Fabricius, 1781 : 404
Fabricius, Fabricius 1787 : 323
Fabricius, Fabricius 1805 : 22
Westwood 1835 : 282
Bittacomorpha
Bittacomorpha clavipes Fabricius, 1890: 230
Aldrich, 1895 : 200
Johnson 1905 : 75
Alexander 1916 : 546
Alexander 1920 : 784
Alexander 1927 : 9
Bittacomorpha clavipes Fabricius, Peus 1968: 11
Alexander 1965 : 98
Alexander 1967 : 171
A revision of Bittacomorphinae with a review of the monophyly of extant subfamilies of Ptychopteridae (Diptera)
Fasbender, Andrew
Courtney, Gregory W.
Zootaxa
2017
4309
1
1
69
LXQ4
Fabricius 1781
Fabricius
1781
[151,627,1453,1479]
Insecta
Ptychopteridae
Bittacomorpha
Animalia
Diptera
29
30
Arthropoda
species
clavipes
Tipula clavipes Fabricius, 1781: 404(original description) Tipula clavipes Fabricius, Fabricius 1787: 323(redescription) Ptychoptera clavipes Fabricius, Fabricius 1805: 22(catalog, redescription) Bittacomorpha clavipesFabricius, Westwood 1835: 282(set as typespecies of Bittacomorpha) Bittacomorpha clavipesFabricius, 1890: 230(key) Bittacomorpha clavipes Aldrich, 1895: 200(original description) Bittacomorpha clavipesFabricius, Johnson 1905: 75(key) Bittacomorpha (Bittacomorpha) clavipesFabricius, Alexander 1916: 546(key) Bittacomorpha clavipesFabricius, Alexander 1920: 784–787 (redescription) Bittacomorpha clavipesFabricius, Alexander 1927: 9(catalog) Bittacomorpha clavipesFabricius, Peus 1968: 11(catalog) Bittacomorpha clavipesFabricius, Alexander 1965: 98(catalog) Bittacomorpha clavipesFabricius, Alexander 1967: 171(illustration)
Diagnosis. Larva( Figs. 5a, 6a–b). Genae round and abbreviate, cranium olive-colored and spotted with black; postgena with rounded anterolateral lobes. Abdominal segments with linear rows of tubercles transverse dorsally and ventrally, longitudinal laterally. Adult( Figs. 7a, 8, 9). Scutum with longitudinal white stripe. Male with paramere apex weakly furcate.
Description. Larva.Dorsal surface of cranium domed, hemispherical, olive-colored, rows of dark spots posteriorly on frontoclypeal apotome and dorsal surface of genae. Genae rounded in dorsal view, not expanded posteriorly, ventroposterior margin weakly and broadly emarginate; subgenal sclerite narrowed at midpoint of length, apex with semicircular anterolateral lobes. Antenna two-segmented, basal segment cylindrical, apical segment weakly fusiform, apex rounded, apical sensilla cluster directed laterally. Abdominal segments with rows of tubercles transverse dorsally and ventrally, longitudinal laterally. Adult. Measurements (n=5).Overall length 10.82mm( 10.51–11.16mm) head width 1.06mm( 1.02–1.12mm); height 0.95mm(0.88–1.0mm), length 0.61mm( 0.57–0.67mm); antennal flagellum length 5.17mm, flagellomere proportions 1.00, 0.82, 0.89, 1.00, 1.00, 0.89, 0.94, 0.89, 0.76, 0.76, 0.64, 0.64, 0.64, 0.51, 0.43, 0.43, 0.38, 0.38, 0.17; maxillary palpus length 1.49mm, maxillary palpus segment proportions 0.24, 1.0, 1.3, 0.96, 1.0; thorax length 1.96mm( 1.65–2.22mm), height 1.78mm( 1.65–1.91mm); wing length 7.26mm( 6.7–7.9mm), width 1.42mm( 1.31–1.6mm), R3+4+5 stem 