Fairmaire 1898b : 468 Hydraenidae of Madagascar (Insecta: Coleoptera) Perkins, Philip D. Zootaxa 2017 2017-11-03 4342 1 1 264 3N394 Fairmaire Fairmaire [151,514,1837,1863] Insecta Hydraenidae Hydraena GBIF Animalia Coleoptera 108 109 Arthropoda species dilutipes     Hydraena dilutipes  Fairmaire 1898b: 468.   Type Material. Lectotype(male): Labels on lectotype: "Madag [illegible]; TYPE [red]; MUSEUM PARIS MADAGASCAR PERRIER DE LA BATHIE COLLECTION LEON FAIRMAIRE 1906; LECTOTYPE  Hydraena dilutipesFairmaire 1898, design. P. D. Perkins 2015[red]". Two female paralectotypes: 1) " Hydraena dilutipes Farm Madag; MUSEUM PARIS MADAGASCAR PERRIER DE LA BATHIE COLLECTION LEON FAIRMAIRE 1906; PARALECTOTYPE  Hydraena dilutipesFairmaire 1898, design. P. D. Perkins 2015[red]". 2) "Madag Perrier; MUSEUM PARIS MADAGASCAR PERRIER DE LA BATHIE COLLECTION LEON FAIRMAIRE 1906; PARALECTOTYPE  Hydraena dilutipesFairmaire 1898, design. P. D. Perkins 2015[red]" (MNHN).   FIGURE 98.  Hydraena dilutipes, lectotype, habitus dorsal, lateral, and ventral.    Specimensexamined (883): Antananarivo, 18.158S 47.2104E Analamanga, Manankazo riverby the bridge of RN4, medium size river over bedrock, elev.  1450 m, 18° 9' S, 47° 12' E,  21 xi 2014, J. Bergsten, R. Bukontaite, J.H. Randriamihaja, T. Ranarilalatiana& S. Holmgren( MAD14-75) (4 NHRS); Manakambahiny, in seepage over exposed granite cliff face, 18° 55' S, 47° 32' E,  12 i 1990, W. E. Steiner(28 USNM);   Antsiranana, 14.4345S 49.7606E Sava Marojejy NP Camp II, wide river forming hygropetric rocks, wide river forming hygropetric rocks and rockpools, elev.  710 m, 14° 26' S, 49° 45' E,  9 xi 2014, J. Bergsten, R. Bukontaite, J.H. Randriamihaja, T. Ranarilalatiana& S. Holmgren( MAD14-54) (3 NHRS);  14.4369S 49.7749E Sava, Marojejy NP,  300mN from Camp I, upstream, waterhole with dead leaves next to the stream, elev.  490 m, 14° 44' S, 49° 46' E,  8 xi 2014, P. D.   Perkins( MAD14-50) (10 MCZ);  14.4369S 49.7749E Sava MarojejyNP  300mN from CampI, upstream, waterhole with dead leaves next to the stream, elev.  490 m, 14° 26' S, 49° 46' E,  8 xi 2014, J. Bergsten, R. Bukontaite, J.H. Randriamihaja, T. Ranarilalatiana& S. Holmgren( MAD14-50) (22 NHRS);  14.5221S 49.7545E Sava, bridge 80kmfrom Sambava, where tributary river enters Lokoho river, rockpools and sandy waterpools at river margin, elev.  120 m, 14° 31' S, 49° 45' E,  17 xi 2014, J. Bergsten, R. Bukontaite, J.H. Randriamihaja, T. Ranarilalatiana& S. Holmgren( MAD14-73) (9 NHRS);  Antsaba, Galoko Mountains, hygropetric cascade, elev.  303 m, 13° 39' S, 48° 45' E,  1 xi–29 xii 2004, M. Balke(MD 016) (124 ZSMC; one male " MNCN AI-547 DNAspecimen"); Diana: Antsaba: Ambolamena mountain,  4kmSE Antsaba village,“Antohatea be Cascade”, S 13.67593 E48.75589, GB nets, sieves, aspirator, forceps: waterfall with hygropetric rocks, elev.  381 m, 13° 41' S, 48° 45' E,  27 xi 2012, J. Bergsten, R. Bukontaite, J.H. Randriamihaja& T. Ranarilalatiana( MAD12-29) (1 NHRS);  Diana: Antsaba: Galoko mountains, S 13.60974 E48.72175, sieves and aspirator: hygropetric rocks and water pools, elev.  263 m, 13° 37' S, 48° 43' E,  25 xi 2012, J. Bergsten, R. Bukontaite, J.H. Randriamihaja& T. Ranarilalatiana( MAD12-26) (1 NHRS);  Forêtd' Antsahabe, 11.4 km275° W Daraina, malaise trap, tropical dry forest, elev.  550 m, 13° 12' S, 49° 33' E,  12 xii 2003, B.L.Fisher(BLF 10117) (1 CAS);  Forêt d'Anabohazo, 21.6 km247° WSW Maromandia, malaise trap- in tropical dry forest, elev.  120 m, 14° 18' S, 47° 54' E,  11–16 iii 2001, Fisher, Griswold et al.