Japalura batangensis Li, Deng, Wu & Wang, 2001 Diploderma batangense ( Li, Deng, Wu & Wang, 2001 ) J. Sich. Teach. Coll. Japalura bengalensis Annandale, 1912 Japalura variegata Gray, 1853 Japalura brevicauda Manthey, Denzer, Hou & Wang, 2012 Diploderma brevicauda ( Manthey, Denzer, Hou & Wang, 2012 ) Japalura brevipes Gressit, 1936 Diploderma brevipes ( Gressitt, 1936 ) J . brevipes J . swinhonis J . brevipes Japalura dasi ( Shah & Kästle, 2002 ) Oriotiaris dasi Shah & Kästle in Schleich & Kästle, 2002 Japalura drukdaypo Wang, Ren, Jiang, Zou, Wu, Che & Siler, 2019 Diploderma drukdaypo ( Wang, Ren, Jiang, Zou, Wu, Che & Siler, 2019 ) Japalura fasciata Mertens, 1926 Diploderma fasciatum ( Mertens, 1926 ) Japalura flaviceps Barbour & Dunn, 1919 Diploderma flaviceps ( Barbour & Dunn, 1919 ) Japalura hamptoni Smith, 1935 Diploderma hamptoni ( Smith, 1935 ) Japalura iadina Wang, Jiang, Siler & Che, 2016 Diploderma iadinum ( Wang, Jiang, Siler & Che, 2016 ) Japalura kumaonensis ( Annandale, 1907 ) Acanthosaura kumaonensis Annandale, 1907 Japalura laeviventris Wang, Jiang, Zou, Siler & Che, 2016 Diploderma laeviventre ( Wang, Jiang, Zou, Siler & Che, 2016 ) Japalura luei Ota, Chen & Shang, 1998 Diploderma luei ( Ota, Chen & Shang, 1998 ) Japalura major ( Jerdon, 1870 ) Oreocalotes major Jerdon, 1870 Japalura makii Ota, 1989 Diploderma makii (Ota, 1989) Japalura micangshanensis Song, 1987 Diploderma micangshanense ( Song, 1987 ) Japalura microlepis Jerdon, 1870 Japalura variegata Gray, 1853 Japalura mitsukurii Stejneger, 1898 Diploderma swinhonis ( Günther, 1864 ) Japalura ngoclinensis Ananjeva, Orlov & Nguyen, 2017 Diploderma ngoclinense ( Ananjeva, Orlov & Nguyen, 2017 ) Japalura otai Mahony, 2009 Cristidorsa otai ( Mahony, 2009 ) Japalura planidorsata Jerdon, 1870 Cristidorsa planidorsata ( Jerdon, 1870 ) Japalura polygonata donan Ota, 2003 Diploderma polygonatum donan ( Ota, 2003 ) Japalura polygonata ishigakiensis van Denburgh, 1912 Diploderma polygonatum ishigakiense (van Denburgh, 1912 ) Proc. Cal. Acad. Sci. Japalura polygonata miyakensis van Denburgh, 1912 Diploderma polygonatum ishigakiense (van Denburgh, 1912 ) Catalogue of type specimens of the agamid lizard genus Japalura s. l. (Squamata: Agamidae: Draconinae) Denzer, Wolfgang Manthey, Ulrich Zootaxa 2019 2019-05-28 4612 1 109 125 Annandale, 1905 Annandale 1905 [151,627,1700,1727] Reptilia Agamidae Japalura Animalia Squamata 4 113 Chordata species andersoniana    Holotype: ZSI 4104, male, Duffla Hills, Assam-Bhutan Frontier, coll. Col. Godwin-Austen    Paratype: NHMUK 1946.8.13.95 (original number NHMUK1906.8.10.5), male, Duffla Hills, Assam-Bhutan Frontier, coll. Col. Godwin-Austen  Type locality: Duffla [= Dafla], Assam/ Bhutanborder (fide Wermuth 1967)   Remarks: According to Das et al.(1998)ZSI 15755 (from ‘Nainital, 6,600 ft.’) is also listed as a type for  J. andersonianain the register, but this specimen was fixed by Annandale (1907)as the holotypeof  J. kumaonensis. As ZSI 15755 was collected by Annandale only in 1906 the specimen cannot possibly have been present in the collection when  J. andersonianawas first described.    Japalura batangensis Li, Deng, Wu & Wang, 2001=   Diploderma batangense( Li, Deng, Wu & Wang, 2001)J. Sich. Teach. Coll.22 (4): 329     Holotype: STCBa-98012, adult male, Wangjiao( 29°17.6′N, 99°5.2′E), Batang Country, Sichuan,  2,400 masl, coll. Wu Yi,  9 Aug. 1998    Allotype: STCBa-98004, adult female; Quxigong Village( 29°46.1′N, 99°3.8′E) Batang Country, Sichuan,  4,100 masl, coll. Wan Yu,  29 July 1998.     