0.60mm, straight; R4+5 stem 2.00mm, R4 after fork 1.25mm, R5 after fork 1.23mm, R3+4+5:R4+5-stem: R4 1.00, 3.33, 2.08, r-m 0.30mm, M distal to r-m 2.97mm, Mdistal:R3+4+5 4.95, 1.00, R4:Mdistal 0.42, 1.00. CuA2 1.78mm; leg femur 1.0, 1.12, 1.27, tibia 0.98, 0.95, 1.19, tarsomeres foreleg 0.53, 0.22, 0.14, 0.05, 0.04, midleg 0.54, 0.21, 0.12, 0.05, 0.03, hind leg 0.58, 0.25, 0.16, 0.04, 0.06; abdomen length 11.87mm( 10.85–13.15mm), male tergite proportions 1.00, 3.13, 2.20, 2.23, 1.88, 0.89, 0.28, male genitalia width 1.03mm, height 0.71mm. Head.Face narrow, incised by clypeus to dorsal margin. Clypeus subcylindrical, lateral margins somewhat pinched at junction with face, shorter than eye, inflated. Hypopharynx subconical triangular, directed anteroventrally. Labium beyond clypeus greater than ½ length; labella pad subequal width to labium. Scutum and prescutum pollinose black, presecutum with white line along medial suture. Wing membrane without macrotrichia, apex of veins R1 and R3, with macrotrichia, others bare. Posthalter palpiform. Male.Epandrium with posterolateral corners simple right angle, posterior margin transverse laterally, with semicircular emargination medially; posteriomedial stylus short, triangular. Internal fold extending medially with slight posterior deflection. Epandrial clasper pad width of subapical clasper. Subepandrial sclerite divided, broadly sickle-shaped, fused to medial margin of base of epandrial clasper, directed towards hypoproct (additional small crescent-shaped weak sclerotization exists between subepandrial sclerites in subepandrial membrane in some specimens). Paramere with lateral spine at 7/8 length supporting lateral membrane; apex subacute triangular spine, with prominent dorsal tooth subapically. Gonostylus apex with or without irregularly spaced trichoid sensilla. Hypandrium terminal division foliform, petiolate base, two flattened lateral lobes directed laterally, posterior margin variable. Lateral ejaculatory process lateral apex broadly triangular, medial apex acute fingerlike; ventral lobe dorsally subacute quadrilateral towards blunt anterior apex, apex with lightly sclerotized region directed laterally, deeply inserted into sperm sac. Aedeagal sclerites widely separated at base, ventral apical apodeme apex pointed, fused to subapical anterior portion of subaedeagal sclerite. Subaedeagal sclerite internal section anterolateral apex obtuse round point, medial margin emarginate. External portion of subaedeagal sclerite extending dorsally beyond apex of dorsal apical apodeme. Female.Epigynium posterior margin V-shaped, medial suture present, faint; hypoproct broad, tongue-like, with sclerotized triangular boat-like sclerite ventrally, cercus dorsal margin straight, apex rounded, ventral margin swollen, potbellied anteriorly, tapering posteriorly. Genital fork spatulate, posterior margin weakly emarginate. Hypogynium a hemi-ring, hypogynial valve a short triangular lobe medially, lateral margins straight and angled dorsally, medial notch square: 0.75x as deep as broad.
