Smeringopus thomensis Simon 1907: 251–252 Revision and cladistic analysis of the Afrotropical endemic genus Smeringopus Simon, 1890 (Araneae: Pholcidae) HUBER, BERNHARD A. Zootaxa 2012 2012-09-07 3461 1 1 138 4XVJR Simon, 1907 Simon 1907 [151,600,1597,1623] Arachnida Pholcidae Smeringopus Animalia Araneae 113 114 Arthropoda species thomensis      Smeringopus thomensis Simon 1907: 251–252.  Type.  Male holotypefrom São Tomé and Principe, São Tomé Island, Ribeira Palma[ 0°21’N, 6°35’E], leg. L. Fea, possibly in Museo Civico di Storia Naturale "Giacomo Doria", Genova, not examined. Diagnosis. Distinguished from similar congeners (other species of the  thomensisspecies group) by shape of embolus (two large rounded apophyses arising from embolus dorsally; Figs. 722, 723) and absence of pair of sclerotized areas on posterior rim of epigynum ( Fig. 709); from  S. principealso by absence of dorsal projection from procursus tip (compare Figs. 720and 725) and much narrower pore plate (compare Figs. 724and 728). Easily distinguished from all other congeners by distinctive color pattern dorsally on abdomen (figure composed of three black elements on posterior half; Fig. 699).   FIGURES 699–708.  Smeringopus thomensisgroup.  S. thomensis(699–700, 705–706),  S. principe(702–704, 707–708), and  S. mayombe(701). 699, 701, 703. Males, dorsal views. 700, 702. Male prosomata, oblique frontal views. 704. Female, ventral view. 705–708. Left male palps, prolateral and retrolateral views. Male (Palha Plantation). Total body length 5.0, carapace width 1.5. Leg 1: 45.3 (11.9 + 0.5 + 10.8 + 20.1 + 2.0), tibia 2: 7.2, tibia 3: 5.1, tibia 4: 7.2; tibia 1 L/d: 64. Habitus as in Fig. 699. Carapace ochre-yellow with median and lateral dark pattern, clypeus distal half darker, without pair of lines, sternum dark brown with some small light spots, legs ochre-yellow, femora and tibiae with dark subdistal rings and light tips, abdomen grey with distinctive dorsal pattern, ventrally with dark anterior and posterior plates and three indistinct bands in median section. Distance PME-PME 125 µm, diameter PME 150 µm, distance PME-ALE 55 µm, distance AME-AME 30 µm, diameter AME 120 µm. Ocular area slightly elevated, secondary eyes with small dark ‘pseudo-lenses’; deep thoracic pit. Chelicerae with pair of small distal apophyses ( Fig. 721), each provided with one modified hair at tip ( Fig. 734). Palps as in Figs. 705 and 706, coxa with distinct retrolateral apophysis, trochanter barely modified, femur with deep retrolateral furrow, proximal rim distinct ( Fig. 732), procursus with distinctive distal processes ( Figs. 719, 720), bulb with complex embolus with distinctive rounded dorsal processes ( Figs. 722, 723). Legs without spines, few vertical hairs, without curved hairs, retrolateral trichobothrium on tibia 1 at 2%; prolateral trichobothrium present on tibia 1. Gonopore with two epiandrous spigots ( Fig. 736); ALS with one widened and one pointed spigot each, other spigots apparently reduced to two tiny remnants (cf. female, Fig. 737).   FIGURES 709–717.  Smeringopus thomensisgroup. Epigyna, ventral views and cleared female genitalia, ventral and dorsal views. 709–711.  S. thomensis. 712–714.  S. principe. 715–717.  S. mayombe. Variation. Tibia 1 in3 other males: 8.9, 10.1, 10.7. Female. In general similar to male; tibia 1 in 5 females: 8.4–9.7 (mean 9.1). Epigynum a simple, roughly triangular plate with pockets ( Figs. 709, 710), posterior plate semicircular; internal genitalia as in Figs. 711, 724, and 739. Distribution. Known from São Tomé Islandonly ( Fig. 718).  Materialexamined.  SÃO TOMÉ AND PRÍNCIPE:  São Tomé: Palha Plantationnear São Tométown ( 0°18.8’N, 6°42.4’E),  12.iv.2001( C.E. Griswold), 4♂ 5♀in CAS.  Ponta Barro Bóró( 0°23.8’N, 6°36.3’E), in culvert along roadside,  27.iv.2001( J.M. Ledford), 1♀3 juvs in CAS.  Augustino Neto, Rio de Oro Plantation( 0°21.9’N, 6°38.7’E),  220 ma.s.l.,  13.iv.2001( C.E. Griswold), 1♀in CAS.  São Tomé, no further locality data,  1.xi.1999(W. Tavernier), 1♂ 1 juv.in MRAC(209235); same data but garden,  17.xi.1999, 1♂ 1 juv.in MRAC(209256). 3747820825 L. Fea Sao Tome and Principe 0.35 Ribeira Palma 1308 6.5833335 Museo Civico di Storia Naturale 113 114 1 Sao Tome Island holotype 3747820723 2001-04-12 CAS C. E. Griswold Sao Tome and Principe 0.31333333 Palha Plantation 130 6.706667 Material 115 116 9 5 4 Sao Tome 3747820521 2001-04-27 CAS J. M. Ledford Sao Tome and Principe 0.39666665 Ponta Barro Boro 130 6.605 115 116 1 1 Sao Tome 3747820649 2001-04-13 CAS C. E. Griswold Sao Tome and Principe 220 0.36499998 Rio de Oro Plantation 130 6.645 Augustino Neto 115 116 1 1 Sao Tome 3747820850 1999-11-01 MRAC Tavernier Sao Tome and Principe Sao Tome 115 116 2 1 Sao Tome 3747820892 1999-11-17 MRAC Tavernier Sao Tome and Principe Sao Tome 115 116 2 1 Sao Tome