Paratypes: STCBa98010; Ba98014; Ba98017, three males;  STCBa98015; Ba98016; Ba98018, three females: collected from the same locality as holotype,  5–9 Aug. 1998.   Type locality: Wangjiao ( 29°17.6′N, 99°5.2′E), Batang Country, Sichuan, 2,400 masl, PR China    Japalura bengalensisAnnandale, 1912=   Japalura variegata Gray, 1853 Rec. Indian Mus. 8(2): 57, pl. V fig. 4     Holotype: ZSI 12564 female, Buxa, Jalpaiguri district, Bengalfrontier of Bhutan  Type locality: Buxa, Jalpaiguri, West Bengal, India   Remarks: In an earlier publication Annandale (1906: 288)referred to this specimen as a male  Japalura yunnanensis. We follow Smith (1935)and consider  J. bengalensisconspecific with  Japalura variegata.    Japalura brevicauda Manthey, Denzer, Hou & Wang, 2012=   Diploderma brevicauda( Manthey, Denzer, Hou & Wang, 2012) Zootaxa 3200: 29     Holotype: AMNH 19879(formerly a paratypeof  J. flaviceps), male, Snow Mt. Village,  2,740 masl( 27°05’N, 100°11’E), 19 kmnorth of Lijiang, coll. R. C. Andrewsand E. Heller,  Nov. 1916   Paratypes. AMNH 19878(formerly a paratypeof  J. flaviceps), female, temple on a mountain (  2,590 m) overlooking Lijiang, coll. R. C. Andrewsand E. Heller,  11. Oct. 1916;  NMW20853, juvenile, Lijiang, coll. von Handel- Mazzetti,  June 1915;  ZMB28932, female, eastern slopes of the Yulong Snow Mountains, near Lijiang(ca. 26°53’N, 100°11’E,>  2,500 masl.), coll. C. Schneider, between  July and Oct. 1914  Type locality: Eastern slopes of Yulong Snow Mountains, vicinity of Lijiang, Lijiang Prefecture, Yunnan, PR China    Japalura brevipesGressit, 1936=   Diploderma brevipes( Gressitt, 1936) Proc. Biol. Soc. Washington49: 117     Holotype: CAS71998, female, Bukai, near Horisha, Central Formosa(altitude  1,200 m), coll. J. L. Gressitt,  16 June 1934    Paratypes: MVZ19308, female;  MVZ 23357–58, one female and one juvenile; same locality as the holotype  Type locality: Bukai (Wu-chieh, Wujie), Yilan, Taiwan   Remarks: Gressitt’s (1936)assignment of paratypesis slightly ambiguous. He mentioned three paratypesalthough he actually lists four: “Three paratypes: two adult females (one in Mus. Vert. Zoöl.) and a juvenile, with the same data; a juvenile specimen from Sakahen, eastern Formosa, alt. 1,100 meters, July 13, 1934”. While it appears that only one specimen should be extant in the MVZ collection, their database lists two further specimens with the same collection data as the holotype(MVZ 23357–58, 23357 listed as paratopotype) and one specimen from Sakahen that has been collected by Gressitt on 13 July 1934(MVZ 23359). As neither the CAS online database nor the handwritten catalogue by Slevin lists further specimens collected by Gressitt we consider MVZ 23357–58 as paratypes. As Gressitt’s statement regarding the number of paratypesis ambiguous we do not list MVZ 23359 as paratypealthough it was part of the original series collected by Gressitt.    J. brevipeshas been treated by most authors as a junior synonym of  J. swinhonissince Okada (1937)considered them conspecific. Ota (1989b)revalidated  J. brevipesto full species rank. For this work he examined the holotype and a paratype( MVZ19308).    