Material examined. COSTA RICA. GUANACOSTE. Bijagua, 22.ii.1991, coll. D.W. Webb& M.E. Irwin[pinned 1 ♂; INHS]. UNITED STATES. ALABAMA. Hale Co. Payne Lake, 24 mi. SE. of Tuscaloosa, 25.v.1978, coll. G.W. Byers, C.W. Young[pinned 1 ♂, 1 ♀on same pin; KU]; Lee Co.Chewacia State Park 4 mi. SW. of Auburn, 26.v.1978, coll. G.W., Byers, C.W. Young[pinned 1 ♀; KU]. ARKANSAS. Hot Spring Co. Malvern, Citypark, 22.v.78, coll. G.W. Byers, C.W. Young[pinned 3 ♂; KU]; Montgomery Co. Gap Creek, 1.3mi. SE. of Joplin, 22.v.1978, coll. G.W. Byers, C.W. Young[pinned 1 ♂; KU]; Gap Creek Picnic Grd. 1 miSE Joplin, 1.vi.1984, coll. S.A. Teale[pinned 1 ♂; KU]; 2 miNW Mount Ida, 1.vi.1984, coll. S.A. Teale[pinned 1 ♂; KU]. COLORADO. Fremont Co., Hayden Creek 4 mi. SW. Coaldale, 13.vii.1974, coll. G.W. Byers, C.W. Young[pinned 8 ♂, 1 ♀; KU]; Gunnison Co. Gothic 23.vii.1952, coll. B. Pruitt[pinned 6 ♂, 1 ♀; UMMZ]; Mineral Co. Wolf CreekCpgr. San Juan Natl. Forest, 25.vii.1981, coll. G.W. Byers, S.A. Teale[pinned 3 ♀; KU]; San Juan Co. 30 mi. N. Durango, 13.viii.1965, coll. G.W. Byers[pinned 1 ♀; KU]. Unknown locality, collector and date [pinned 1 ♂; WSU]. CONNECTICUT. Fairfield Co.Redding, 2.viii.1928, coll. A.L. Melander[pinned 1 ♂; USNM]; GEORGIA. Butts Co.Unknown locality and collector, 6.v.1973[pinned 2 ♂; FMNH]; Fulton Co. East Point, 15.v.1965, coll. B. Harrison[pinned 1 ♀; KU]; Lumpkin Co.#5 De Soto Falls, 23.vi.1967, coll. G.W. Byers[pinned 2 ♂; KU]; Oconee Co. 2mi. N. Watkinsville, 3.x.1977, coll. G.W. Byers[pinned 2 ♂; KU]; Union Co. Lake Winfield Scott, 23.vi.1967, coll. G.W. Byers[pinned 1 ♂; KU]. ILLINOIS. Cook Co. Palos Park, 20.vi.1909, coll. W.J. Gerhard[pinned 1 ♂; FMNH]; Dupage Co. Argonne Natl. Lab, 11.ix.1974, coll. D. Pearson[pinned 1 ♂; FMNH]; Kane Co. 23.vii.1972, coll. D. Pearson[pinned 2 ♂, 4 ♀, additional dates July 23, 1972– July 27, 1973; FMNH]. INDIANA. Lagrange Co. Pigeon River, 4.ix.1928 B.E.M.[pinned 1 ♀, UMMZ]; LaPorte Co. Michigan City, 2.ix.1955, coll. H. Dybas[pinned 1 ♂; FMNH]; IOWA. Dickinson Co. Iowa Lakeside Lab, 18.vii.1959, J.L. Laffoon[pinned 1 ♂; ISIC]; Iowa Lakeside Lab Hillside Seepnear marsh, 17.v.2010, 43.383374°N, 95.179974°W, coll. G.W. Courtney, A. Fasbender[pinned 2 ♀; ISIC]; Silver LakeFen 17.v.2010, 43.437472°N, 95.365009°W, coll. G.W. Courtney, A. Fasbender[pinned 11 ♂, 9 ♀; alcohol: 4 ♂, 3 ♀, 3 larvae, adultsreared; ISIC]; Guthrie Co.Springbrook State Park, 24.viii.1972, A. Stewart Cott[pinned 1 ♂, 1♀; ISIC], 19.vii.1960, coll. J. Laffoon[pinned 1 ♂; ISIC]; Story Co. Ames, 17.v.1964, coll. W.S. Craig[pinned 43 ♂, 2 ♀; ISIC]. KANSAS. Douglas Co. Lawrence, Snow Hall, 8.vii.1987, J. Gelhaus[pinned 1 ♀; KU] 19.v.1986, coll. B.N. Danforth[pinned 1♀; KU]. MAINE. Hancock Co. Bar Harbor, 5.viii.1936, coll. A.E. Brower[pinned 2 ♂; USNM]; Ellsworth, 12.ix.1967, coll. P.J. Spangler[pinned 1 ♀; USNM]; Penobscot