Cicadas (Hemiptera: Cicadidae) from Camiguin of Mindanao Province and Dinagat Island in the Philippines, with a new genus and three new species
Lee, Young June
Marshall, David C.
Mohagan, Alma
Hill, Kathy B. R.
Zootaxa
2016
4097
2
187
202
8HBWN
Buckton, 1889
Buckton
1889
[226,551,1907,1933]
Insecta
Cicadidae
Animalia
Hemiptera
8
195
Arthropoda
tribe
Cicadettini
Remarks."Huechysaria" (= Huechysini Distant, 1905) was proposed by Distant (1905b)with Huechys Amyot & Audinet-Serville, 1843as the typegenus mainly by the opaque or semiopaque wings. However, the molecular phylogenetic study by Marshall et al. (2015)suggests that Huechysshould be placed within Cicadettini. This suggested placement is well supported morphologically by the presence of a pair of flat upper lobes, a welldeveloped dorsal beak, a duck-bill-shaped median lobe of uncus, and a pair of clasper-like processes, which are commonly seen in the male genitalia of Huechysas well as that of the typegenus of Cicadettini, CicadettaKolenati, 1857. Huechysalso has the following characteristics, which were listed as distinguishing characters of Cicadettiniby Moulds (2005): pygofer upper lobes flat; and pseudoparameres present (see Fig. 4). Here, Huechysini syn. nov.is synonymized with Cicadettini. Moulds (2005)also proposed as a diagnostic character for Cicadettinithe timbals not extending ventrally beyond the wing bases although he later found the opposite character-state in UradolichosMoulds, 2012, which he placed in Cicadettini. Because the timbals slightly extend below the wing bases ventrally in specimens of Huechys parvula Haupt, 1924listed below and some examined specimens of KosemiaMatsumura, 1927and TettigettaKolenati, 1857in the first author's care, this character may have to be excluded from the characters for diagnosing Cicadettini.