Japalura dasi( Shah & Kästle, 2002)see  Oriotiaris dasiShah & Kästle in Schleich & Kästle, 2002   Japalura drukdaypo Wang, Ren, Jiang, Zou, Wu, Che & Siler, 2019=   Diploderma drukdaypo( Wang, Ren, Jiang, Zou, Wu, Che & Siler, 2019) Zootaxa 4544: 508     Holotype: KIZ 027616, adult male, Chaya County, Chamdo, Tibet, China( 30.7294°N, 97.3808°E, elevation 3,310 m, WGS 84), coll. K. Wang,  19 June 2016    Paratypes: KIZ 027617–19,  KIZ 027629–30, adult females, coll. K. Wang& G. Nima,  19 June 2016;  KIZ 027628, adult male, coll. K. Wang& G. Nima,  21 June 2016; all with same locality data as holotype;  KIZ 016486, adult female, Kanuo, Chamdo, Tibet, China( 31.0433°N, 97.2239°E, elevation 3,245 m, WGS 84), coll. D. Zou& F. Yan,  13 May 2015  Type locality: Chaya County, Chamdo, Tibet, PR China   Remarks: Although in an earlier paper Wang et al (2018)resurrected the genus  Diplodermatheir new species ( Wang et al.2019) is described as  Japalura s. l.and the previous taxonomic paper is not mentioned at all. Based on their description and apparent close relationship to  flavicepswe assign the species  drukdaypoto the genus  Diploderma. It should also be noted that within the publication the year of description is given as 2016 although the paper was only published in 2019.    Japalura fasciata Mertens, 1926=   Diploderma fasciatum( Mertens, 1926) Senckenbergiana 8 (3–4): 146     Holotype: SMF 9871, male, H. Fruhstorfer  Type locality: not known.   Remarks: The typespecimen was purchased by the Senckenberg Museum from H. Fruhstorfer, Tonkin(N Vietnam) [Boettger (1904)]. This apparently led Smith (1935)to state “ Tongking” as the typelocality.    Japalura flaviceps Barbour & Dunn, 1919=   Diploderma flaviceps( Barbour & Dunn, 1919) Proc. New Englandzool. Club 7: 16     Holotype: MCZR-12469, adult male, sandy shores of the Tung Riverin western Szechuan, coll. W. R. Zappey,  7. Feb 1908.    Paratypes: MCZ R12470–73, all listed as skeletons, same data as the holotype;  Type locality: Tung River (= Dadu He), Western Szechuan (= Sichuan), China   Remarks: Barbour & Dunn (1919)listed three further paratypesfrom the Snow Mountains near Lichiangfu in northern Yunnan, not far from the Szechuan frontier AMNH collection. Of these AMNH 19879 was later designated as the holotypefor  Japalura brevicauda Manthey, Denzer, Hou & Wang, 2012and AMNH 19878 as a paratypefor that species. The whereabouts of the third specimen is unknown.    Japalura hamptoni Smith, 1935=   Diploderma hamptoni( Smith, 1935) Fauna Brit. India: 175     Holotype: 1946.8.14.1 (formerly NHMUK1908.9.18.1), male, Mogok, Upper Burma, coll. H. Hampton, 1908  Type locality: Mogok, Pyin Oo Lwin District, Mandalay Region, Myanmar    Japalura iadina Wang, Jiang, Siler & Che, 2016=   Diploderma iadinum( Wang, Jiang, Siler & Che, 2016) Zool. Res. 37(1): 51     Holotype: KIZ 019321, adult male, Lancang Valleyat Ninong, Deqin, northwest Yunnan, China( 28.370255° N, 98.865287° E),  2,062 masl, coll. K. Wang,  27 May 2015  Allotopotype:  KIZ09398, adult female, coll. D.-H. Zou, same data as the holotype   Paratopotypes: KIZ 09401–03, 019322, 019323, 019325–28, all adult males, coll. K. Wang, K. Jiang and D.- H. Zou   Type locality: Lancang Valley at Ninong, Deqin, northwest Yunnan, China( 28.370255° N, 98.865287° E)     Japalura kumaonensis( Annandale, 1907)see  Acanthosaura kumaonensis Annandale, 1907   Japalura laeviventris Wang, Jiang, Zou, Siler & Che, 2016=   Diploderma laeviventre( Wang, Jiang, Zou, Siler & Che, 2016) Zool. Res. 37(1): 42     Holotype: KIZ 014038, adult male, near the Nujiang Bridge in the upper Nujiang Valleyat Baxoi, Qamdo, eastern Tibet, coll. K. Jiang,  3 July 2013  Paratopotypes: KIZ014037, adult male; KIZ0141041.043, three adult females; all coll. K. Jiang, K. Wang & Y.