Amphibians and squamate reptiles from the late early Miocene (MN 4) of Béon 1 (Montréal-du-Gers, southwestern France) Rage, Jean-Claude Bailon, Salvador Geodiversitas 2005 27 3 413 441 [332,427,1004,1029] Amphibia Ranidae Rana Animalia Anura 7 420 Chordata species undetermined  MATERIAL EXAMINED. — 2 angulosplenials (Béon 2004 LT 21), 6 presacral vertebrae (Béon 2004 LT 22), 6 sacral vertebrae (Béon 2004 LT 23), 6 humeri (Béon 2004 LT 24-26), and 1 ilium (Béon 2004 LT 27).  DESCRIPTION The specimens clearly show the characteristics of the genus  Rana. More specifically, several features are typical of species belonging to the “green frogs group”, or “water frogs” (i.e. synklepton  Rana esculentasensu Dubois & Günther 1982). The processus coronoideusof the angulosplenial forms a well developed vertical lamina whose dorsal edge is convex; its extent is similar to that of green frogs. Moreover, the lateral surface of the angulosplenial that is located below the sulcus Meckeli, at the level of the processus coronoideus, is nearly vertical and limited ventrally by an elongate external mandibular ridge as in green frogs ( Bailon 1999). The vertebrae and humeri show all of the characteristics of the genus  Rana. All available humeri are small, they likely belonged to juvenile individuals ( Fig. 3A, B). The only available ilium is damaged ( Fig. 3C), but the depth of the supracetabular fossa and the thickness of the ilio-ischiatic face also point to green frogs ( Bailon 1991, 1999).  COMMENTS Green frogs are comparatively frequent in Tertiary localities. The earliest fossils were report- ed from the early Oligocene ( Sanchiz et al. 1993), perhaps the late Eocene ( Rage 1984a; Holman & Harrison 1999). Several species from the Miocene of Europe likely belong to this assemblage ( Rage & Roček 2003), but Sanchiz (1998b)regarded all of them as nomina dubia. Today, the green frogs group includes true species and hybridogenetic “species” that form a complex assemblage. Dubois & Günther (1982)named this assemblage “Synklepton  Rana esculenta”. The morphology of these frogs is very homogeneous and the osteology of several recently recognized living species is unknown. Consequently, it is practically impossible to distinguish extinct species within this complex.