New species and records of heterostigmatic mites (Acari: Prostigmata: Heterostigmata) phoretic on scarabaeid dung beetles (Coleoptera: Scarabaeidae) from northwestern Iran Sobhi, Mohammad Hajiqanbar, Hamidreza Mortazavi, Azim Zootaxa 2017 4276 3 427 434 Lombardini 1949 [572,671,1715,1741] Arachnida Dolichocybidae Pavania GBIF Animalia Actinedida 4 431 Arthropoda genus    1. Prodorsum with three pairs of setae ( sc1 absent)...................................... (  gymnopleurispecies group) 2  - Prodorsum with four pairs of setae (including sc1)............................................................ 4    2. Setae   v1longer than distance between their bases; setae elonger than f; setae h2 15 times longer than h1 ( Iran, phoretic on Gym- nopleurus  mopsus persianusReitter)..................................  P. gymnopleuri Hajiqanbar & Khaustov, 2010  - Setae  v1shorter than distance between their bases; setae enever longer than f; setae h2 at most seven times longer than h1 … 3    3. Genu Iwith one seta ( v'); dorsal idiosomal setae smooth; setae c1 longer than c2; setae c1 and dpointed ( Iran, phoretic on  G. mopsus persianus)................................................  P. sabzevarensis Hajiqanbar & Khaustov, 2010  - Genu I with two setae ( v', l'); dorsal idiosomal setae weakly barbed; setae c2 longer than c1; setae c1 and ddistinctly blunt-ended  ( Iran, phoretic on  Onthophagus gibbosus gibbosus(Scriba),  O. taurus(Schreber)and  O. amyntas alces(Fabricius)).........................................................................  P. onthophagi Hajiqanbar & Khaustov, 2010  4. Setae sc1 seta-like ( Iran, phoretic on  O. vitulusFabricius)................................................................................................... (  setiformisspecies group),  P. setiformisLoghmani & Hajiqanbar, 2013  - Setae sc1 as capitate trichobothria....................................................(  fusiformisspecies group) 5    5. Some setae on tarsi I ( pv’, pv”, u’, u”) and II ( pv’, pv”, u’, tc”) modified, lanceolate ( Iran, phoretic on  Scarabaeus pius(Illiger))..........................................................  P. lanceolata Bahramian & Hajiqanbar, 2015  - Setae on tarsi I-II not modified, seta-like................................................................... 6    6. Setae h2 less than three times longer than h1; setae h1 longer than e( Ukraine, Slovakia, phoretic on  Copris lunarisL.).....................................................................................  P. riparia Sevastianov, 1980  - Setae h2 four to five times longer than h1; setae h1 shorter than or subequal to e..................................... 7   7. Setae eas long as f; setae on epimeres II subequal (Tajikistan, phoretic on  Onthophagussp.)....................................................................................................  P. tadjikistanica Sevastianov, 1980  - Setae flonger than e; setae on epimeres II not subequal........................................................ 8    8. Setae fslightly (at most 1.35 times) longer than e; distance d -dtwice as long as setae d( Iran, phoretic on  Euoniticellus fulvus(Goeze))............................................................  P. khiavensisSobhi & Hajiqanbar sp. nov.  - Setae fat least twice longer than e; distance d -ddistinctly more than two times longer than setae d..................... 9    9. Setae c2 longer than c1; setae sc2 more than two times longer than c1 ( Italy, phoretic on  Scarabaeus sacerL.)..........................................................................................  P. fusiformisLombardini, 1949  - Setae c1 longer than c2; setae sc2 less than two times longer than c1 (Iran, phoretic on  Scarabaeusspp.).......................................................................................  P. kamalii Hajiqanbar & Khaustov, 2010 1563872044 v 1 Iran Iran 4 431 1 1563872035 Genu I Iran Iran 4 431 1 1563872046 Iran Iran 5 432 1 1563872037 Iran Iran 5 432 1 1563872039 Ukraine Ukraine 5 432 1 1563872038 Iran Iran 5 432 1 1563872042 Italy Italy 5 432 1