-Q. Sun  Type locality: near the Nujiang Bridge in the upper Nujiang Valley at Baxoi (=Basu), Qamdo (=Changdu), eastern Tibet(= Xizang), China( 30.10034° N, 97.22787° E, 2739 masl)    Japalura luei Ota, Chen & Shang, 1998=   Diploderma luei( Ota, Chen & Shang, 1998) Copeia 1998 (3): 649     Holotype: NTNUB 2235, adult male, Mt. Taipingshan, Nanao Hsiang, Ilan Pref., Taiwancoll. S.-L. Chen,  5 Sept. 1995    Paratypes: KUZ R32929–930, 32940, 32953,  NMNS 2816, five adult males;  KUZR32939, one immature male, all having the same data as the holotype;  TM RL0425, adult female, same locality as the holotype, coll. G. Shang,  8 July 1996  Type locality: Mt. Taipingshan, Nanao Hsiang, Ilan Pref., Taiwan( 24°31’N, 121°30’E, 1950–2000 masl)     Japalura major( Jerdon, 1870)see  Oreocalotes major Jerdon, 1870   Japalura makiiOta, 1989=   Diploderma makii(Ota, 1989)  Copeia 1989 (3): 570     Holotype: OMNH(Osaka) R2565, adult male, Chitou, Nantou Pref., Taiwan, ca.  1,500masl, coll. H. Ota,  13 Aug 1986    Paratypes: KUZ 7877, 7940, 7975, 8954;  AMNHR-  130926;  NHMUK 1987.2183,  MCZ 172743,  USNM 281851, eight males;  OMNH R2566,  KUZ 7857, two females; all having same collection data as holotype  Type locality: Chitou, Nantou Pref., TaiwanProvince, China, about 1500 masl, 23° 42’ N, 120° 47’ E    Japalura micangshanensis Song, 1987=   Diploderma micangshanense( Song, 1987) Acta Herp. Sinica 6 (1): 59 (in Chinese), 64 (in English)     Holotype: SIZ82158 ( Shaanxi Institute of Zoology) adult female, Qingmuchuan, Ningqiang County, Shaanxi, alt.  700 m, collected by the author  16. June 1982    Allotype: SIZ82132, adult male, alt.  750 m,  8. June, 1982     Paratypes: 1 subadultmale ( SIZ82119)and  15 adultfemales ( SIZ82092, 82120, 82125–6, 82135–8, 82152, 82159–60, 82186, 82193, 82254–5) from the same locality,  25. May to 28. June 1982   Type locality: Qingmuchuan, Ningqiang County., Shaanxi Province, China, 700 masl   Remarks: The catalogue numbers for the allotypeand paratypesare only mentioned in the Chinese part of the description.    Japalura microlepis Jerdon, 1870=   Japalura variegata Gray, 1853 Proc. Asiat. Soc. Bengal, 1870: 76     Holotype: NHMUK 1946.8.14.10 (original number NHMUK1870.11.29.44), female (Boulenger: 309, specimen g), Sikkim, pres. T. C. Jerdon  Type locality: Sikkim   Remarks: Jerdon (1870)only had a single specimen for the original description which therefore becomes the holotype. This specimen corresponds to specimen g in Boulenger (1885: 309). See also comments under  J. variegata. We follow Boulenger (1885)and consider  Japalura microlepisconspecific with  J. variegata.    Japalura mitsukurii Stejneger, 1898=   Diploderma swinhonis( Günther, 1864) J.  Coll. Sci. Univ. Tokyo12: 218     Holotype: USNM 36505, male, Botel Tobago Island, coll. T. Tada, 1896/97    Paratype: USMN 36506, female;  CAS27257 (formerly cotype e = presumably Science College Tokyo SCT 24e), male, Botel Tobago Island; SCT 24a, male; SCT 24b, female; SCT 24c, male; SCT 24d, male; all coll. T. Tada, 1896/97  Type locality: Botel Tobago Island (= Lanyu Island)   Remarks: Alternatively TIU ( TokyoImperial University; now University of Tokyo, Science College Museum, SCT) is used as the collection acronym (see Frost 2019).  