The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species (Coleoptera, Gyrinidae, Haliplidae, Noteridae, Dytiscidae) Ramahandrison, Andriamirado Tahina Rakouth, Bakolimalala Manuel, Michael ZooKeys 2022 2022-11-02 1127 1 60 7SVCF 04747A9A-EF84-51F3-9CBE-78770B59079E Sharp, 1882 Sharp 1882 Insecta Dytiscidae Cybister CoL Animalia Cybister operosus Coleoptera 0 1 Arthropoda species operosus  Type locality. Madagascar.  Material examined.  1 ♀: MAK-50.  Distribution. Madagascar (widespread but localised) ( Guignot 1959-1961; Bukontaite et al. 2015).  Habitat in study area. This species was captured only once, in northern inner Makay, in a large temporary pond located in a shallow open area on sand. This single capture does not reflect the usual habitat preferences of this species. According to our observations in Isalo and Ankarafantsika, this is a lotic species (an exceptional ecology for the genus) inhabiting the margins of streams and rivers with some vegetation and / or tree roots and / or plant debris, in well-preserved forested or semi-forested environments.  Subfamily Dytiscinae, tribe Aciliini