Co. Orono, 18.viii.1981, coll. S. Struad[pinned 1 ♂; KU]. MICHIGAN. Calhoun Co. Battle Creek, 5.vii.1979, G.W. Byers[pinned 1 ♀; KU]; Battle Creek, date and collector unknown [pinned 3 ♂, 5 ♀; WSU]; Cheboygan Co. Cochran Lake, 19.vii.1957, coll. R.E. Beer[pinned 1 ♂; KU]; 4mi. N. of Topinbee, 8.vii.1961, coll. H. Willis[pinned 1 ♀; KU]; No collector, vii.1931[pinned 1 ♂, KU]; Emmet Co. 4mi. W. Pellston, 15.vii.1969[pinned 3 ♂, 2 ♀; UMMZ]; Gratiot Co. Ithaca, 13.vii.1959, coll. V. Potter[pinned 3 ♂, 5 ♀; UMMZ]; Jackson Co. 7.v.1939, coll. F. Awnerman[pinned 1 ♀, UMMZ]; Kalamazoo Co. Gull LakeBiol. Sta., 4.vii.1965, coll. B. Hlavac, T. Hlavac[pinned 1 ♀; FMNH]; Schoolcraft Co. 14.viii.1951, coll. J.S. Rogers[pinned 1 ♂; UMMZ]. MINNESOTA. Clearwater Co. Itasca State Park, 2.ix.1950, coll. J.L. Laffoon[pinned 6♂, 1 ♀; ISIC]. MISSISSIPPI. Lafayette Co. Puskus Lake, 8mi. NE. Oxford, 23.v.1978, G.W. Byers, C.W. Young[pinned 1 ♀; KU]; Warren Co. Vicksburg, 23.ix.1977, coll. G.W. Byers[pinned 1 ♂; KU]. NEBRASKA. Cherry Co. Valentine, 9.vi.1950, coll. Hicks, Slater, J.L. Laffoon, multiple dates 9–11.vi.1950[pinned 6 ♂, 5 ♀; ISIC]; Sheridan Co. Rushville, Malaise Trap, coll. R.W. Dawson, 12.vii.1967[pinned 1 ♂; WSU]. NEW YORK. Chenango Co. Smyrna, 16.viii.25, coll. A.C. Weed[pinned 1 ♀; FMNH]; Clinton Co. Saranac Inn, 1900, coll. O.S. Wescott[pinned 5 ♂; KU]; Hamilton Co. 7 milesS. of Long Lake, 26.viii.1956, coll. W.E. LaBerge[pinned 2 ♂]; Wayne Co. North Rose, 17.vi.1927, Rawson-MacMillan Exped., coll. A.C. Weed[pinned 1 ♀; FMNH]. NORTH CAROLINA. Buncombe Co. Asheville, on store window, 29.iv.62, coll. R.D. Peterson[pinned 1 ♀; ISIC]; Swain Co.1.2, mi. NE. Clingman’s Dome, GSMNP, 15.vi.1967, coll. G.W. Byers[pinned 1 ♂; KU]. OHIO. Portage Co. 4 mi. E Kent, 8.viii.1967, coll. M. Duckenonod[pinned 1 ♀; USNM]. PENNSYLVANIA. Berks Co. French Creek Park, 7.vii.1960, coll. G.W. Byers, multiple dates 7–10.vii.1960[pinned 30 ♂, 2 ♀; KU]; 1mi. SE. Reading, 15.viii.1956, G.W. Byers[pinned 3♂, 1 ♀; KU]; Franklin Co. Caledonia St. Park, 2.vii.1960, coll. G.W, Byersand party [pinned 2 ♂, KU]; Luzerne Co. Hazelton, 8.vii.1979, coll. G.W. Byers[pinned 1 ♂; KU]. SOUTH CAROLINA. Abbeville Co. 5 mi. S. Abbeville, 29.v.1978, coll. G.W. Byers, C.W. Young[pinned 2 ♂, 1 ♀; KU]; Oconee Co. Westminster, 30.v.1978, coll. G.W., Byers, C.W. Young[pinned 1 ♀; KU]; Oconee State Park, Small lake 20.viii.1939, coll. B.E. Montgomery#3974 [pinned 1 ♂; UMMZ]; Pickens Co. Clemson U. Experimental Forest, Six-Mile Creek, 17.iii.2005, 34.75°N, 82.85°W, coll. G.W. Courtney[alcohol 5 pupae; ISIC] coll. M.J. Petersen[ 1 ♂, 2 pupae; ISIC]; SOUTH DAKOTA Pennington Co. Black Hills Nat. For., Streamnr. South Rochford Road, Sweeping, 44.109621°N 103.734564°W, A Fasbender 29.vi.2010[alcohol 3 ♂; ISIC]; Custer Co. Custer State Park, French Creek Picnic, 30.vi.2008, 43.718°N, 