Orectochilus cardiophorus Régimbart Peschet 1923:125 Orectochilus salvazai Peschet Ochs 1930: 33 Orectochilus salvazai Ochs, 1940: 19 Review of the Patrus landaisi species group from Thailand with description of three new species (Coleoptera, Gyrinidae) Suksai, Benjamart Gustafson, Grey T. Sites, Robert W. Sangpradub, Narumon Zootaxa 2021 2021-06-25 4991 3 561 591 (Ochs, 1940) Ochs 1940 [151,498,1050,1077] Insecta Gyrinidae Patrus GBIF Animalia Coleoptera 21 582 Arthropoda species salvazai  ( Figs. 2G, 3G, 4G, 11, 13B, 13C, 13E, 13F, 14B)     Orectochilus cardiophorusRégimbart:  Peschet 1923:125(misidentification)    Orectochilus salvazaiPeschet:  Ochs 1930: 33( nomen nudum)     Orectochilus salvazai Ochs, 1940: 19(valid original description)     Typelocality.‘Sala-Nam-Pik’, Luang Prabang, Laos.  Material examined.   THAILAND: Chiang Mai Province: Doi Inthanon Natl.Park., stream from Huai Sai Luang Waterfall, 18°31’N 98°27’E,  1060 ma.s.l.,  8.V.2002, leg. UMC & CMU teams, L-396, 1 male( THNHM),  2 females( THNHM, UMC);  Doi Suthep Natl. Park., Monthathan Waterfall, 18°49’N 98°55’E,  700 ma.s.l.,  14.II.2003, leg. CMU team, 1 female( UMC);  Haow River, Sop Poeng Subdistrict, Mae Taeng District, 19°07’20.0”N 98°46’44.7”E,  505 ma.s.l.,  8.I.2018, leg. B. Suksai, C. Yatfung& S. Yoolong, 1 male( KKU).   Chiang Rai Province: Nam Kuen Stream, Ban Huai Nam Kuen, Mae Chedi Subdistrict, Wiang Pa Pao District, 19°07’22.3”N 99°19’43.3”E,  1327 ma.s.l.,  11.XI.2018, leg. B. Suksai, S. Kullasoot& P. Sapewisut, 1 male, 1 female( THNHM);  Streamclose to Ban Phithak Thai, Tap Tao Subdistrict, Thoeng District, 19°45’00.55”N 100°23’59.53”E,  1089 ma.s.l.,  2.V.2019, leg. B. Suksai, P. Sapewisut& N. Plangklang, 1 female( THNHM);  Streamclose to Ban Thai Samakkhi, Tap Tao Subdistrict, Thoeng District, 19°44’13.3”N 100°24’02.2”E,  1299 ma.s.l.,  28.V.2017, leg. P. Sapewisut, 1 male( KKU),  2 females( KKU, THNHM); same locality except for,  10.XI.2018, leg. B. Suksai, S. Kullasoot& P. Sapewisut, 1 male, 2 females( THNHM); same locality except for,  2.V.2019, leg. B. Suksai, P. Sapewisut& N. Plangklang, 8 males( KKU, THNHM),  2 females( GTGC).   Nan Province: Doi Phuka Natl. Park., creek at Namtok Ton Tong, 19°12’N 101°04’E,  900 ma.s.l.,  13.III.2002, leg. CMU teams, 1 male, 1 female( UMC);  Khun Sathan Natl. Park., Ban Khun Sathan, San Ta Subdistrict, Na Noi District, 18°16’24.12”N 100°29’10.80”E,  1474 ma.s.l.,  30.V.2017, leg. P. Sapewisut, 3 females( KKU, THNHM); same locality except for, Ban Mueng Li, Mueng Li Subdistrict, Na Muen District, 18°15’08.44”N 100°29’35.77”E,  786 ma.s.l.,  29.V.2017, leg. P. Sapewisut, 1 male( KKU),  3 females( KKU, THNHM); same locality except for, Khun Sathan Watershed Management Station, San Ta Subdistrict, Na Noi District, 18°16’43.67”N 100°30’14.34”E,  1334 ma.s.l.,  30.V.2017, leg. P. Sapewisut, 1 male( THNHM); same locality except for, Watersupply Weir, Khun Sathan National Park, San Ta Subdistrict, Na Noi District, 18°16’42.06”N 100°30’26.95”E,  1356 ma.s.l.,  30.V.2017, leg. P. Sapewisut, 2 males, 2 females( KKU, THNHM);  Mae Charim Natl. Park., Huay Taostream, Beside of Ban Nampang-Romklao Road, close to Huay Tao Ranger Station, 18°35’01.33”N 101°03’52.52”E,  898 ma.s.l.,  2.I.2013, leg. K. Rattanachan, 1 female( KKU).   