Stejneger (1898)stated that the collection consisted of eight specimens. In his Herpetology of Japan(1907: 191) he only listed seven specimens, the ones listed above. He designated USNM 36505 as the type, which we understand is actually the holotype. Consequently, the remaining specimens are considered paratypes. Stejneger (1907)erroneously stated that the collection date was May 1899, i. e. after the original publication. Van Denburgh (1912)synonymized  J. mitsukuriiwith  J. swinhonisand treated it as conspecific with or a subspecies of the latter. Most subsequent authors followed this classification until Ota (1988b)initially showed that  J. mitsukuriirepresents a valid species and that  J. swinhonis formosensishad to be considered conspecific with  J. mitsukurii. In a later publication Ota (1991a)revised this classification and stated that  J. mitsukurii, and consequently  J. swinhonis formosensis, constitutes a junior synonym of  J. swinhonis(=  Diploderma swinhonis).    Japalura ngoclinensis Ananjeva, Orlov & Nguyen, 2017=   Diploderma ngoclinense( Ananjeva, Orlov & Nguyen, 2017) AHR 8(1): 15     Holotype: ZISP 26230, adult female, above Mang Xang Village, Ngoc Linh Mountain, Dac Glei District, Kon Tum Province, Vietnam( 15°05’ N 107°57’ E,  1650 masl), coll. A. V. Abramov,  25 Mar. 2004    Paratypes: ZISP 26231, adult female and VNMN 3110, subadult female, same data as the holotype  Type locality: above Mang Xang Village, Ngoc Linh Mountain, Dac Glei District, Kon Tum Province, Vietnam( 15°05’ N 107°57’ E, 1,650 masl)   Remarks: Ananjeva et al (2017)used two different museum acronyms (VNMH and VNMN) for the Vietnamese collection where one of the paratypeshas been deposited. Hitherto this collection has not been listed in previous herpetological publications. As the full name of the collection is given as VietnamNational Museum of Nature under author affiliations we consider VNMN as the correct acronym.  Wang et al.(2018)treat this species as conspecific with  Diploderma splendidum( Barbour & Dunn, 1919)    Japalura otai Mahony, 2009=   Cristidorsa otai( Mahony, 2009) Zootaxa 2212: 43     Holotype: ZSI 25217, male, “ Sairep, Lunglet District, Mizoram”, coll. S.S. Saha,  19 Apr. 1995    Paratypes: ZSI 25215 and 25216, two females, locality data as for holotype, coll. S. S. Saha,  16 and 19 April 1995  Type locality: Sairep ( 22°49’0 N, 92°49’0 E), Lunglei District, central Mizoram, northeast India    Japalura planidorsata Jerdon, 1870=   Cristidorsa planidorsata( Jerdon, 1870) Proc. Asiat. Soc. Bengal, March 1870: 76     Syntypes: 1946.8.30.86–87 (formerly NHMUK(18)70.11.29.20–21), Khasi Hills, pres. T. C. Jerdon   Type locality: Khasi Hills   Remarks: As Jerdon (1870)had two specimens but as he did not base his original description on any particular specimen we consider them as syntypes.    Japalura polygonata donan Ota, 2003=   Diploderma polygonatum donan( Ota, 2003) Curr. Herp. 22 (2): 63, fig.2     Holotype: KUZ R234, adult male, Mt. Urabudake( 24°27’N, 123°00’E,  130 masl), Yonagunijima Island, coll. H. Ota,  17 Aug. 1981    Paratypes: KUZ R102, adult female, Higawa, Yonagunijima Island, coll. H. Ota,  30 Mar. 1981;  KUZ R231, 238, two adult females, same data as the holotype;  KUZ R415, adult female, between Higawaand Arakawabana, Yonagunijima Island, coll. H. Ota,  7 Mar. 1982;  KUZ R971, 972, 975, 979, 997, four adult males and one adult female, between Mt Urabudakeand Sonai, Yonagunijima Island, coll. T. Hayashi, K. Kawamura, S. Kobayashi, A. Mori, and H. Ota,  26 Mar. 1983;  KUZ R1382, 1420, two adult females, Kubura, Yonagunijima Island, coll. H. Ota,  27 July 1983;  KUZR  2000–2003, 2164, 2876, 3261-3263, 3265, 3269, 3270, 3281-3283, 3289, 3291-3294, 3982, 4517, 4518, 22 adultmales, two adult females, and three juveniles, Mt. Urabudake, coll. H. Ota,  15 July 1984; KUZ R28117, 28118, two adult females, Kubura, coll. H. Ota,  25 June 1993; KUZ R56152-56156, 56158-56167, 15 adultmales, Mt. Urabudake, coll. H. Ota,  25 March 2003  Typelocality: Mt. Urabudake( 24°27’N, 123°00’E,  130 masl), Yonagunijima Island, Japan    Japalura polygonata ishigakiensisvan Denburgh, 1912 =   Diploderma polygonatum ishigakiense(van Denburgh, 1912)Proc. Cal. Acad. Sci., 4 (3): 211     Holotype: CAS21354, Ishigaki Shima, Loo Choo Isls. Japan, coll. V. Kühne,  25 May – 2 June 1910    Paratypes: CAS 21355–361, Ishigaki Shima, Loo Choo Islands. Japan, coll. V. Kühne,  25 May – 2 June 1910;  CAS 21362–377, Funaoke, Iriomote Shima, Loo Choo Islands. Japan, coll. V. Kühne,  24–25 May 1910.  