103.4891°W, coll. J.D. Petersen[alcohol: 1 ♀, ISIC]; Custer State Park, Sylvan Lake, 30. vi.20 0 8, 43.8459°N, 103.5602°W, coll.: J.D. Petersen[alcohol 1 ♀; ISIC]. TENNESSEE Fentress Co. Allardt, 2.vi.1978, coll. G.W., Byers, C.W. Young[pinned 2 ♂; KU]; Sevier Co. Fighting Creek, Great Smoky Mts. 7.vii.1958, 35.67833°N, 83.555°W, J.L. Laffoon[pinned 1 ♂; ISIC]. TEXAS. Bastrop Co. Bastrop St. Park, 20.iii.1976, E.M. May[pinned 1 ♂; KU]; Nacogdoches Co. Nacogdoches, 12.x.1960, coll. R. Oliver, multiple dates 8.iv.1960– 6.v.1962[pinned 7 ♂, 1 ♀; KU]. VERMONT Orleans Co. 5 mi. NNE. Orleans, 19.vi.1982, coll. G.W. Byers[pinned 1 ♀; KU]. VIRGINIA. Arlington Co. Alexandria, 15.vi.1973, coll. B.M. O’Connor[pinned 1 ♀; UMMZ]; Fairfax Co. Lake Accotink, 5.viii.1972, coll. B.M. O’Connor[pinned 1 ♀; UMMZ]; Giles Co. Mt. Lake Biol. Stn. 6–12.viii.1979, coll. G.W. Byers#163 [pinned 1 ♂; KU]; Prince William Co. Woodbridge, 28.viii.1946, coll. L.D. Beamer[pinned 1 ♂; KU]. WISCONSIN. Vilas Co. Squaw Lake, 16.viii.1954, coll. J.L. Laffoon[pinned 2 ♂; ISIC]. WYOMING. Albany Co. Medicine Bow Nat. For. North Laramie Riverat Sand Lake Rd. Beaver Dam, 21.vi.2012 41.351557°N, 106.166511°W, coll. A Fasbender[pinned 1 ♀; ISIC]. Taxonomic notes.This species can be easily differentiated from its congener Bittacomorpha occidentalisby the medial white stripe on the scutum, shortened and narrowed clypeus, and paramere with lateral spur and apical dorsal tooth. Larvae can be distinguished in dorsal view, the head capsule of Bi. clavipesbeing ovoid and that of Bi. occidentalistriangular. The holotype of Bittacomorpha clavipesis not listed in the collection of Fabricius type material at ZMUC, and may be lost or destroyed. Fortunately, the distinctive character of the inflated 1 st tarsomereallows the determination of the genus without contention, and material from the range of Bittacomorpha occidentaliswould not have been available to Fabricius at the time.
Distribution. Fig. 18a.
2556241568
1991-02-22
INHS
D. W. Webb & M. E. Irwin
Costa Rica
Bijagua
Bijagua
31
32
1
1
GUANACOSTE
1586248125
1978-05-25
KU
G. W. Byers & C. W. Young
United States of America
Hale Co.
SE. of Tuscaloosa
Payne Lake
31
32
2
1
1
ALABAMA
2556241576
1978-05-26
KU
G. W., Byers & C. W. Young
United States of America
SW. of Auburn
Lee Co. Chewacia State Park
31
32
1
1
ALABAMA
2556241570
1978-05-22
KU
G. W. Byers & C. W. Young
United States of America
Hot Spring Co.
City
Malvern
31
32
3
3
ARKANSAS
1586248136
1978-05-22
KU
G. W. Byers & C. W. Young
United States of America
Montgomery Co.
SE. of Joplin
Gap Creek
31
32
1
1
ARKANSAS
2556241581
1984-06-01
KU
S. A. Teale
United States of America
Gap Creek
31
32
1
1
ARKANSAS
2556241583
[498,1262,1531,1556]
1984-06-01
KU
S. A. Teale
United States of America
Mount Ida
31
32
1
1
ARKANSAS
2556241551
1974-07-13
KU
G. W. Byers & C. W. Young
United States of America
Fremont Co.