Diagnosis.Size: Male: 12.0– 13.7 mm, female: 10.6–12.9 mm; body form in lateral view weakly dorsoventrally convex, more depressed anteriorly than posteriorly; elytral glabrous region of male ( Figs. 2G, 11A) cordiform, along suture extending ca. 7/9 length of elytra, laterally expanded in anterior 2/3 to similar width as pronotal glabrous region, posterior 1/3 acuminate with weakly rounded margins; female elytral glabrous region ( Figs. 3G, 11E) similar to male, except lateral expansion of glabrous region is stronger, surpassing lateral margin of pronotal glabrous region, apical acumination weaker; elytral apex of both sexes similar, weakly rounded in appearance, without apicolateral sinuation, epipleural angle indistinct; male protibia ( Fig. 4G) cleaver shaped, constricted in basal 1/2, protibial apex truncate with prominent distolateral angle; male protarsus ( Fig. 4G) ca. 1/2 length of protibia, protarsomeres width weakly decreasing apically; mesoventrite without medial pit; aedeagus ( Fig. 11B– D) with median lobe constricted medially, apex with rounded lateral margins, tip papilliform, in lateral view tip dorsally up-turned ( Fig. 11C); female reproductive tract ( Fig. 11G) without additional sclerotized structure between gonocoxae ( Fig. 11F), fertilization duct as in Fig. 11H.  Differential diagnosis.  Patrus salvazaiis most similar to  P. phetchabunensis  sp. nov., but can be distinguished from it by the characters given in the differential diagnosis of the latter.   Distribution.This species was originally described from Luang Prabang Provincewith several additional records from Laos( Peschet 1923; Ochs 1930). Ochs (1930)reported the first records of  P. salvazaifrom Doi Prae Muang mountain in Chiang Mai Province, Thailand. Here we add additional localities for  P. salvazaiacross the mountains of northern Thailandin Chiang Raiand NanProvinces ( Fig. 14B). In Thailand,  P. salvazaihas been found coexisting with  P. apicalisin streams at high elevations ( Fig. 13B) as well as with  P. landaisifrom a shallow stream with stagnant water or with slow current ( Fig. 13C). These two species were also found in puddles in a drying stream bed ( Fig. 13E–F).   Discussion.The name  Orectochilus salvazaiwas first published by Ochs (1930)in his catalog of Indian insects, for specimens that Peschet (1923)misidentified as  Orectochilus cardiophorus( Régimbart, 1888). This renders  Orectochilus salvazai Ochs, 1930a nomen nudumas it was not accompanied by a description. Ochs (1940, p. 19), explains later that he attributed the name  O. salvazaito Peschet in both his prior checklist (1930) and present work (1940) because, subsequent to the aforementioned misidentification ( Peschet, 1923), Peschet had recognized that these specimens actually belonged to a new species he dubbed “  salvazai” and had provided Ochs specimens identified as such. Because Peschet had not previously published a description of  P. salvazai,when Ochs (1940)used the name  Orectochilus salvazaiand included both a description of the species, a differential diagnosis, and a mention of examined material, the name was made available and is attributable to him