Two new species of Megarthrus (Coleoptera, Staphylinidae, Proteininae) from Mexican Cloud Forests Rodríguez, William David Navarrete-Heredia, José L. Arriaga-Varela, Emmanuel Cuccodoro, Giulio Zootaxa 2020 2020-05-15 4778 3 571 583 8Y69S Rodriguez, Navarrete-Heredia & Arriaga-Varela Rodriguez, Navarrete-Heredia & Arriaga-Varela 2020 [151,1027,1592,1620] Insecta Staphylinidae Megarthrus Animalia Coleoptera 7 578 Arthropoda species cavianae sp. nov.  ( Figs 27-39) http://zoobank.org:act: F4814244-2EA7-493D-8086-01CC754585C0   Type material.( 3♂, 2♀).  Holotype  ♂: labeled “ MÉXICO: Veracruz, San Andrés Tlalnelhuayocan, Piedra Parada, 19°30’58.6’’N 97°00’34.4’’W, 1522 msnm, Fragmento de Bosque Mesófilo de Montañaripario, Cernido de Hojarasca húmeda, extraída en Winkler,  5.V.2013, F. Alvarado, E. Arriaga, R. Chaverocols”, in CZUG.   Paratypes(4): labeled “ MÉXICO: Veracruz, Xico, Matlalapa, 19°26’24.8»N 97°02’37.2»W, 1532 msnm. Fragmento de Bosque Mesófilo de Montañaripario, Cernido de Hojarasca húmeda, extraída en embudo de Berlesse,  20.VI.2012. F. Escobar, F. Alvarado, R. Chaverocols”,  1♀in CZUG,  1♂in IEXA, and 1 ♂, 1♀in MHNG.   Description.Similar to  M. chiapasfrom which it differs as follows: Habitus as in Fig. 29. Combined length of pronotum and elytra = 1.0 - 1.4 mm; maximal pronotal width = 0.7 - 1.1 mm. Antennae as in Fig. 36. Elytra with humeral callus strongly convex; disc with humeral callus strongly convex, two conspicuous swellings similar to humeral callus along anterior and posterior portions of suture, moderately depressed posteriorly along lateral edge. Male: Frontoclypeal area not modified. Protarsomere 1 lacking ventral patch of modified adhesive setae. Mesofemora ( Fig. 33) as long as metafemora. Mesotibiae ( Fig. 34) shorter than metatibiae ( Fig. 35), the latter slightly compressed subapically. Peg-like setae present only on mesotrochanters ( Fig. 33; arranged in a single row), on distal two thirds of mesotibiae ( Fig. 34; grouped in a field) and on distal half of metatibiae ( Fig. 35; grouped in a field). Apical segments of abdomen similar to those in Figs 10-15 & 18; tergite VIII as in Figs 38, with apex sinuate in dorsal view. Aedeagus as in Figs 32 & 37, with dorsal valve triangular.   FIGURES 29–31.  Megarthrus cavianae  sp. nov., female: habitus in dorsal view (29); abdominal sternite VIII in ventral views (30); apical segments of abdomen in lateral view (31). Scale bars; vertical = 0.5 mm (29); 25–26. horizontal = 0.2 mm (30, 31).   FIGURES 32–38.  Megarthrus cavianae  sp. nov., male: aedeagus in ventral (32) and lateral (37) views; mesofemur and mes- otrochanter (33); mesotibia (34); metatibia (35); antenna (36); abdominal tergite VIII in dorsal view (38). Scale bars = 0.2 mm: 32–35, 37, 38 (horizontal); 36 (vertical).  Female: Abdominal tergite forming a medioapical projection, with dorsal outline subangulate subapically in lateral view VIII ( Fig. 31). Sternite VIII as in Fig. 30. Genital segments similar to those in Figs 19-21; gonocoxal plate without mediodorsal ridge.  Comparisons and diagnostic notes.  Megarthrus cavianaeRodríguez, Navarrete-Heredia & Arriaga-Varela  sp. nov.is similar to  M. chiapasCuccodoro  sp. nov., from which it differs by its more pronounced elytral relief, as well as several other features detailed in Comparisons and diagnostic notesunder  M. chiapas.   Distribution and natural history.  Megarthrus cavianaeRodríguez, Navarrete-Heredia & Arriaga-Varela  sp. nov.is apparently restricted to Mountain Cloud Forests in the south of the Transmexican Volcanic Belt ( Fig. 39). The specimens were collected from sifted leaf litter in riparian remnants of cloud forests at an elevation of approximately 1500 m.   Etymology.The specific name is a composite word in honor to Carmen Moreno Jiménez, Virginiade Jesús Rodríguez, Ana Cricelia Rodríguez, and Ana María Jiménez, in recognition of their unconditional support of the first author. 2620268301 2013-05-05 CZUG F. Alvarado & E. Arriaga & R. Chavero Mexico 19.516277 Winkler 1 -97.00955 Fragmento de Bosque Mesofilo de Montana 7 578 1 1 Veracruz de Ignacio de la Llave holotype 2620268303 2012-06-20 F. Escobar & F. Alvarado & R. Chavero Mexico 19.440222 Berlesse 1 -97.04366 Fragmento de Bosque Mesofilo de Montana 7 578 1 Veracruz de Ignacio de la Llave paratype 2620268306 [550,700,1910,1936] CZUG Mexico 7 578 1 1 Veracruz de Ignacio de la Llave paratype 2620268302 [711,852,1910,1937] IEXA Mexico 7 578 1 1 Veracruz de Ignacio de la Llave paratype