(BLF 3336) (1 CAS);  MarojejyNP, camp 2, hygropetric habitat, 14° 26' S, 49° 46' E,  30 x 2007, J. Stastny(1 NHRS);  Montagne d'Ambre, Grande Cascade, elev.  740 m, 13° 34' S, 49° 10' E,  1 xi–30 xii 2004, M. Balke(MD 004) (20 ZSMC);  Sambava: MarojejyNP, Camp2 long waterfall along smooth rock area (hygropetric), N:-14.43400, E:49.75633, elev.  717 m, 14° 26' S, 49° 45' E,  7 xii 2006, Isambert et al.(P57 Bi12) (8 NHRS);  Sava, Marojejy NP,  800mN from Camp I, Humbertwaterfall; 14.4333S 49.773E, waterfall, elev. 550 m, 14° 26' S, 49° 46' E, 8–12 xi 2014, J. Bergsten, R. Bukontaite, J.H. Randriamihaja, T. Ranarilalatiana & S. Holmgren (MAD14-48) (12 NHRS); Marojejy NP, 800mN from Camp I, Humbert waterfall; 14.4333S 49.773E, ex. bedrock pools at side of waterfall, elev. 550 m, 14° 26' S, 49° 46' E, 8 xi 2014, P. D. Perkins (MAD14-48) (256 MCZ);   Atsinanana, NP Midongy, 23° 30' S, 46° 55' E,  17 viii 2011, (collector not given on label) (4 NHRS);   Fianarantsoa, 20.1711S 47.091E, Amoron'i, ManiaUpstream Talaviana riverfrom the crossing with RN7, 37km S of Antsirabe, rocky river with sandy bottom and some hygropetric rocks, elev.  1360 m, 20° 10' S, 47° 5' E,  1 xi 2014, P. D. Perkins( MAD14-01) (1 MCZ);  21.2581S 47.4213E, Vatovavy-Fitovinany, Ranomafana NP, Namorona riverby bridge at park entrance, rockpools and hygropetric sites by river, elev.  890 m, 21° 15' S, 47° 25' E, P. D. Perkins( MAD14-09) (37 MCZ);  7 kmW Ranomafana, from seepage over sunlit rock at river, montane rainforest, elev.  900 m, 21° 16' S, 47° 25' E,  6 ix 1993, W. E. Steiner& M. Stebbins(32 USNM); same locality, W. E. Steiner& J. E. Cadle(41 USNM);  7 kmW Ranomafana, on seepage over exposed rocks at bank of Namorona River, elev.  900 m, 21° 16' S, 47° 25' E,  1–7 iii 1990, W. E. Steiner(44 USNM);  Ionilahy(  Fianarantsoa), first left affl. of Riv. Ionilahyupstr. railroad bridge to Manakara,  200 m, 23.5°C, 0.088 mS/cm; hygropetric, elev.  200 m, 21° 43' S, 47° 38' E,  11 viii 2001, Gerecke & Goldschmidt (MD 20c) (4 NMW);  Ionilahy(  Fianarantsoa) Mahavavona, spring 3 Exp.NW, (hygropetric on rocks exposed to sun in rice field), elev.  284 m, 21° 42' S, 47° 38' E,  20 iv 2011, R. Gerecke(MD 218) (1 NMW);  Namorona River, 7 kmW Ranomafana, from semi-shaded temporary pond in montane rain forest, elev.  1100 m, 21° 15' S, 47° 25' E,  1–7 xi 1988, W. E. Steiner(55 USNM);  Namorona River, 7 kmW Ranomafana, on seepage over exposed rock at bank of Namorona River, elev.  900 m, 21° 16' S, 47° 25' E,  8– 21 x 1988, W. E. Steiner(38 USNM); Ranomafana, Namorona main river, rockpools, elev. 927 m, 21° 15' S, 47° 25' E, 1 xi–30 xii 2004, M. Balke (MD 029) (93 ZSMC);  Vatovavy Fitovinany: RanomafanaNP: Namorona riverby the bridge below park entrance: S 21.25809 E0 47.42165, GB Nets& sieves, rockpools, elev.  920 m, 21° 15' S, 47° 25' E,  1 xi 2011, J. Bergsten, R. Bukontaite, T. Ranarilalatiana, & J.H. Randriamihaja( MAD11-09) (9 NHRS);  Melaky: Tsingy de BemarahaNP. S 18.75875 E0 44.71648, elev.  112 m, 18° 46' S, 44° 43' E,  17 xii 2009, J. Bergsten, N. Jönsson, T. Ranarilalatiana, HJ. Randriamihaja( MAD09-68) (1 NHRS);   Toliara, Andhahela( Tulear), Isaka, stream exp. W 1 kmN from the vill., elev.  250 m, 24° 48' S, 46° 51' E,  7 ix 2001, Gerecke & Goldschmidt (MD 63) (1 NMW);  Ifaty, under seaweed drift on sand beach, 23° 9' S, 43° 37' E,  19 ix 1993, W. E. Steiner& R. Andriamasimanana(1 USNM);  Tsimelahy( Tulear), Riv. Antarantsa, ca. 1 kmupstream from village, elev.  300 m, 24° 56' S, 46° 38' E,  4 ix 2001, Gerecke & Goldschmidt (MD 58) (20 NMW).   