Fabricius, 1781 : 404 Fabricius, Fabricius 1787 : 323 Fabricius, Fabricius 1805 : 22 Westwood 1835 : 282 Bittacomorpha Bittacomorpha clavipes Fabricius, 1890: 230 Aldrich, 1895 : 200 Johnson 1905 : 75 Alexander 1916 : 546 Alexander 1920 : 784 Alexander 1927 : 9 Bittacomorpha clavipes Fabricius, Peus 1968: 11 Alexander 1965 : 98 Alexander 1967 : 171 A revision of Bittacomorphinae with a review of the monophyly of extant subfamilies of Ptychopteridae (Diptera) Fasbender, Andrew Courtney, Gregory W. Zootaxa 2017 4309 1 1 69 LXQ4 Fabricius 1781 Fabricius 1781 [151,627,1453,1479] Insecta Ptychopteridae Bittacomorpha Animalia Diptera 29 30 Arthropoda species clavipes     Tipula clavipes  Fabricius, 1781: 404(original description)    Tipula clavipes  Fabricius, Fabricius 1787: 323(redescription)    Ptychoptera clavipes  Fabricius, Fabricius 1805: 22(catalog, redescription)  Bittacomorpha clavipesFabricius,  Westwood 1835: 282(set as typespecies of  Bittacomorpha)  Bittacomorpha clavipesFabricius, 1890: 230(key)    Bittacomorpha clavipes  Aldrich, 1895: 200(original description)    Bittacomorpha clavipesFabricius,  Johnson 1905: 75(key)    Bittacomorpha (Bittacomorpha) clavipesFabricius,  Alexander 1916: 546(key)  Bittacomorpha clavipesFabricius,  Alexander 1920: 784–787 (redescription)  Bittacomorpha clavipesFabricius,  Alexander 1927: 9(catalog)   Bittacomorpha clavipesFabricius, Peus 1968: 11(catalog)    Bittacomorpha clavipesFabricius,  Alexander 1965: 98(catalog)    Bittacomorpha clavipesFabricius,  Alexander 1967: 171(illustration)   Diagnosis. Larva( Figs. 5a, 6a–b). Genae round and abbreviate, cranium olive-colored and spotted with black; postgena with rounded anterolateral lobes. Abdominal segments with linear rows of tubercles transverse dorsally and ventrally, longitudinal laterally. Adult( Figs. 7a, 8, 9). Scutum with longitudinal white stripe. Male with paramere apex weakly furcate.   Description. Larva.Dorsal surface of cranium domed, hemispherical, olive-colored, rows of dark spots posteriorly on frontoclypeal apotome and dorsal surface of genae. Genae rounded in dorsal view, not expanded posteriorly, ventroposterior margin weakly and broadly emarginate; subgenal sclerite narrowed at midpoint of length, apex with semicircular anterolateral lobes. Antenna two-segmented, basal segment cylindrical, apical segment weakly fusiform, apex rounded, apical sensilla cluster directed laterally. Abdominal segments with rows of tubercles transverse dorsally and ventrally, longitudinal laterally. Adult. Measurements (n=5).Overall length 10.82mm( 10.51–11.16mm) head width 1.06mm( 1.02–1.12mm); height 0.95mm(0.88–1.0mm), length 0.61mm( 0.57–0.67mm); antennal flagellum length 5.17mm, flagellomere proportions 1.00, 0.82, 0.89, 1.00, 1.00, 0.89, 0.94, 0.89, 0.76, 0.76, 0.64, 0.64, 0.64, 0.51, 0.43, 0.43, 0.38, 0.38, 0.17; maxillary palpus length 1.49mm, maxillary palpus segment proportions 0.24, 1.0, 1.3, 0.96, 1.0; thorax length 1.96mm( 