following Article 13.1 of The International Code of Zoological Nomenclature( ICZN 1999), not Peschet. A similar situation surrounds the name  Orectochilus peguensis( Ochs, 1940), another member of the  P. landaisigroup (see checklist here below), in that its first use is in Ochs’ 1930 catalog as  O. cardiophorus peguensis(a nomen nudum) attributed to Peschet with the name being made available subsequently by the description given in Ochs (1940, p. 17) and is thus attributed to Ochs 1940, not Peschet. [151,750,1281,1308] Laos Luang Prabang 21 582 1 Louangphabang holotype 2002-05-08 THNHM Thailand 1060 18.516666 Huai Sai Luang Waterfall 1275 98.45 Doi Inthanon Natl. Park. 21 582 1 1 Chiang Mai 2002-05-08 THNHM, UMC Thailand 1060 true 18.516666 Huai Sai Luang Waterfall 1275 98.45 Doi Inthanon Natl. Park. 21 582 2 2 Chiang Mai 2003-02-14 UMC Thailand 700 18.816668 Monthathan Waterfall 1274 98.916664 Doi Suthep Natl. Park. 21 582 1 1 Chiang Mai 2018-01-08 KKU B. Suksai & C. Yatfung & S. Yoolong Thailand Haow River 505 19.122223 Mae Taeng District 1 98.77908 Sop Poeng Subdistrict 21 582 1 1 Chiang Mai 2018-11-11 THNHM B. Suksai & S. Kullasoot & P. Sapewisut Thailand Nam Kuen Stream 1327 19.122862 Mae Chedi Subdistrict 1 99.3287 Ban Huai Nam Kuen 21 582 2 1 1 Chiang Rai 2019-05-02 THNHM B. Suksai & P. Sapewisut & N. Plangklang Thailand Stream 1089 19.750153 Tap Tao Subdistrict 1 100.399864 Ban Phithak Thai 21 582 1 1 Chiang Rai 2017-05-28 KKU P. Sapewisut Thailand Stream 1299 19.737028 Tap Tao Subdistrict 1 100.40061 Ban Thai Samakkhi 21 582 1 1 Chiang Rai [312,635,1677,1704] 2017-05-28 KKU, THNHM P. Sapewisut Thailand Stream 1299 true 19.737028 Tap Tao Subdistrict 1 100.40061 Ban Thai Samakkhi 21 582 2 2 Chiang Rai 2018-11-10 THNHM B. Suksai & S. Kullasoot & P. Sapewisut Thailand Stream 1299 true 19.737028 Tap Tao Subdistrict 1 100.40061 Ban Thai Samakkhi 21 582 3 2 1 Chiang Rai 2019-05-02 KKU, THNHM B. Suksai & P. Sapewisut & N. Plangklang Thailand Stream 1299 true 19.737028 Tap Tao Subdistrict 1 100.40061 Ban Thai Samakkhi 21 582 8 8 Chiang Rai [592,805,1749,1775] 2019-05-02 GTGC B. Suksai & P. Sapewisut & N. Plangklang Thailand Stream 1299 true 19.737028 Tap Tao Subdistrict 1 100.40061 Ban Thai Samakkhi 21 582 2 2 Chiang Rai 2002-03-13 UMC Thailand 900 19.2 Namtok Ton Tong 1273 101.066666 Doi Phuka Natl. Park. 21 582 2 1 1 Nan 2017-05-30 KKU, THNHM P. Sapewisut Thailand Khun Sathan Natl. Park. 1474 18.273367 San Ta Subdistrict 1 100.48633 Ban Khun Sathan 21 582 3 3 Nan 2017-05-29 KKU P. Sapewisut Thailand Ban Mueng Li 786 18.252344 Na Muen District 1 100.49326 Mueng Li Subdistrict 21 582 1 1 Nan [246,565,1929,1956] 2017-05-29 KKU, THNHM P. Sapewisut Thailand Ban Mueng Li 786 true 18.252344 Na Muen District 1 100.49326 Mueng Li Subdistrict 21 582 3 3 Nan 2017-05-30 THNHM P. Sapewisut Thailand Khun Sathan Watershed Management Station 1334 true 18.278797 Na Noi District 1 100.50398 San Ta Subdistrict 21 582 1 1 Nan 2017-05-30 KKU, THNHM P. Sapewisut Thailand Water 1356 true 22 583 18.278349 Khun Sathan National Park 1 100.507484 Weir 21 582 4 2 2 Nan 2013-01-02 KKU K. Rattanachan Thailand Mae Charim Natl. Park. 898 18.583704 Beside of Ban Nampang-Romklao Road 1 101.06459 Huay Tao 22 583 1 1 Nan