Coluber sansaniensis Lartet, 1851: 40
Pylmophis sansaniensis
Rochebrune 1880: 282
sansaniensis
Lydekker 1888: 251
Natrix sansaniensis
Rage 1981: 538-540
Augé & Rage 2000: 288- 299
Ivanov 2002: 523-525
Natrix
N . sansaniensis
Szyndlar & Schleich 1993: 17-20
Natrix sansaniensis
Natrix
N . sansaniensis
Natrix
N . sansaniensis
Natrix
N . sansaniensis
Natrix
N . sansaniensis
N . merkurensis
N . sansaniensis
Natrix merkurensis
Amphibians and squamate reptiles from the late early Miocene (MN 4) of Béon 1 (Montréal-du-Gers, southwestern France)
Rage, Jean-Claude
Bailon, Salvador
Geodiversitas
2005
27
3
413
441
Ivanov 2002: 523-525
(Lartet, 1851)
Lartet
1851
[758,1127,1292,1317]
Reptilia
Natricidae
Natrix
Animalia
Squamata
16
429
Chordata
species
sansaniensis
pars Coluber sansaniensis Lartet, 1851: 40. Pylmophis sansaniensis– Rochebrune 1880: 282, 283, pl. 12, fig. 11. Pilemophis sansaniensis– Lydekker 1888: 251. Natrix sansaniensis– Rage 1981: 538-540, fig. 1A; 1984b: 48, 49, fig. 3A. — Augé & Rage 2000: 288- 299, figs 21, 22. — Ivanov 2002: 523-525, fig. 7. pars Natrixaff. N. sansaniensis– Szyndlar & Schleich 1993: 17-20, fig. 4A-H.
MATERIAL EXAMINED. — 144 trunk vertebrae (Béon 2004 LT 65-68; Béon 91 E3 SN 1).
COMMENTS
Natrix sansaniensiswas revised and redescribed by Rage (1981)and Augé & Rage (2000). The species appears as a typical member of Natrix. It is mainly characterized by its high neural spines and the peculiar morphology of its hypapophyses (unusual combination of a curved anteroventral border and a pointed tip). N. sansaniensiswas reported from Sansan (MN 6, France), the typelocality, and Merkur-North (MN 3, Czech Republic) ( Augé & Rage 2000; Ivanov 2002). The species is perhaps present in the early middle Miocene of Vieux-Collonges ( Natrixaff. N. sansaniensis; Ivanov 2000). Natrixcf. N. sansaniensiswas reported from La Grive M (MN 7+8; France) by Ivanov (2002)and from Mátraszölös 1 (MN 6 or 7+8; Hungary) by Gál et al. (1999). Moreover, numerous vertebrae from Petersbuch 2 (MN 4; Germany) were referred to as Natrixaff. N. sansaniensisby Szyndlar & Schleich (1993); but, after the description of the species N. merkurensisby Ivanov (2002)it may be supposed that a part of the vertebrae assigned to the fossil from Petersbuch 2 really belongs to N. sansaniensis(see below: Natrix merkurensis).