1.65–2.22mm), height 1.78mm( 1.65–1.91mm); wing length 7.26mm( 6.7–7.9mm), width 1.42mm( 1.31–1.6mm), R3+4+5 stem 0.60mm, straight; R4+5 stem 2.00mm, R4 after fork 1.25mm, R5 after fork 1.23mm, R3+4+5:R4+5-stem: R4 1.00, 3.33, 2.08, r-m 0.30mm, M distal to r-m 2.97mm, Mdistal:R3+4+5 4.95, 1.00, R4:Mdistal 0.42, 1.00. CuA2 1.78mm; leg femur 1.0, 1.12, 1.27, tibia 0.98, 0.95, 1.19, tarsomeres foreleg 0.53, 0.22, 0.14, 0.05, 0.04, midleg 0.54, 0.21, 0.12, 0.05, 0.03, hind leg 0.58, 0.25, 0.16, 0.04, 0.06; abdomen length 11.87mm( 10.85–13.15mm), male tergite proportions 1.00, 3.13, 2.20, 2.23, 1.88, 0.89, 0.28, male genitalia width 1.03mm, height 0.71mm. Head.Face narrow, incised by clypeus to dorsal margin. Clypeus subcylindrical, lateral margins somewhat pinched at junction with face, shorter than eye, inflated. Hypopharynx subconical triangular, directed anteroventrally. Labium beyond clypeus greater than ½ length; labella pad subequal width to labium. Scutum and prescutum pollinose black, presecutum with white line along medial suture. Wing membrane without macrotrichia, apex of veins R1 and R3, with macrotrichia, others bare. Posthalter palpiform. Male.Epandrium with posterolateral corners simple right angle, posterior margin transverse laterally, with semicircular emargination medially; posteriomedial stylus short, triangular. Internal fold extending medially with slight posterior deflection. Epandrial clasper pad width of subapical clasper. Subepandrial sclerite divided, broadly sickle-shaped, fused to medial margin of base of epandrial clasper, directed towards hypoproct (additional small crescent-shaped weak sclerotization exists between subepandrial sclerites in subepandrial membrane in some specimens). Paramere with lateral spine at 7/8 length supporting lateral membrane; apex subacute triangular spine, with prominent dorsal tooth subapically. Gonostylus apex with or without irregularly spaced trichoid sensilla. Hypandrium terminal division foliform, petiolate base, two flattened lateral lobes directed laterally, posterior margin variable. Lateral ejaculatory process lateral apex broadly triangular, medial apex acute fingerlike; ventral lobe dorsally subacute quadrilateral towards blunt anterior apex, apex with lightly sclerotized region directed laterally, deeply inserted into sperm sac. Aedeagal sclerites widely separated at base, ventral apical apodeme apex pointed, fused to subapical anterior portion of subaedeagal sclerite. Subaedeagal sclerite internal section anterolateral apex obtuse round point, medial margin emarginate. External portion of subaedeagal sclerite extending dorsally beyond apex of dorsal apical apodeme. Female.Epigynium posterior margin V-shaped, medial suture present, faint; hypoproct broad, tongue-like, with sclerotized triangular boat-like sclerite ventrally, cercus dorsal margin straight, apex rounded, ventral margin swollen, potbellied anteriorly, tapering posteriorly. Genital fork spatulate, posterior margin weakly emarginate. Hypogynium a hemi-ring, hypogynial valve a short triangular lobe medially, lateral margins straight and angled dorsally, medial notch square: 0.75x as deep as broad.  