NHMW Leder Georgia Likhskiy Khrebet 26 27 6 [854,1394,1535,1560] NHMW Leder Azerbaijan Lenkoran 26 27 2 Lankaran NHMW Leder Azerbaijan Lerik 26 27 1 Lerik MHNG, NHMW Talysh Azerbaijan 26 27 4 2008-06-06 Putz Iran Ramsar Co. 1050 36.85 Eshkatechal 1185 50.55 Mts. 26 27 1 Mazandaran 2008-05-31 Putz Iran Babol Co. 840 36.183334 Mts. 1189 52.65 26 27 1 Alborz 2008-06-05 Putz Iran Tonekabon Co. 940 36.55 Khanian 1186 50.833332 Mts. 26 27 2 Alborz 2008-05-29 Putz Iran Sari Co. 1530 36.066666 Mohammabad 1190 53.166668 Mts. 26 27 4 Alborz 2008-05-29 Putz Iran Sari Co. 920 36.166668 Mohammabad 1189 53.166668 Mts. 26 27 1 Alborz 2008-06-04 MNB Putz Iran Chalus Co. 280 36.65 Abbasabad 1186 51.166668 Mts. 26 27 1 Alborz 1977-07-24 MHNG Huber Iran 27 28 Taquiabad 26 27 3 Mazandaran [564,785,255,280] Iran Gilan 27 28 1 Gilan [174,603,416,440] NHMW Leder 27 28 1 NHMW Leder & Reitter 27 28 6 1875-05-01 MHNG Leder 27 28 1