Smeringopus hypocrita Simon 1910: 190 Smeringopus similis Kraus 1957: 230–232 S. hypocrita S. hypocrita Revision and cladistic analysis of the Afrotropical endemic genus Smeringopus Simon, 1890 (Araneae: Pholcidae) HUBER, BERNHARD A. Zootaxa 2012 2012-09-07 3461 1 1 138 4XVJQ Kraus, 1957 Kraus 1957 [151,554,1658,1685] Arachnida Pholcidae Smeringopus Animalia Araneae 110 111 Arthropoda species similis   Figs. 563, 586–587, 606–607, 686–692   “  Smeringopus hypocrita” (misidentification):  Simon 1910: 190(female from Kubub only).     Smeringopus similis Kraus 1957: 230–232, figs. 34–40. Types. All types from Namibia:  Male holotypefrom Otjozondjupa Region, Okahandja[ 21°59’S, 16°55’E],  v.1955( Gaerdes), in SMF(RII 10626; only one palp, chelicerae missing).  4♀ 1 juv. paratypes(3 vials), same data, in SMF (RII 11054–56).  1♀2 juvs paratypesfrom Omaheke Region, Gobabis [ 22°27’S, 18°58’E], date and collector not given, in SMF (RII 7677).  1♀ paratypefrom Kubub [locality not identified, see Note under  S. hypocritaredescription], iv.1904(L. Schultze), misidentified by E. Simon as  S. hypocrita, in ZMB (10461).  3♂ 7♀ paratypesfrom Oshikoto Region, Otjikoto, 20 kmW Tsumeb [ 19°11.5’S, 17°33.0’E], 16.vi.1911(W. Michaelsen), in ZMH.  1 juv. paratypefrom Otjozondjupa Region, Grootfontein [ 10°34’S, 18°07’E], 7./ 11.vi.1911(W. Michaelsen), in ZMH.  1 juv. paratypefrom Otjozondjupa Region, Okaputo, 64 kmSW Otavi [ 20°07’S, 16°58’E], 5.vi.1911(W. Michaelsen), in ZMH. All types examined.  Diagnosis. Distinguished from similar congeners by tip of procursus (distinctive sclerotized and membranous elements; Figs. 686–688), shape of embolus ( Fig. 689–691; distinctive dorsal view); from  S. peregrinoidesalso by more downward directed cheliceral apophyses (similar  S. peregrinus; cf. Figs. 614, 615) and absence of v-shaped or u-shaped structure frontally in female internal genitalia ( Fig. 606); from  S. atomariusand  S. uisib(which have overlapping geographic distributions) also by larger epigynal pockets closer together ( Fig. 606). Male ( paratypefrom Otjikoto). Total body length 6.9, carapace width 2.2. Leg 1: 46.2 (12.4 + 0.9 + 12.3 + 17.9 + 2.7), tibia 2: 8.9, tibia 3: 6.8, tibia 4: 9.6; tibia 1 L/d: 50. Habitus as in Fig. 563. Color pattern similar  S. peregrinoidesbut less distinct (possibly artificially), clypeus lines wider, light marks on sternum larger, ventral abdominal pattern limited to pair of interrupted lines. Distance PME-PME 160 µm, diameter PME 150 µm, distance PME-ALE 70 µm, distance AME-AME 45 µm, diameter AME 140 µm. Ocular area slightly elevated, secondary eyes with ‘pseudo-lenses’; deep thoracic pit. Chelicerae very similar  S. peregrinus(cf. Figs. 614, 615). Palps as in Figs. 586 and 587, coxa with distinct retrolateral apophysis and shallow wide furrow, trochanter barely modified, femur with retrolateral furrow with distinct rim proximally, procursus with distinctive distal elements ( Figs. 686–688), bulb with distinctively shaped prolateral process on embolus ( Figs. 689–691). Legs without spines, few vertical hairs, with curved hairs on tibiae and metatarsi 1 and 2, retrolateral trichobothrium on tibia 1 at 3%; prolateral trichobothrium present on tibia 1. Variation. Some males with small conical projection ventrally on embolus (arrow in Fig. 689) and slightly more slender distal apophysis of procursus (in dorsal view). Males from Otavi