Material examined.   COSTA RICA. GUANACOSTE. Bijagua,  22.ii.1991, coll. D.W. Webb& M.E. Irwin[pinned 1 ♂; INHS].   UNITED STATES. ALABAMA.  Hale Co. Payne Lake,  24 mi. SE. of Tuscaloosa,  25.v.1978, coll. G.W. Byers, C.W. Young[pinned 1 ♂, 1 ♀on same pin; KU];   Lee Co.Chewacia State Park  4 mi. SW. of Auburn,  26.v.1978, coll. G.W., Byers, C.W. Young[pinned 1 ♀; KU].  ARKANSAS.  Hot Spring Co. Malvern, Citypark,  22.v.78, coll. G.W. Byers, C.W. Young[pinned 3 ♂; KU];   Montgomery Co. Gap Creek,  1.3mi. SE. of Joplin,  22.v.1978, coll. G.W. Byers, C.W. Young[pinned 1 ♂; KU];  Gap Creek Picnic Grd.  1 miSE Joplin,  1.vi.1984, coll. S.A. Teale[pinned 1 ♂; KU];   2 miNW Mount Ida,  1.vi.1984, coll. S.A. Teale[pinned 1 ♂; KU].  COLORADO.  Fremont Co., Hayden Creek  4 mi. SW. Coaldale,  13.vii.1974, coll. G.W. Byers, C.W. Young[pinned 8 ♂, 1 ♀; KU];   Gunnison Co. Gothic  23.vii.1952, coll. B. Pruitt[pinned 6 ♂, 1 ♀; UMMZ];   Mineral Co. Wolf CreekCpgr. San Juan Natl. Forest,  25.vii.1981, coll. G.W. Byers, S.A. Teale[pinned 3 ♀; KU];   San Juan Co.  30 mi. N. Durango,  13.viii.1965, coll. G.W. Byers[pinned 1 ♀; KU]. Unknown locality, collector and date [pinned 1 ♂; WSU].  CONNECTICUT.  Fairfield Co.Redding,  2.viii.1928, coll. A.L. Melander[pinned 1 ♂; USNM];  GEORGIA.  Butts Co.Unknown locality and collector,  6.v.1973[pinned 2 ♂; FMNH];   Fulton Co. East Point,  15.v.1965, coll. B. Harrison[pinned 1 ♀; KU];   Lumpkin Co.#5 De Soto Falls,  23.vi.1967, coll. G.W. Byers[pinned 2 ♂; KU];   Oconee Co.  2mi. N. Watkinsville,  3.x.1977, coll. G.W. Byers[pinned 2 ♂; KU];   Union Co. Lake Winfield Scott,  23.vi.1967, coll. G.W. Byers[pinned 1 ♂; KU].  ILLINOIS.  Cook Co. Palos Park,  20.vi.1909, coll. W.J. Gerhard[pinned 1 ♂; FMNH];   Dupage Co. Argonne Natl. Lab,  11.ix.1974, coll. D. Pearson[pinned 1 ♂; FMNH];   Kane Co.  23.vii.1972, coll. D. Pearson[pinned 2 ♂, 4 ♀, additional dates  July 23, 1972–  July 27, 1973; FMNH].  INDIANA.  Lagrange Co. Pigeon River,  4.ix.1928 B.E.M.[pinned 1 ♀, UMMZ];   LaPorte Co. Michigan City,  2.ix.1955, coll. H. Dybas[pinned 1 ♂; FMNH];  IOWA.  Dickinson Co. Iowa Lakeside Lab,  18.vii.1959, J.L. Laffoon[pinned 1 ♂; ISIC];  Iowa Lakeside Lab Hillside Seepnear marsh,  17.v.2010, 43.383374°N, 95.179974°W, coll. G.W. Courtney, A. Fasbender[pinned 2 ♀; ISIC];  Silver LakeFen  17.v.2010, 43.437472°N, 95.365009°W, coll. G.W. Courtney, A. Fasbender[pinned 11 ♂, 9 ♀; alcohol: 4 ♂, 3 ♀, 3 larvae, adultsreared; ISIC];   Guthrie Co.Springbrook State Park,  24.viii.1972, A. Stewart Cott[pinned 1 ♂, 1♀; ISIC],   19.vii.1960, coll. J. Laffoon[pinned 1 ♂; ISIC];   Story Co. Ames,  17.v.1964, coll. W.S. Craig[pinned 43 ♂, 2 ♀; ISIC].  KANSAS.  Douglas Co. Lawrence, Snow Hall,  8.vii.1987, J. Gelhaus[pinned 1 ♀; KU]   19.v.1986, coll. B.N. Danforth[pinned 1♀; KU].  