Coaldale
Hayden Creek
31
32
9
1
8
COLORADO
2556241571
[216,1078,1603,1628]
1952-07-23
UMMZ
B. Pruitt
United States of America
Gothic
Gunnison Co.
31
32
7
1
6
COLORADO
2556241584
1981-07-25
KU
G. W. Byers & S. A. Teale
United States of America
Mineral Co.
San Juan Natl. Forest
Wolf Creek
31
32
3
3
COLORADO
2556241543
1965-08-13
KU
G. W. Byers
United States of America
San Juan Co.
Durango
31
32
1
1
COLORADO
2556241574
WSU
United States of America
31
32
1
1
COLORADO
1586248158
[246,1432,1711,1736]
1928-08-02
USNM
A. L. Melander
United States of America
Fairfield Co. Redding
Fairfield Co. Redding
31
32
1
1
CONNECTICUT
1586248179
[151,1159,1747,1772]
1973-05-06
FMNH
United States of America
Butts Co.
Butts Co.
31
32
2
2
GEORGIA
1586248156
1965-05-15
KU
B. Harrison
United States of America
East Point
Fulton Co.
31
32
1
1
GEORGIA
1586248141
1967-06-23
KU
G. W. Byers
United States of America
De Soto Falls
Lumpkin Co.
31
32
2
2
GEORGIA
1586248137
[286,1231,1819,1844]
1977-10-03
KU
N. Watkinsville & G. W. Byers
United States of America
Oconee Co.
31
32
2
2
GEORGIA
2556241552
1967-06-23
KU
G. W. Byers
United States of America
Union Co.
Lake Winfield Scott
Union Co. Lake
31
32
1
1
GEORGIA
2556241572
1909-06-20
FMNH
W. J. Gerhard
United States of America
Cook Co.
Palos Park
Palos Park
31
32
1
1
ILLINOIS
2556241559
1974-09-11
FMNH
D. Pearson
United States of America
Dupage Co.
Argonne Natl. Lab
31
32
1
1
ILLINOIS
1586248161
1972-07-23
1973-07-27
1972-07-23
FMNH
D. Pearson
United States of America
Kane Co.
31
32
6
4
2
ILLINOIS
1586248147
[256,1169,1963,1988]
1928-09-04
1955-09-02
1928-09-04
UMMZ
B. E. M.
United States of America
Lagrange Co.
Pigeon River
Pigeon River
31
32
1
1
1
INDIANA
1586248132
1955-09-02
1955-09-02
1928-09-04
FMNH
H. Dybas
United States of America
LaPorte Co.
Michigan City
Lagrange Co. Pigeon River
31
32
1
1
1
INDIANA
2556241586
1959-07-18
ISIC
J. L. Laffoon
United States of America
Dickinson Co.
32
33
Iowa Lakeside Lab
31
32
1
1
IOWA
2556241541
2010-05-17
ISIC
G. W. Courtney & A. Fasbender
United States of America
43.383373
Hillside Seep
1
-95.17998
Iowa Lakeside Lab
32
33
2
2
IOWA
1586248142
2010-05-17
ISIC
G. W. Courtney & A. Fasbender
United States of America
43.437473
Silver Lake
1
-95.365005
Silver Lake
32
33
31
1
12
3
15
IOWA
2556241534
1972-08-24
ISIC
A. Stewart Cott
United States of America
Guthrie Co. Springbrook State Park
Guthrie Co. Springbrook State Park
32
33
2
1
1
IOWA
2556241573
1960-07-19
ISIC
J. Laffoon
United States of America
32
33
1
1
IOWA
1586248145
[299,1143,295,320]
1964-05-17
ISIC
W. S. Craig
United States of America
Ames
Story Co.
32
33
45
2
43
IOWA
2556241565
1987-07-08
KU
J. Gelhaus
United States of America
Douglas Co.
Snow Hall
Lawrence
32
33
1
1
KANSAS
2556241553
[880,1433,332,357]
1986-05-19
KU
B. N. Danforth
United States of America
32
33
1
1
KANSAS
2556241540
[151,1169,367,392]
1936-08-05
A. E. Brower
United States of America
Hancock Co.
