Calliaxina Ngoc-Ho, 2003: 493–494 Sakai, 2011: 497–498 Calliaxina Paraglypturus Türkay and Sakai, 1995 Calliax Eucalliax Manning and Felder, 1991 Calliaxina Calliax Calliaxina Eucalliaxiopsis Sakai, 2011 Calliaxina Eucalliaxiopsis C. bulimba C. kensleyi C. novaebritanniae C. sakaii C. aequimana C. inaequimana C. panglaoensis C. jonesi C. punica Calliaxina C. cearaensis Eucalliaxiopsis A new classification of Callianassidae and related families (Crustacea: Decapoda: Axiidea) derived from a molecular phylogeny with morphological support Poore, Gary C. B. Dworschak, Peter C. Robles, Rafael Mantelatto, Fernando L. Felder, Darryl L. Memoirs of Museum Victoria 2019 Mem. Mus. Vic. 2019-12-31 78 73 146  Ngoc-Ho, 2003: 493–494  Nguyen Ngoc-Ho 2003 [817,929,1382,1404] Malacostraca Callianassidae Calliaxina GBIF Animalia Decapoda 53 126 Arthropoda genus     Calliaxina  Ngoc-Ho, 2003: 493–494.—  Sakai, 2011: 497–498.— Hyžný, 2012: 55–56.— Sakai and Türkay, 2014: 191.— Ngoc-Ho, 2014:  549.— Poore and Dworschak, 2017: 120(partim).— Sakai,2018: 739–741.  Calliax.— Sakai, 2005b: 196(partim, not  Calliaxde Saint Laurent, 1973).  Calliamina Sakai and Türkay, 2014: 190(misspelling).   Typespecies.  Calliax punicade Saint Laurent and Manning, 1982, by original designation.  Diagnosis. Cardiac sulci present. Pleomere 1 with pair of sternal plates anterior to pleopods 1. Maxilliped 3 ischium and merus broad, less than twice as long as wide at their articulation. Both chelipeds with palms as long as wide in both sexes, minor slightly smaller if at all. Male pleopod 1article 1 with distal long setae; article 2 blade-like with bifid apex (sometimes obscure), orarticle 2 much shorter, digitiform (sometimes fused); with appendix interna (present at least as hooks on endopod). Male pleopod 2 appendix interna digitiform; appendix masculina a lobe fused to appendix interna attached midway on endopod margin. Telsonwidest anteriorly, more or less semicircular, curving to broad convex apex.  Remarks.  Calliaxinadiffers from  Eucalliaxiopsisin having a tapering telson without a transverse ridge, and at least a vestige of an appendix interna on pleopod 1 of the male. All species have an exopod on maxilliped 3, sometimes vestigial, whereas only one species of  Eucalliaxiopsisdoes.    Calliaxinawas erected by Ngoc-Ho (2003)for three species previously included by Sakai (1999a)in  Paraglypturus Türkay and Sakai, 1995and later (Sakai, 2005) in  Calliax. Ngoc-Ho (2003)tabulated differences between these genera but did not include  Eucalliax Manning and Felder, 1991, which she diagnosed separately in her table. Her diagnosis relied importantly on the presence of an exopod on maxilliped 3, equal and similar chelipeds, appendix interna on pleopods 1 and 2 of the male. Some of her key features are difficult to quantify, notably the shape of the front and anterolateral projections of the carapace, and length of the appendix interna on pleopods 3–5. Sakai (2005) synonymised  Calliaxinawith  Calliaxwithout discussing any morphological similarities or differences. Later, Sakai (2011)revived  Calliaxinawhich he had earlier synonymised in new sense (sensu nov.) by including other species with similar maxillipeds 3 and with one or two cardiac sulci, although some lack maxillipedal 3 exopods. At the same time, he erected a monotypic genus  Eucalliaxiopsis Sakai, 2011, relying on the possession of unique male pleopods (see below). As already explained, Robles et al. (in press) divided the species allocated by Sakai (2011)to  Calliaxinaand  Eucalliaxiopsisinto two groups. Their molecular analysis grouped  C. bulimba, C. kensleyi,  C. novaebritanniaeand  C. sakaiiinto a one clade, and  C. aequimana,  C. inaequimana,  C. panglaoensis,  C. jonesiand three others into a second clade. The morphological treatment includes  C. punica, typespecies of  Calliaxinain the first and  C. cearaensis, typespecies of  Eucalliaxiopsisin the second. These two names are applied to these clades.