MAINE.  Hancock Co. Bar Harbor,  5.viii.1936, coll. A.E. Brower[pinned 2 ♂; USNM];  Ellsworth,  12.ix.1967, coll. P.J. Spangler[pinned 1 ♀; USNM];   Penobscot Co. Orono,  18.viii.1981, coll. S. Struad[pinned 1 ♂; KU].  MICHIGAN.  Calhoun Co. Battle Creek,  5.vii.1979, G.W. Byers[pinned 1 ♀; KU];  Battle Creek, date and collector unknown [pinned 3 ♂, 5 ♀; WSU];   Cheboygan Co. Cochran Lake,  19.vii.1957, coll. R.E. Beer[pinned 1 ♂; KU];   4mi. N. of Topinbee,  8.vii.1961, coll. H. Willis[pinned 1 ♀; KU]; No collector,  vii.1931[pinned 1 ♂, KU];   Emmet Co.  4mi. W. Pellston,  15.vii.1969[pinned 3 ♂, 2 ♀; UMMZ];   Gratiot Co. Ithaca,  13.vii.1959, coll. V. Potter[pinned 3 ♂, 5 ♀; UMMZ];   Jackson Co.  7.v.1939, coll. F. Awnerman[pinned 1 ♀, UMMZ];   Kalamazoo Co. Gull LakeBiol. Sta.,  4.vii.1965, coll. B. Hlavac, T. Hlavac[pinned 1 ♀; FMNH];   Schoolcraft Co.  14.viii.1951, coll. J.S. Rogers[pinned 1 ♂; UMMZ].  MINNESOTA.  Clearwater Co. Itasca State Park,  2.ix.1950, coll. J.L. Laffoon[pinned 6♂, 1 ♀; ISIC].  MISSISSIPPI.  Lafayette Co. Puskus Lake,  8mi. NE. Oxford,  23.v.1978, G.W. Byers, C.W. Young[pinned 1 ♀; KU];   Warren Co. Vicksburg,  23.ix.1977, coll. G.W. Byers[pinned 1 ♂; KU].  NEBRASKA.  Cherry Co. Valentine,  9.vi.1950, coll. Hicks, Slater, J.L. Laffoon, multiple dates  9–11.vi.1950[pinned 6 ♂, 5 ♀; ISIC];   Sheridan Co. Rushville, Malaise Trap, coll. R.W. Dawson,  12.vii.1967[pinned 1 ♂; WSU].  NEW YORK.  Chenango Co. Smyrna,  16.viii.25, coll. A.C. Weed[pinned 1 ♀; FMNH];   Clinton Co. Saranac Inn, 1900, coll. O.S. Wescott[pinned 5 ♂; KU];   Hamilton Co.  7 milesS. of Long Lake,  26.viii.1956, coll. W.E. LaBerge[pinned 2 ♂];   Wayne Co. North Rose,  17.vi.1927, Rawson-MacMillan Exped., coll. A.C. Weed[pinned 1 ♀; FMNH].  NORTH CAROLINA.  Buncombe Co. Asheville, on store window,  29.iv.62, coll. R.D. Peterson[pinned 1 ♀; ISIC];   Swain Co.1.2, mi. NE. Clingman’s Dome, GSMNP,  15.vi.1967, coll. G.W. Byers[pinned 1 ♂; KU].  OHIO.  Portage Co.  4 mi. E Kent,  8.viii.1967, coll. M. Duckenonod[pinned 1 ♀; USNM].  PENNSYLVANIA.  Berks Co. French Creek Park,  7.vii.1960, coll. G.W. Byers, multiple dates  7–10.vii.1960[pinned 30 ♂, 2 ♀; KU];   1mi. SE. Reading,  15.viii.1956, G.W. Byers[pinned 3♂, 1 ♀; KU];   Franklin Co. Caledonia St. Park,  2.vii.1960, coll. G.W, Byersand party [pinned 2 ♂, KU];   Luzerne Co. Hazelton,  8.vii.1979, coll. G.W. Byers[pinned 1 ♂; KU].  SOUTH CAROLINA.  Abbeville Co.  5 mi. S. Abbeville,  29.v.1978, coll. G.W. Byers, C.W. Young[pinned 2 ♂, 1 ♀; KU];   Oconee Co. Westminster,  30.v.1978, coll. G.W., Byers, C.W. Young[pinned 1 ♀; KU];  Oconee State Park, Small lake  20.viii.1939, coll. B.E. Montgomery#3974 [pinned 1 ♂; UMMZ];   Pickens Co. Clemson U. Experimental Forest, Six-Mile Creek,  17.iii.2005, 34.75°N, 82.85°W, coll. G.W. Courtney[alcohol 5 pupae; ISIC] coll. M.J. Petersen[ 1 ♂, 2 pupae; ISIC];  SOUTH DAKOTA  Pennington Co. Black Hills Nat. For., Streamnr. South Rochford Road, Sweeping, 44.109621°N 103.734564°W, A Fasbender  29.vi.2010[alcohol 3 ♂; ISIC];   Custer Co. Custer State Park, French Creek Picnic,  30.vi.2008, 43.718°N, 103.4891°W, coll. J.D. Petersen[alcohol: 1 ♀, ISIC];  Custer State Park, Sylvan Lake, 30.  