Bar Harbor
32
33
2
2
MAINE
1586248152
1967-09-12
1981-08-18
1967-09-12
USNM
P. J. Spangler
United States of America
Ellsworth
Penobscot Co.
32
33
1
1
1
MAINE
1586248157
[631,1432,404,429]
1981-08-18
1981-08-18
1967-09-12
KU
S. Struad
United States of America
Orono
Penobscot Co.
32
33
1
1
1
MAINE
1586248146
[151,1073,439,464]
1979-07-05
KU
G. W. Byers
United States of America
Calhoun Co.
Battle Creek
32
33
1
1
MICHIGAN
2556241546
WSU
United States of America
Battle Creek
Battle Creek
32
33
8
5
3
MICHIGAN
2556241569
[557,1431,476,501]
1957-07-19
KU
R. E. Beer
United States of America
Cheboygan Co.
Cochran Lake
Cochran Lake
32
33
1
1
MICHIGAN
2556241544
[151,883,511,536]
1961-07-08
KU
H. Willis
United States of America
Topinbee
32
33
1
1
MICHIGAN
2556241585
[895,1347,511,536]
1931-07
KU
United States of America
32
33
1
1
MICHIGAN
1586248165
1969-07-15
UMMZ
W. Pellston
United States of America
Emmet Co.
Emmet Co.
32
33
5
2
3
MICHIGAN
1586248174
1959-07-13
UMMZ
V. Potter
United States of America
Ithaca
Gratiot Co.
32
33
8
5
3
MICHIGAN
1586248151
[471,1191,583,608]
1939-05-07
UMMZ
F. Awnerman
United States of America
Jackson Co.
32
33
1
1
MICHIGAN
1586248176
1965-07-04
FMNH
B. Hlavac & T. Hlavac
United States of America
Kalamazoo Co.
Gull Lake
Gull Lake
32
33
1
1
MICHIGAN
1586248197
1951-08-14
UMMZ
J. S. Rogers
United States of America
Schoolcraft Co.
32
33
1
1
MICHIGAN
1586248168
1950-09-02
ISIC
J. L. Laffoon
United States of America
Clearwater Co.
Itasca State Park
Itasca State Park
32
33
7
1
6
MINNESOTA
1586248173
1978-05-23
KU
G. W. Byers & C. W. Young
United States of America
Lafayette Co.
Oxford
Puskus Lake
32
33
1
1
MISSISSIPPI
1586248159
[455,1266,727,752]
1977-09-23
KU
G. W. Byers
United States of America
Warren Co.
Vicksburg
32
33
1
1
MISSISSIPPI
2556241564
1950-06-09
1950-06-11
1950-06-09
ISIC
Hicks & Slater & J. L. Laffoon
United States of America
Cherry Co.
Valentine
Valentine
32
33
11
5
6
NEBRASKA
2556241556
[229,1267,799,824]
1967-07-12
Malaise Trap
WSU
R. W. Dawson
United States of America
Rushville
Sheridan Co.
32
33
1
1
NEBRASKA
2556241535
1925-08-16
FMNH
A. C. Weed
United States of America
Chenango Co.
Smyrna
32
33
1
1
NEW YORK
2556241554
1900
KU
O. S. Wescott
United States of America
Saranac Inn
Clinton Co.
32
33
5
5
NEW YORK
2556241577
[463,1430,871,896]
1956-08-26
W. E. LaBerge
United States of America
Long Lake
Hamilton Co.
32
33
2
2
NEW YORK
2556241560
[151,1329,908,933]
1927-06-17
FMNH
A. C. Weed
United States of America
North Rose
Wayne Co.
32
33
1
1
NEW YORK
2556241557
1962-04-29
ISIC
R. D. Peterson
United States of America
Buncombe Co.
Asheville
32
33
1
1
NORTH CAROLINA
1586248175
1967-06-15
KU
G. W. Byers
United States of America
Swain Co.
Clingman's Dome
32
33
1
1
NORTH CAROLINA
2556241531
1967-08-08
USNM
M. Duckenonod
United States of America
Portage Co.
Kent
Kent
32
33
1
1
OHIO
2556241539
1960-07-07
1960-07-10
1960-07-07
KU
G. W. Byers
United States of America
Berks Co.
