Hydraenidae of Madagascar (Insecta: Coleoptera) Perkins, Philip D. Zootaxa 2017 2017-11-03 4342 1 1 264 3N3FN Perkins, 2017 Perkins 2017 [151,408,152,178] Insecta Hydraenidae Hydraena GBIF Animalia Coleoptera 115 116 Arthropoda species pilotumida sp. nov.     TypeMaterial. Holotype(male):  Fianarantsoa, 7 kmW Ranomafana, from seepage over sunlit rock at river, montane rainforest, elev.  900 m, 21° 16' S, 47° 25' E,  6 ix 1993, W. E. Steiner& M. Stebbins( USNM).   Paratypes(94): Samedata as holotype(5 USNM); Samedata except W. E. Steiner& J. E. Cadle(6 USNM);   Fianarantsoa, 7 kmW Ranomafana, on seepage over exposed rocks at bank of Namorona River, elev.  900 m, 21° 16' S, 47° 25' E,  1–7 iii 1990, W. E. Steiner(10 USNM);  Namorona River, 7 kmW Ranomafana, from semi-shaded temporary pond in montane rain forest, elev.  1100 m, 21° 15' S, 47° 25' E,  8–21 x 1988, W. E. Steiner(5 USNM);  Namorona River,  7 kmW Ranomafana, on seepage over exposed rock at bank of Namorona River, elev.  900 m, 21° 16' S, 47° 25' E,  1–7 xi 1988, W. E. Steiner(2 USNM); Ranomafana, Sakaroa R., CascadeRiana, elev.  994 m, 21° 16' S, 47° 25' E,  1 xi–30 xii 2004, M. Balke(MD 030) (15 ZSMC); Vatovavy-Fitovinany, Ranomafana NP, Shakaroawaterfall, 2kmfrom park entrance, 21.2631S, 47.4132E, waterfall and hygropetric rock with some dead leaves, 21° 16' S, 47° 25' E,  4 xi 2014, P. D. Perkins( MAD14-13) (1 MCZ);   Toamasina, 18.8282S 48.4398E Alaotra MangoroAndasibe- MantadiaNP, Mantadiawaterfall Beanamalao, 7kmN of park entrance "circuit chute sacre", waterfall with hygropetric rocks, elev.  990 m, 18° 49' S, 48° 26' E,  28 xi 2014, J. Bergsten, R. Bukontaite, J.H. Randriamihaja, T. Ranarilalatiana & S. Holmgren ( MAD14-84) (50 NHRS).   Differential Diagnosis.One of the larger members of the group (ca. 2.02 mm). Males differentiated from others by the form of the metatibiae: slightly arcuate, medial margin with a large brush of setae that begins slightly before midlength and continues almost to the apex. The habitus is similar to that of  H. pilobova, but the male metatibiae and the sixth ventrite differ in the two species. The aedeagi also show a relationship, but differ in shapes of several parts ( Figs. 105, 106).   Description. Size: holotype(length/width, mm): body (length to elytral apices) 2.02/0.84; head width 0.47; pronotum 0.48/0.60, PA 0.47, PB 0.53; elytra 1.29/0.84. Dorsum dark brown, head darkest; legs brown to dark brown; maxillary palpi brown to light brown, distal ½ of last palpomere darker. Head and pronotum densely moderately coarsely punctate. Pronotal punctures larger than those of head; interstices varying from narrow ridges anteriorly and posteriorly to ca. 1xpd on disc, shining. Mentum sparsely moderately finely punctulate, very weakly shining; postmentum rugulose, dull. Genae roundly raised, smooth and shining, without posterior ridge. Pronotum bisinuate laterally, distinctly emarginate between anterior angle and widest part of pronotum; anterior margin straight behind eyes, emarginate behind frons; PF1 very shallow; PF2 shallow, very shallowly confluent medially forming very weak U-shaped impression; PF3 moderately deep; PF4 very shallow, nearly absent. Elytra weakly arcuate laterally; summit of posterior declivity near distal 2/3; lateral explanate margins moderately narrow; serial punctures round, separated by ca. 1xpd, slightly larger than pronotal punctures, some punctures random near scutellum. Intervals very weakly raised, not forming distinct unbroken straight lines, shining, ca. 1xpd. Apices in dorsal aspect broadly conjointly rounded, in posterior aspect margins forming moderately deep angle with one another. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1.5/2.2/1. P1 laminate; median carina sinuate in profile. P2 l/w ca. 2/1, sides parallel, apex blunt and raised slightly above metaventral intercoxal process. Plaques large, narrowly separated, widest at anterior 1/3, anteriorly and posteriorly rounded, more sharply so posteriorly, converging slightly anteriorly, not raised, located at sides of median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at arcuate posterior margin ca. 2xP2. Protibia very weakly arcuate, almost straight, slightly widened at ca. distal ¼ and here bearing on medial surface row of short spines, one larger than the others. Mesotibia straight. Metatibia slightly arcuate, medial margin with large brush of setae that begins slightly anterior to midlength and continues almost to apex. Metafemur lateral margin strongly arcuate, medial margin strongly sinuate. Abdominal apex with deep apicomedian concavity, upper margin bearing short setae and lower margin asymmetrical, with small produced point off center. Ventrite 6 with pair of low submedian tumidities that bear short setae.   Etymology.Named in reference to the pubescent tumidities of abdominal ventrite six. 1697157641 1993-09-06 USNM W. E. Steiner & M. Stebbins 900 -21.266666 Ranomafana 1264 47.416668 115 116 2 Fianarantsoa holotype 1697157654 USNM Same & W. E. Steiner & J. E. Cadle Same 115 116 2 Fianarantsoa paratype 1697157676 1990-03-01 1990-03-07 1990-03-01 USNM W. E. Steiner 900 -21.266666 Namorona River 1264 47.416668 Ranomafana 115 116 1 Fianarantsoa paratype 1697157652 1988-10-08 1988-10-21 1988-10-08 USNM W. E. Steiner 1100 -21.25 Ranomafana 1264 47.416668 Namorona River 115 116 1 Fianarantsoa paratype 1697157646 1988-11-01 2014-11-04 1988-11-01 USNM, ZSMC, MCZ W. E. Steiner & Ranomafana & Sakaroa R. & M. Balke & Vatovavy-Fitovinany & Ranomafana NP & Shakaroa & P. D. Perkins 900 -21.266666 Namorona River 1264 47.416668 Namorona River 115 116 MAD14-13 1 Cascade paratype 1697157648 2014-11-28 NHRS Alaotra Mangoro 990 -18.816668 Mantadia 1274 48.433334 Mantadia 115 116 MAD14-84 1 Toamasina paratype