vi.20 0 8, 43.8459°N, 103.5602°W, coll.: J.D. Petersen[alcohol 1 ♀; ISIC].  TENNESSEE  Fentress Co. Allardt,  2.vi.1978, coll. G.W., Byers, C.W. Young[pinned 2 ♂; KU];   Sevier Co. Fighting Creek, Great Smoky Mts.  7.vii.1958, 35.67833°N, 83.555°W, J.L. Laffoon[pinned 1 ♂; ISIC].  TEXAS.  Bastrop Co. Bastrop St. Park,  20.iii.1976, E.M. May[pinned 1 ♂; KU];   Nacogdoches Co. Nacogdoches,  12.x.1960, coll. R. Oliver, multiple dates  8.iv.1960–  6.v.1962[pinned 7 ♂, 1 ♀; KU].  VERMONT  Orleans Co.  5 mi. NNE. Orleans,  19.vi.1982, coll. G.W. Byers[pinned 1 ♀; KU].  VIRGINIA.  Arlington Co. Alexandria,  15.vi.1973, coll. B.M. O’Connor[pinned 1 ♀; UMMZ];   Fairfax Co. Lake Accotink,  5.viii.1972, coll. B.M. O’Connor[pinned 1 ♀; UMMZ];   Giles Co. Mt. Lake Biol. Stn.  6–12.viii.1979, coll. G.W. Byers#163 [pinned 1 ♂; KU];   Prince William Co. Woodbridge,  28.viii.1946, coll. L.D. Beamer[pinned 1 ♂; KU].  WISCONSIN.  Vilas Co. Squaw Lake,  16.viii.1954, coll. J.L. Laffoon[pinned 2 ♂; ISIC].  WYOMING.  Albany Co. Medicine Bow Nat. For. North Laramie Riverat Sand Lake Rd. Beaver Dam,  21.vi.2012 41.351557°N, 106.166511°W, coll. A Fasbender[pinned 1 ♀; ISIC].  Taxonomic notes.This species can be easily differentiated from its congener  Bittacomorpha occidentalisby the medial white stripe on the scutum, shortened and narrowed clypeus, and paramere with lateral spur and apical dorsal tooth. Larvae can be distinguished in dorsal view, the head capsule of  Bi. clavipesbeing ovoid and that of  Bi. occidentalistriangular. The holotype of  Bittacomorpha clavipesis not listed in the collection of Fabricius type material at ZMUC, and may be lost or destroyed. Fortunately, the distinctive character of the inflated 1 st tarsomereallows the determination of the genus without contention, and material from the range of  Bittacomorpha occidentaliswould not have been available to Fabricius at the time.   Distribution. Fig. 18a. 2556241568 1991-02-22 INHS D. W. Webb & M. E. Irwin Costa Rica Bijagua Bijagua 31 32 1 1 GUANACOSTE 1586248125 1978-05-25 KU G. W. Byers & C. W. Young United States of America Hale Co. SE. of Tuscaloosa Payne Lake 31 32 2 1 1 ALABAMA 2556241576 1978-05-26 KU G. W., Byers & C. W. Young United States of America SW. of Auburn Lee Co. Chewacia State Park 31 32 1 1 ALABAMA 2556241570 1978-05-22 KU G. W. Byers & C. W. Young United States of America Hot Spring Co. City Malvern 31 32 3 3 ARKANSAS 1586248136 1978-05-22 KU G. W. Byers & C. W. Young United States of America Montgomery Co. SE. of Joplin Gap Creek 31 32 1 1 ARKANSAS 2556241581 1984-06-01 KU S. A. Teale United States of America Gap Creek 31 32 1 1 ARKANSAS 2556241583 [498,1262,1531,1556] 