The Ants (Hymenoptera: Formicidae) of Bol’shaya Svetlovodnaya (Late Eocene of Sikhote-Alin, Russian Far East)
Dlussky, G. M.
Rasnitsyn, A. P.
Perfilieva, K. S.
Caucasian Entomological Bulletin
2015
11
1
131
152
Dlussky & Rasnitsyn & Perfilieva, 2015
Dlussky & Rasnitsyn & Perfilieva
2015
[952,1197,1465,1486]
Insecta
Formicidae
Ypresiomyrma
GBIF,CoL
Animalia
Hymenoptera
8
139
Arthropoda
species
orientalis
sp. n.
( Fig. 18, 19)
Material.Holotype PIN 3429/1109 (print and counterprint), dorsal imprint of gyne without head. Paratype: PIN 3429/1198, forewing.
Description.Gyne. Judging by the remained parts body length approximately 17.5 mm. Mesosoma robust, 1.5 times as long as wide. Petiole nodiform, a little longer than wide. 1st gastral (abdominal III) segment 0.8 times as long and 0.7 times as wide as 2nd gastral (abdominal IV) segment. Forewing with closed cells1+2r, 3r, rm and mcu. Cell3r elongated, 4.5–5times as long as wide. Cell1+2r shorted, 2–2.2times as long as wide. The meeting of crossvein rs-m and RS is distal to 2r-rs, section 4RS nearly as long as cross-vein rs-m. Section 2-3RS bent to form rounded obtuse angle in its basal third and so cell rm hexogonal, twice as long as wide. Transverse rs-m position at level of pterostigma top. Section1M 2–2.5times as long as1RS. Cell mcu pentagonal. Cross-veins cu-a and section 1M meet M+Cu at the same point, essentially forming a single crossvein, so Icu=1. Measurements, mm. Holotype PIN 3429/1109: AL6; PtL1.4; PtW1.3; FWL9.8. Paratype PIN 3429/1198: FWL9.6.
Comparison.Wing venation and short the 1st gastral segment indicate unquestionably a position of the fossils in the subfamily Myrmeciinae. The gaster lacking a constriction between the 1st and the 2nd segments in combination with a robust body are further characteristic of the genus Ypresiomyrma. The new species well differs from Y. rebekkaein much less size of the petiolar node. No reliable difference from the North American species are found because of different (lateral) burial position of Y.bartlettiand Y.orbiculataand incomplete presentation of their wings. Yet a distinct species status of the new species is beyond doubts because all other congeners are distinctly more old (known from the Early and Middle Eocene).
Figs 16–21. Species of genera Lasius, Ypresiomyrma, Casaleiaand Cerapachyinaeinc. gen. 16–17– Lasiussp. B: 16– PIN 3429/1111, male, 17 – PIN 3429/102, gyne; 18–19– Ypresiomyrma orientalis sp.n.: 18– holotype PIN 3429/1109, gyne, left forewing from counterprint, 19–PIN 3429/1198, forewing; 20– Cerapachyinae inc. gen., PIN 3429/1132, gyne or worker; 21– Casaleia orientalis sp.n., holotype PIN 3429/1174, gyne. Рис. 16–21. Виды родов Lasius, Ypresiomyrma, Casaleiaи Cerapachyinae inc. gen. 16–17– Lasiussp. B: 16– PIN 3429/1111, самец, 17 – PIN 3429/102, самка; 18–19– Ypresiomyrma orientalis sp.n.: 18– голотип PIN 3429/1109, самка, левое переднее крыло реконструировано по обратному отпечатку, 19 – PIN 3429/1198, переднее крыло; 20 – Cerapachyinae inc. gen., PIN 3429/1132, самка или рабочий; 21– Casaleia orientalis sp.n., голотип PIN 3429/1174, самка.
Etymology. Orientalisis the Latin for eastern.