and Andara-Kavango and two malesfrom Otjikoto with many light brown marks on femora; specimens (male and females) from Otavi also otherwise with much more distinct color pattern (possibly the normal condition). Tibia 1 in 8 males: 6.3–14.8 (mean 10.7); in holotype: 14.8. Female. In general similar to male; tibia 1 in 22 females: 7.2–12.7 (mean 10.1). Epigynum a rather pale indistinct plate with distinct, round pockets ( Fig. 606); most females from the two eastern-most localities ( Andara-Kavangoand Sitwa) have smaller pockets wider apart and may thus not be conspecific with the males from these localities; internal genitalia as in Figs. 607and 692(very similar  S. kalomo,  S. chibububo). Distribution. Widely distributed in northern Namibia( Fig. 608).  Materialexamined.  NAMIBIA:  Otjozondjupa: Okahandja, 1♂ holotypeand 4♀ 1 juv. paratypesabove. Grootfontein, Totes Tal Cave( 19°56’S, 17°23’E), in cave,  11.iv.1991( J. Irish), 1♀in NMBA(5729).  Grootfontein, 1 juv. paratypeabove. Okaputo:  1 juv. paratypeabove.  Omaheke: Gobabis,  1♀2 juvs paratypesabove.  Oshikoto: Otjikoto,  3♂ 7♀ paratypesabove. [Near] Otavi, Ghaub Cave[ 19°29.0’S, 17°46.7’E],  29.iv.1972( P. Wrede, P. Strinati),  1♂ 2♀+ juvs in MHNG.   Omusati: Etosha National Park, Bitterwater[~ 18°56’S, 15°06’E],  16.iii.1998( A. Russell-Smith), 1♂in ZFMK( Ar8564).  Etosha National Park,  iii.1998( A. Russell-Smith), 1♂in MRAC(215622).   Okawango: Andara-Kavango, Okavangoriv. [ 18°04’S, 21°27’E], 1979 ( M.E. Baddeley), 1♂ 6♀in MRAC(152787).   Caprivi: Sitwanear Choy,  10 kmS Kongola[ 17°52’S, 23°22’E], under bark,  19.x.1987(R. Jocqué), 1♂ 2♀ 1 juv.in MRAC(168618).   Khomas: Windhoek[ 22°34’S, 17°05’E], date and collector not given, 1♂in SMF(part of RII 6618; misidentified by Kraus 1957as  S. atomarius). Unidentified locality: Kubub: 1♀ paratypeabove. 3747820486 1955-05 SMF -21.983334 Gaerdes 1262 16.916666 Okahandja 110 111 1 1 Otjozondjupa Region holotype 3747820898 110 111 paratype 3747820854 -22.45 18.966667 110 111 paratype 3747820587 110 111 paratype 3747820492 111 112 -19.191668 17.55 110 111 paratype 3747820589 -10.566667 18.116667 111 112 paratype 3747820386 -20.116667 16.966667 111 112 paratype 3747820858 1991-04-11 NMBA J. Irish Namibia Material -19.933332 Grootfontein 1270 17.383333 Okahandja 112 113 7 5 1 Otjozondjupa holotype 3747820832 Namibia Grootfontein 112 113 1 1 Otjozondjupa paratype 3747820953 [521,1001,836,860] Namibia Grootfontein 112 113 1 1 Otjozondjupa paratype 3747820603 Namibia Grootfontein 112 113 1 1 3 Otjozondjupa paratype 3747820870 1972-04-29 MHNG P. Wrede & P. Strinati Namibia -19.483334 Ghaub Cave 127 17.778334 Otavi 112 113 10 7 3 Otjozondjupa paratype 3747820695 [256,522,908,932] 1972-04-29 MHNG P. Wrede & P. Strinati Namibia -19.483334 Grootfontein 127 17.778334 Otavi 112 113 3 2 1 Otjozondjupa paratype 3747820369 1998-03-16 ZFMK A. Russell-Smith & Ar Namibia -18.933332 Bitterwater 1273 15.1 Etosha National Park 112 113 1 1 Omusati paratype 3747820900 1998-03 MRAC A. Russell-Smith Namibia Etosha National Park 112 113 1 1 Omusati paratype 3747820728 MRAC M. E. Baddeley Namibia -18.066668 Andara-Kavango 1276 21.45 Okawango 112 113 7 6 1 Kavango West paratype 3747820724 1987-10-19 R, MRAC Jocque Namibia -17.866667 Choy 1277 23.366667 Sitwa 112 113 4 2 1 Zambezi paratype 3747820463 SMF Namibia -22.566668 Windhoek 1259 17.083334 112 113 RII 6618 1 1 Khomas paratype