1984-06-01 KU S. A. Teale United States of America Mount Ida 31 32 1 1 ARKANSAS 2556241551 1974-07-13 KU G. W. Byers & C. W. Young United States of America Fremont Co. Coaldale Hayden Creek 31 32 9 1 8 COLORADO 2556241571 [216,1078,1603,1628] 1952-07-23 UMMZ B. Pruitt United States of America Gothic Gunnison Co. 31 32 7 1 6 COLORADO 2556241584 1981-07-25 KU G. W. Byers & S. A. Teale United States of America Mineral Co. San Juan Natl. Forest Wolf Creek 31 32 3 3 COLORADO 2556241543 1965-08-13 KU G. W. Byers United States of America San Juan Co. Durango 31 32 1 1 COLORADO 2556241574 WSU United States of America 31 32 1 1 COLORADO 1586248158 [246,1432,1711,1736] 1928-08-02 USNM A. L. Melander United States of America Fairfield Co. Redding Fairfield Co. Redding 31 32 1 1 CONNECTICUT 1586248179 [151,1159,1747,1772] 1973-05-06 FMNH United States of America Butts Co. Butts Co. 31 32 2 2 GEORGIA 1586248156 1965-05-15 KU B. Harrison United States of America East Point Fulton Co. 31 32 1 1 GEORGIA 1586248141 1967-06-23 KU G. W. Byers United States of America De Soto Falls Lumpkin Co. 31 32 2 2 GEORGIA 1586248137 [286,1231,1819,1844] 1977-10-03 KU N. Watkinsville & G. W. Byers United States of America Oconee Co. 31 32 2 2 GEORGIA 2556241552 1967-06-23 KU G. W. Byers United States of America Union Co. Lake Winfield Scott Union Co. Lake 31 32 1 1 GEORGIA 2556241572 1909-06-20 FMNH W. J. Gerhard United States of America Cook Co. Palos Park Palos Park 31 32 1 1 ILLINOIS 2556241559 1974-09-11 FMNH D. Pearson United States of America Dupage Co. Argonne Natl. Lab 31 32 1 1 ILLINOIS 1586248161 1972-07-23 1973-07-27 1972-07-23 FMNH D. Pearson United States of America Kane Co. 31 32 6 4 2 ILLINOIS 1586248147 [256,1169,1963,1988] 1928-09-04 1955-09-02 1928-09-04 UMMZ B. E. M. United States of America Lagrange Co. Pigeon River Pigeon River 31 32 1 1 1 INDIANA 1586248132 1955-09-02 1955-09-02 1928-09-04 FMNH H. Dybas United States of America LaPorte Co. Michigan City Lagrange Co. Pigeon River 31 32 1 1 1 INDIANA 2556241586 1959-07-18 ISIC J. L. Laffoon United States of America Dickinson Co. 32 33 Iowa Lakeside Lab 31 32 1 1 IOWA 2556241541 2010-05-17 ISIC G. W. Courtney & A. Fasbender United States of America 43.383373 Hillside Seep 1 -95.17998 Iowa Lakeside Lab 32 33 2 2 IOWA 1586248142 2010-05-17 ISIC G. W. Courtney & A. Fasbender United States of America 43.437473 Silver Lake 1 -95.365005 Silver Lake 32 33 31 1 12 3 15 IOWA 2556241534 1972-08-24 ISIC A. Stewart Cott United States of America Guthrie Co. Springbrook State Park Guthrie Co. Springbrook State Park 32 33 2 1 1 IOWA 2556241573 1960-07-19 ISIC J. Laffoon United States of America 32 33 1 1 IOWA 1586248145 [299,1143,295,320] 1964-05-17 ISIC W. S. Craig United States of America Ames Story Co. 32 33 45 2 43 IOWA 2556241565 1987-07-08 KU J. Gelhaus United States of America Douglas Co. Snow Hall Lawrence 32 33 1 1 KANSAS 2556241553 [880,1433,332,357] 1986-05-19 KU B. N. Danforth United