Heimatomyces paradoxus Peyr., Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Wien . Mathematisch-naturwissenschaftliche Classe. Abteilung 1 68: 251 ( Peyritsch 1873 ) Heimatomyces unciger Thaxt. Thaxter 1895: 478 Chitonomyces unciger (Thaxt.) Thaxt. Thaxter 1896: 288 Chitonomyces truncatus Speg. ( Spegazzini 1915b: 47 ) Laboulbeniomycetes (Fungi, Ascomycota) of Denmark Santamaria, Sergi Pedersen, Jan European Journal of Taxonomy 2021 2021-11-26 781 1 425 5Y7QF (Peyr.) Thaxt. [556,1031,264,291] Laboulbeniomycetes Laboulbeniaceae Chitonomyces Fungi Laboulbeniales 105 106 Ascomycota species paradoxus  MB#170184  Fig. 21E    Memoirs of the American Academy of Arts and Sciences12: 287 ( Thaxter 1896: 287). – Basionym:    Heimatomyces paradoxusPeyr., Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften.  Wien. Mathematisch-naturwissenschaftliche Classe. Abteilung 168: 251 ( Peyritsch 1873)[MB#249906]. –     Type: [Not designated] [Ind. loc.] [Type lost?!] “Mit der vorhergehenden Art auf  Laccophilus minutusSturmund  L. hyalinusDegeer”. [ Austria]     Heimatomyces uncigerThaxt.(  Thaxter 1895: 478, as uncigerus) [MB#245659]. –   Chitonomyces unciger(Thaxt.) Thaxt.(  Thaxter 1896: 288, as uncigerus) [MB#175610]     Chitonomyces truncatusSpeg. ( Spegazzini 1915b: 47)[MB#176303]    Diagnostic features The ascogenous cavity inside the perithecium appears nearly horizontal, strongly curved, u-shaped. The ostiole is surrounded by obtuse, usually prominent and asymmetrical lips ( Fig. 21E, *). The perithecium bears a variably elongated horn-like outgrowth arising from the upper outer wall cell of the anterior row. [Detailed descriptions: Thaxter 1916, 1924; Sugiyama 1973; Majewski 1994b; Santamaria 2001b, 2003; De Kesel & Werbrouck 2008; Goldmann & Weir 2012]    Distribution and hosts More common than  Ch. melanuruswith which it often coexists on the same host individual, occupying a similar position but just below it. Reported on  Laccophilus(Col. Dytiscidae) in America: Grenada, Jamaica, USA, Trinidad & Tobago( Santamaria et al.1991), Bahamas( Haelewaters et al.2017); Asia: Indonesia, Japan, Taiwan, Thailand( Santamaria et al.1991), Korea( Lee & Kim 1990), China( Lee et al.1995), Turkey( Rossi 2016), Cambodia( Try et al.2017); Africa: Sierra Leone( Rossi & Leonardi 2018); and Europe: Austria( type), Finland, France, Germany, Hungary, Italy, Poland, Russia, Spain, United Kingdom, former Yugoslavia( Santamaria et al.1991), Belgium( De Kesel & Werbrouck 2008), Sweden( Huggert 2010).    Collections examined from Denmark  On  Laccophilus minutus(Linnaeus, 1758)(Col. Dytiscidae)   DENMARK–  Bornholm(B)• Raghammer Odde; 55°1.193′ N, 14°55.777′ E; VA99;  22 Jun. 2018;  JP1146; JPdet.; ZMUC C-F-123668. –   Nordvestsjaelland(NWZ)• Klintebjerg ved Klint; 55°57.321′ N, 11°34.197′ E; PH60;  16 Oct. 2017;  JP589; JPdet.; ZMUC C-F-123078. –   Vestjylland(WJ)• Simmelmose; 55°46.299′ N, 9°3.519′ E; NG08;  2 Jun. 2019;  JP1478; JPdet.; ZMUC C-F-124063.    Remarks First record from Denmark. Three forms of this species may be perfectly distinguished by morphological traits, depending on the hosts, being  Laccophilus minutus,  L. poecilus(=  L. variegatus) and  L. hyalinus. These forms were described and illustrated by Majewski (1994b)and Santamaria (2001b, 2003). Further studies supported by molecular analysis might clarify the taxonomic position of these forms. Danish thalli belong to the form growing on  L. minutus. 4988642159 Austria Austria 105 106 1 type 4988642074 2018-06-22 ZMUC JP Denmark JP 55.019882 Raghammer Odde 1 14.929617 105 106 ZMUC C-F-123668 1 Bornholm 4988642250 2017-10-16 JP Denmark JP 55.95535 Klintebjerg ved Klint 1 11.56995 105 106 ZMUC C-F-123078 1 Nordvestsjaelland 4988640642 2019-06-02 JP Denmark JP 55.77165 Simmelmose 1 9.05865 105 106 ZMUC C-F-124063 1 Vestjylland