French Creek Park
32
33
32
2
30
PENNSYLVANIA
2556241563
1956-08-15
KU
G. W. Byers
United States of America
Reading
32
33
4
1
3
PENNSYLVANIA
1586248138
1960-07-02
1979-07-08
1960-07-02
KU
G. W, Byers
United States of America
Franklin Co.
Caledonia St. Park
32
33
2
2
PENNSYLVANIA
1586248164
[467,1324,1124,1149]
1979-07-08
1979-07-08
1960-07-02
KU
G. W. Byers
United States of America
Luzerne Co.
Hazelton
32
33
1
1
PENNSYLVANIA
1586248169
1978-05-29
1978-05-30
1978-05-29
KU
S. Abbeville & G. W. Byers & C. W. Young
United States of America
Abbeville Co.
Abbeville Co.
32
33
3
1
2
SOUTH CAROLINA
2556241542
[151,1133,1196,1221]
1978-05-30
1978-05-30
1978-05-29
KU
G. W., Byers & C. W. Young
United States of America
Oconee Co.
Westminster
32
33
1
1
2
SOUTH CAROLINA
2556241575
1939-08-20
UMMZ
B. E. Montgomery
United States of America
Small lake
Oconee State Park
32
33
1
1
SOUTH CAROLINA
1586248200
2005-03-17
ISIC
G. W. Courtney & M. J. Petersen
United States of America
34.75
Clemson U.
718
-82.85
Pickens Co.
32
33
8
1
7
SOUTH CAROLINA
2556241548
2010-06-29
ISIC
United States of America
Pennington Co.
44.10962
Stream
1
-103.734566
Black Hills Nat. For.
32
33
3
3
SOUTH DAKOTA
1586248185
2008-06-30
ISIC
J. D. Petersen
United States of America
Custer Co.
43.718
French Creek Picnic
55
-103.4891
Custer State Park
32
33
1
1
SOUTH DAKOTA
1586248166
vi. 2
ISIC
J. D. Petersen
United States of America
43.8459
Custer State Park
6
-103.5602
32
33
1
1
SOUTH DAKOTA
1586248195
[227,1307,1447,1472]
1978-06-02
KU
G. W., Byers & C. W. Young
United States of America
Fentress Co.
Allardt
32
33
2
2
TENNESSEE
1586248202
1958-07-07
ISIC
J. L. Laffoon
United States of America
Sevier Co.
35.67833
Great Smoky Mts.
45
-83.555
Fighting Creek
32
33
1
1
TENNESSEE
1586248189
1976-03-20
KU
E. M. May
United States of America
Bastrop Co.
Bastrop St. Park
32
33
1
1
TEXAS
2556241567
1960-04-08
1962-05-06
1960-04-08
KU
R. Oliver
United States of America
Nacogdoches Co.
Nacogdoches
32
33
8
1
7
TEXAS
1586248178
1982-06-19
KU
G. W. Byers
United States of America
Orleans Co.
Orleans
Orleans
32
33
1
1
VERMONT
1586248198
1973-06-15
UMMZ
B. M. O'Connor
United States of America
Arlington Co.
Alexandria
32
33
1
1
VIRGINIA
1586248201
1972-08-05
UMMZ
B. M. O'Connor
United States of America
Fairfax Co.
Lake Accotink
32
33
1
1
VIRGINIA
1586248199
1979-08-06
1979-08-12
1979-08-06
KU
G. W. Byers
United States of America
Giles Co.
Mt. Lake Biol. Stn.
32
33
1
1
VIRGINIA
1586248191
[216,1153,1700,1725]
1946-08-28
KU
L. D. Beamer
United States of America
Prince William Co.
Woodbridge
32
33
1
1
VIRGINIA
1586248171
1954-08-16
ISIC
J. L. Laffoon
United States of America
Vilas Co.
Squaw Lake
Squaw Lake
32
33
2
2
WISCONSIN
2556241579
2012-06-21
ISIC
A Fasbender
United States of America
Albany Co.
41.35156
Sand Lake Rd. Beaver Dam
1
-106.16651
Medicine Bow Nat. For. North Laramie River
32
33
1
1
WYOMING