States of America 32 33 1 1 KANSAS 2556241540 [151,1169,367,392] 1936-08-05 A. E. Brower United States of America Hancock Co. Bar Harbor 32 33 2 2 MAINE 1586248152 1967-09-12 1981-08-18 1967-09-12 USNM P. J. Spangler United States of America Ellsworth Penobscot Co. 32 33 1 1 1 MAINE 1586248157 [631,1432,404,429] 1981-08-18 1981-08-18 1967-09-12 KU S. Struad United States of America Orono Penobscot Co. 32 33 1 1 1 MAINE 1586248146 [151,1073,439,464] 1979-07-05 KU G. W. Byers United States of America Calhoun Co. Battle Creek 32 33 1 1 MICHIGAN 2556241546 WSU United States of America Battle Creek Battle Creek 32 33 8 5 3 MICHIGAN 2556241569 [557,1431,476,501] 1957-07-19 KU R. E. Beer United States of America Cheboygan Co. Cochran Lake Cochran Lake 32 33 1 1 MICHIGAN 2556241544 [151,883,511,536] 1961-07-08 KU H. Willis United States of America Topinbee 32 33 1 1 MICHIGAN 2556241585 [895,1347,511,536] 1931-07 KU United States of America 32 33 1 1 MICHIGAN 1586248165 1969-07-15 UMMZ W. Pellston United States of America Emmet Co. Emmet Co. 32 33 5 2 3 MICHIGAN 1586248174 1959-07-13 UMMZ V. Potter United States of America Ithaca Gratiot Co. 32 33 8 5 3 MICHIGAN 1586248151 [471,1191,583,608] 1939-05-07 UMMZ F. Awnerman United States of America Jackson Co. 32 33 1 1 MICHIGAN 1586248176 1965-07-04 FMNH B. Hlavac & T. Hlavac United States of America Kalamazoo Co. Gull Lake Gull Lake 32 33 1 1 MICHIGAN 1586248197 1951-08-14 UMMZ J. S. Rogers United States of America Schoolcraft Co. 32 33 1 1 MICHIGAN 1586248168 1950-09-02 ISIC J. L. Laffoon United States of America Clearwater Co. Itasca State Park Itasca State Park 32 33 7 1 6 MINNESOTA 1586248173 1978-05-23 KU G. W. Byers & C. W. Young United States of America Lafayette Co. Oxford Puskus Lake 32 33 1 1 MISSISSIPPI 1586248159 [455,1266,727,752] 1977-09-23 KU G. W. Byers United States of America Warren Co. Vicksburg 32 33 1 1 MISSISSIPPI 2556241564 1950-06-09 1950-06-11 1950-06-09 ISIC Hicks & Slater & J. L. Laffoon United States of America Cherry Co. Valentine Valentine 32 33 11 5 6 NEBRASKA 2556241556 [229,1267,799,824] 1967-07-12 Malaise Trap WSU R. W. Dawson United States of America Rushville Sheridan Co. 32 33 1 1 NEBRASKA 2556241535 1925-08-16 FMNH A. C. Weed United States of America Chenango Co. Smyrna 32 33 1 1 NEW YORK 2556241554 1900 KU O. S. Wescott United States of America Saranac Inn Clinton Co. 32 33 5 5 NEW YORK 2556241577 [463,1430,871,896] 1956-08-26 W. E. LaBerge United States of America Long Lake Hamilton Co. 32 33 2 2 NEW YORK 2556241560 [151,1329,908,933] 1927-06-17 FMNH A. C. Weed United States of America North Rose Wayne Co. 32 33 1 1 NEW YORK 2556241557 1962-04-29 ISIC R. D. Peterson United States of America Buncombe Co. Asheville 32 33 1 1 NORTH CAROLINA 1586248175 1967-06-15 KU G. W. Byers United States of America Swain Co. Clingman's Dome 32 33 1 1 NORTH CAROLINA 2556241531 1967-08-08 USNM M. Duckenonod United States of America Portage Co. Kent Kent 32 33 1 1 OHIO 