Type locality: Ishigaki Shima and Iriomote Shima, Ryukyu Islands, Japan   Remarks: The original description was first published by van Denburgh in a privately printed paper published also in 1912: “Advance diagnoses of new reptiles and amphibians from the Loo Choo Islands and Formosa”.    Japalura polygonata miyakensisvan Denburgh, 1912=   Diploderma polygonatum ishigakiense(van Denburgh, 1912) Proc. Cal. Acad. Sci., 4(3): 211     Holotype: CAS21353, Miyako Shima, Loo Choo Islands, coll. V. Kühne,  23–24 May 1910    Paratypes: CAS 21346–352, Miyako Shima, Loo Choo Islands. Japan, coll. V. Kühne,  23–24 May 1910  Type locality: Miyako Shima, Ryukyu Islands, Japan   Remarks: The original description was first published by van Denburgh in a privately printed paper published also in 1912: “Advance diagnoses of new reptiles and amphibians from the Loo Choo Islands and Formosa”. In his guide book about Japanese amphibians and reptiles Matsumoto (1979)did not list  Japalura polygonata miyakensis. Although Matsumoto (l. c.) did not explicitly synonymize the subspecies he treated poulations from Miyako Shima as identical with  Japalura polygonata ishigakiensis. Apart from Ota (1991b, 2003) there appears to be no further mention of Matsumoto‘s publication in herpetological literature.   Japalura polygonata xanthostomaOta, 1991=   Diploderma polygonatum xanthostoma(Ota, 1991)Herpetologica47: 288, fig.4     Holotype: KUZ R9855, adult male, Waishuangchi, Taipei Prefecture, Taiwan(25’06’ N, 121’33’ E), elevation approximately  100 m, coll. H. Ota,  13 June 1988    Paratypes: KUZR  1600–1602, 1609–1610, all coll. T. Hikida, Aug 1976;  KUZ R1649, 1657,  2058–2059all coll. T. Hikida, April 1977;  KUZ R2715, 2809, 29192923, 2933,  2978–2979, 3036, 3056, 3219, 3964, 4360–4361, all coll. H. Ota, Oct 1984;  KUZ R6707–6710, 7519–7534, 7537–7545, 7846, 7851–7855, 7861–7871, 7880–7882, 7884, 7901, 7922, 7941–7942, 7952, 7961, 7973, 7991–7992, 8009–8010, 8019–8020, 8066–8067, 8099, 8420, 8446–8452, all coll. H. Ota,  July and Aug. 1986;  KUZ R9775–9780, 9842–9854, 9856–9861, 13010–13019, all coll. H. Ota,  May and June 1988;  101 adultmales, 10 immature males, 30 adultfemales and seven immature females; same locality data as the holotype;  KUZ R8424–8431, Tansui, Taipei Pref., coll. H. Ota,  Aug 1986;  KUZ R6914–6915, 6943–6944, 6947, Tungao, Ilan Pref., coll. H. Ota,  July 1986;  AMNHR-77093, Pei-tou, Formosa, coll.  12 Sep 1956  Type locality: Waishu-angchi, Taipei, Taiwan 2244393302 NHMUK Duffla Hills & Assam-Bhutan Frontier & Col. Godwin-Austen 4 113 NHMUK 1946.8 1 paratype 2244393325 1998-08-09 STC Wu Yi 2400 29.293333 Batang Country 122 99.08667 Wangjiao 5 114 1 Sichuan holotype 2244393283 1998-07-29 STC Wan Yu 4100 29.768333 Batang Country 122 99.06333 Quxigong Village 5 114 1 Sichuan allotype 2244393307 [199,867,441,467] STC 5 114 1 paratype 2244393312 1998-08-05 1998-08-09 1998-08-05 STC Wu Yi 2400 29.293333 Batang Country 122 99.08667 Wangjiao 5 114 1 Sichuan paratype 2244393280 [199,1110,656,683] Bhutan