2556241539 1960-07-07 1960-07-10 1960-07-07 KU G. W. Byers United States of America Berks Co. French Creek Park 32 33 32 2 30 PENNSYLVANIA 2556241563 1956-08-15 KU G. W. Byers United States of America Reading 32 33 4 1 3 PENNSYLVANIA 1586248138 1960-07-02 1979-07-08 1960-07-02 KU G. W, Byers United States of America Franklin Co. Caledonia St. Park 32 33 2 2 PENNSYLVANIA 1586248164 [467,1324,1124,1149] 1979-07-08 1979-07-08 1960-07-02 KU G. W. Byers United States of America Luzerne Co. Hazelton 32 33 1 1 PENNSYLVANIA 1586248169 1978-05-29 1978-05-30 1978-05-29 KU S. Abbeville & G. W. Byers & C. W. Young United States of America Abbeville Co. Abbeville Co. 32 33 3 1 2 SOUTH CAROLINA 2556241542 [151,1133,1196,1221] 1978-05-30 1978-05-30 1978-05-29 KU G. W., Byers & C. W. Young United States of America Oconee Co. Westminster 32 33 1 1 2 SOUTH CAROLINA 2556241575 1939-08-20 UMMZ B. E. Montgomery United States of America Small lake Oconee State Park 32 33 1 1 SOUTH CAROLINA 1586248200 2005-03-17 ISIC G. W. Courtney & M. J. Petersen United States of America 34.75 Clemson U. 718 -82.85 Pickens Co. 32 33 8 1 7 SOUTH CAROLINA 2556241548 2010-06-29 ISIC United States of America Pennington Co. 44.10962 Stream 1 -103.734566 Black Hills Nat. For. 32 33 3 3 SOUTH DAKOTA 1586248185 2008-06-30 ISIC J. D. Petersen United States of America Custer Co. 43.718 French Creek Picnic 55 -103.4891 Custer State Park 32 33 1 1 SOUTH DAKOTA 1586248166 vi. 2 ISIC J. D. Petersen United States of America 43.8459 Custer State Park 6 -103.5602 32 33 1 1 SOUTH DAKOTA 1586248195 [227,1307,1447,1472] 1978-06-02 KU G. W., Byers & C. W. Young United States of America Fentress Co. Allardt 32 33 2 2 TENNESSEE 1586248202 1958-07-07 ISIC J. L. Laffoon United States of America Sevier Co. 35.67833 Great Smoky Mts. 45 -83.555 Fighting Creek 32 33 1 1 TENNESSEE 1586248189 1976-03-20 KU E. M. May United States of America Bastrop Co. Bastrop St. Park 32 33 1 1 TEXAS 2556241567 1960-04-08 1962-05-06 1960-04-08 KU R. Oliver United States of America Nacogdoches Co. Nacogdoches 32 33 8 1 7 TEXAS 1586248178 1982-06-19 KU G. W. Byers United States of America Orleans Co. Orleans Orleans 32 33 1 1 VERMONT 1586248198 1973-06-15 UMMZ B. M. O'Connor United States of America Arlington Co. Alexandria 32 33 1 1 VIRGINIA 1586248201 1972-08-05 UMMZ B. M. O'Connor United States of America Fairfax Co. Lake Accotink 32 33 1 1 VIRGINIA 1586248199 1979-08-06 1979-08-12 1979-08-06 KU G. W. Byers United States of America Giles Co. Mt. Lake Biol. Stn. 32 33 1 1 VIRGINIA 1586248191 [216,1153,1700,1725] 1946-08-28 KU L. D. Beamer United States of America Prince William Co. Woodbridge 32 33 1 1 VIRGINIA 1586248171 1954-08-16 ISIC J. L. Laffoon United States of America Vilas Co. Squaw Lake Squaw Lake 32 33 2 2 WISCONSIN 2556241579 2012-06-21 ISIC A Fasbender United States of America Albany Co. 41.35156 Sand Lake Rd. Beaver Dam 1 -106.16651 Medicine Bow Nat. For. North Laramie River 32 33 1 1 WYOMING