Bengal Buxa Jalpaiguri district 5 114 1 holotype 2244393276 1916-11 AMNH R. C. Andrews & E. Heller 2740 27.083334 Snow Mt. Village 1239 100.183334 Lijiang 5 114 AMNH 19879 2 holotype 2244393260 1916-10-11 AMNH Lijiang & R. C. Andrews & E. Heller 2590 5 114 AMNH 19878 2 paratype 2244393308 1915-06 NMW Lijiang Mazzetti 5 114 1 paratype 2244393304 1914-07 1914-10-31 1914-07 ZMB C. Schneider 2500 26.883333 Yulong Snow Mountains 1239 100.183334 Lijiang 5 114 1 paratype 2244393259 1934-06-16 CAS J. L. Gressitt Taiwan Central 1200 Bukai Horisha 5 114 1 holotype 2244393310 [199,567,1413,1439] MVZ 5 114 1 paratype 2244393303 [578,1429,1412,1439] 1934-06-16 MVZ J. L. Gressitt Taiwan Central 1200 Bukai Horisha 5 114 MVZ 23357-58 1 paratype 2244393318 [151,483,1881,1907] MVZ 5 114 1 paratype 2244393213 2016-06-19 KIZ, WGS K. Wang China Chaya County 3310 30.7294 Chamdo 6 97.3808 6 115 KIZ 027616, WGS 84 1 Xizang holotype 2244393199 [199,525,333,359] KIZ 6 115 KIZ 027617-19 1 paratype 2244393201 [535,1376,333,359] 2016-06-19 KIZ K. Wang & G. Nima 6 115 KIZ 027629-30 1 paratype 2244393226 2016-06-21 KIZ K. Wang & G. Nima 6 115 KIZ 027628 1 paratype 2244393231 [847,1281,369,395] 2016-06-21 KIZ K. Wang & G. Nima 6 115 1 holotype 2244393209 2015-05-13 KIZ, WGS D. Zou & F. Yan China 3245 31.0433 Kanuo 6 97.2239 Chamdo 6 115 KIZ 016486, WGS 84 1 Xizang paratype 2244393203 [199,685,797,823] SMF H. Fruhstorfer 6 115 SMF 9871 1 holotype 2244393205 1908-02-07 MCZ W. R. Zappey Tung River western Szechuan 6 115 1 holotype 2244393198 [199,1089,1121,1147] 1908-02-07 MCZ W. R. Zappey Tung River western Szechuan 6 115 R12470-73 1 paratype 2244393207 NHMUK H. Hampton Myanmar Burma 6 115 1 holotype 2244393229 2015-05-27 KIZ K. Wang China Deqin 2062 28.370255 Lancang Valley 1 98.86529 Ninong 6 115 KIZ 019321 1 Yunnan holotype 2244393216 [368,1132,1733,1759] 2015-05-27 KIZ D. - H. Zou China Deqin 2062 28.370255 Lancang Valley 1 98.86529 Ninong 6 115 1 Yunnan 2244393224 2013-07-03 KIZ K. Jiang upper Nujiang Valley 7 116 KIZ 014038 1 Xizang holotype 2244393206 1995-09-05 NTNUB S. - L. Chen Taiwan Mt. Taipingshan 7 116 NTNUB 2235 1 holotype 2244393210 [199,729,584,611] KUZ 7 116 R32929-930, 32940, 32953 1 paratype 2244393275 [739,1092,585,611] NMNS 7 116 NMNS 2816 1 paratype 2244393200 1995-09-05 KUZ S. - L. Chen Taiwan Mt. Taipingshan 7 116 1 paratype 2244393273 1996-07-08 TM G. Shang Taiwan Mt. Taipingshan 7 116 RL0425 1 paratype 2244393270 1986-08-13 OMNH H. Ota Taiwan 1500 Chitou 7 116 1 Nantou holotype 2244393266 [199,687,980,1007] KUZ 7 116 KUZ 7877, 7940, 7975, 8954 1 paratype 2244393278 [701,913,981,1007] 26-09-13 AMNH 7 116 1 paratype 2244393313 [928,1171,981,1007] NHMUK 7 116 NHMUK 1987.2183 1 paratype 2244393223 [1182,1344,981,1007] MCZ 7 116 MCZ 172743 1 paratype 2244393233 USNM 7 116 USNM 281851 1 paratype 2244393214 [392,566,1016,1043] OMNH 7 116 R2566 1 paratype 2244393298 [574,697,1016,1043] KUZ 7 116 KUZ 7857 1 paratype 2244393289 [708,1341,1017,1043] 1986-08-13 OMNH H. Ota Taiwan 1500 Chitou 7 116 1 Nantou paratype 2244393271 1982-06-16 SIZ Ningqiang County 700 Qingmuchuan Institute of Zoology 7 116 1 Shaanxi holotype 2244393326 [199,856,1268,1295] 8. June SIZ 750 7 116 1 allotype 2244393279 [199,701,1305,1332] SIZ 7 116 1 1 paratype 2244393316 1982-05-25 1982-06-28 1982-05-25 SIZ 7 116 82120, 82125-6, 82135-8, 82152, 82159-60, 82186, 82193, 82254-5 15 15 paratype 2244393265 NHMUK T. C. Jerdon Sikkim 7 116 NHMUK 1946.8 1 Sikkim holotype 2244393285 [176,427,1873,1899] Coll. Sci. Univ. Tokyo 7 116 1 Tokyo 2244393274 [199,1047,1908,1935] USNM T. Tada Botel Tobago Island 7 116 USNM 36505 1 holotype 2244393277 [199,579,1944,1971] USMN 7 116 USMN 36506 1 paratype 2244393262 CAS Science College 7 116 SCT 24 1 Tokyo paratype 2244393230 SCT T. Tada Botel Tobago Island 7 116 1 Tokyo paratype 2244393288 2004-03-25 ZISP A. V. Abramov Vietnam Dac Glei District 1650 15.083333 Mang Xang Village 1286 107.95 Ngoc Linh Mountain 8 117 ZISP 26230 1 Kon Tum holotype 2244393296 [199,760,801,827] VNMN 8 117 ZISP 26231 1 paratype 2244393295 [767,1316,801,827] 2004-03-25 VNMN A. V. Abramov Vietnam Dac Glei District 1650 15.083333 Mang Xang Village 1286 107.95 Ngoc Linh Mountain 8 117 1 Kon Tum paratype 2244393315 [199,1265,1196,1223] 1995-04-19 S. S. Saha Lunglet District Sairep 8 117 1 Mizoram holotype 2244393284 1995-04-16 1995-04-19 1995-04-16 S. S. Saha Lunglet District Sairep 8 117 1 Mizoram paratype 2244393322 [199,1333,1448,1475] NHMUK Khasi Hills & T. C. Jerdon 8 117 1 syntype 2244393306 1981-08-17 KUZ H. Ota 130 24.45 Mt. Urabudake 1251 123.0 Yonagunijima Island 8 117 R234 1 holotype 2244393321 [199,1290,1773,1799] 1981-03-30 KUZ H. Ota Higawa Yonagunijima Island 8 117 R102 1 paratype 2244393294 1981-08-17 KUZ H. Ota 130 24.45 Mt. Urabudake 1251 123.0 Yonagunijima Island 8 117 R231, 238 1 paratype 2244393264 1982-03-07 KUZ H. Ota Yonagunijima Island Higawa Arakawabana 8 117 R415 1 paratype 2244393317 1983-03-26 KUZ T. Hayashi & K. Kawamura & S. Kobayashi & A. Mori & H. Ota Yonagunijima Island Mt Urabudake Sonai 8 117 R971, 972, 975, 979, 997 1 paratype 2244393251 1983-07-27 KUZ H. Ota Kubura Yonagunijima Island 8 117 R1382, 1420 1 paratype 2244393263 [313,526,1953,1979] 2000 2003 2000 KUZ 8 117 1 paratype 2244393300 1984-07-15 H. Ota Mt. Urabudake 8 117 22 22 paratype 2244393305 1993-06-25 2003-03-25 1993-06-25 KUZ Kubura & H. Ota Japan Yonagunijima Island 130 24.45 Mt. Urabudake 1251 123.0 Mt. Urabudake 9 118 R56152-56156, 56158-56167 15 15 2244393286 [199,1326,404,431] 1910-05-25 1910-06-02 1910-05-25 CAS V. Kuhne Japan Japan 9 118 1 holotype 2244393292 [199,1431,440,467] 1910-05-25 1910-06-02 1910-05-25 CAS V. Kuhne Japan Loo Choo Islands 9 118 CAS 21355-361 1 paratype 2244393311 [151,1312,476,503] 1910-05-24 1910-05-25 1910-05-24 CAS V. Kuhne Japan Loo Choo Islands 9 118 CAS 21362-377 1 paratype 2244393267 [199,1227,764,791] 1910-05-23 1910-05-24 1910-05-23 CAS V. Kuhne Loo Choo Islands 9 118 1 holotype 2244393256 [199,1372,800,827] 1910-05-23 1910-05-24 1910-05-23 CAS V. Kuhne Japan Loo Choo Islands 9 118 CAS 21346-352 1 paratype 2244393297 1988-06-13 KUZ H. Ota Taiwan 100 Waishuangchi 9 118 R9855 1 Taipei holotype 2244393250 [199,690,1269,1295] 1600 - 1602, 1609 - 1610 KUZ 9 118 1 paratype 2244393249 [1046,1400,1268,1295] 2058 2059 2058 KUZ 9 118 R1649, 1657 1 paratype 2244393320 [462,1011,1304,1331] 2978 2979 2978 KUZ 9 118 R2715, 2809, 2919 1 paratype 2244393254 1986-07 1986-08-31 1986-07 KUZ H. Ota 9 118 R6707-6710, 7519-7534, 7537-7545, 7846, 7851-7855, 7861-7871, 7880-7882, 7884, 7901, 7922, 7941-7942, 7952, 7961, 7973, 7991-7992, 8009-8010, 8019-8020, 8066-8067, 8099, 8420, 8446-8452 1 paratype 2244393323 1988-05 1988-06-31 1988-05 KUZ H. Ota 9 118 R9775-9780, 9842-9854, 9856-9861, 13010-13019 1 paratype 2244393248 1988-05 1988-06-31 1988-05 KUZ H. Ota 9 118 131 131 holotype 2244393252 [637,1365,1485,1511] 1986-08 KUZ H. Ota Tansui 9 118 R8424-8431 1 Taipei paratype 2244393247 1986-07 KUZ H. Ota Ilan Pref. Tungao 9 118 R6914-6915, 6943-6944, 6947 1 Taipei paratype 2244393255 1956-09-12 AMNH Taiwan Pei-tou 9 118 1 paratype