Differential Diagnosis.Closely related to  H. dilutipoides, differing therefrom by the larger size (ca. 1.81 mmvs. 1.73), the more coarsely densely punctate pronotum, and having the elytral intervals strongly raised and forming unbroken lines. The plaques of  H. dilutipoidesare more widely separated, especially posteriorly, and more sharply rounded at the posterior edge. Males of both species have large metatibial brushes. The aedeagi also show a relationship, having a large, flat distal extension of the main piece; the complex gonopore bearing distal piece differs significantly in the two species ( Figs. 100–102).   Description. Size: Lectotype(length/width, mm): body (length to elytral apices) 1.81/0.76; head width 0.43; pronotum 0.42/0.60, PA 0.44, PB 0.53; elytra 1.11/0.76. Dorsum dark brown, head darkest; legs brown; maxillary palpi brown to light brown, distal ½ of last palpomere darker. Mentum sparsely finely punctulate, shining; postmentum rugulose, dull. Genae raised, weakly rugulose, dull, without distinct posterior ridge. Head and pronotum coarsely densely punctate, punctures of pronotum slightly larger than those of head; interstices narrow ridges, shining. Pronotum slightly sinuate laterally; anterior margin straight behind eyes, emarginate behind frons; PF1 very shallow; PF2 shallow, very shallowly confluent medially forming weak U-shaped impression; PF3 moderately deep; PF4 absent. Elytra weakly arcuate laterally; summit of posterior declivity near distal 2/3; lateral explanate margins moderately narrow; serial punctures deeply impressed, round, and closely spaced, slightly larger than pronotal punctures. Intervals raised, narrow, shining, ca. 0.5xpd. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming very shallow angle with one another. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/2/1.3. P1 laminate; median carina sinuate in profile. P2 l/w ca. 2/1, sides parallel, apex blunt, apex raised slightly above mesoventral intercoxal process. Plaques large, anteriorly and posteriorly rounded, parallel, anterior at level of surrounding cuticle, gradually elevating posteriorly such that posterior border is slightly raised above surrounding cuticle, located at sides of median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at slightly arcuate posterior margin ca. 1.5x P2. Protibia moderately wide, slightly arcuate on medial margin, slightly more so on lateral margin. Mesotibia straight; medial margin with 7–8 very short spines (microslide mount), margin not notched between spines. Metatibia slightly arcuate, distal ½ of medial margin slightly emarginate and with large brush of setae. Abdominal apex slightly asymmetrical; last tergite with apicomedian notch. Females average slightly smaller than males; male metafemur longer and more sinuately shaped than in female; female metatibia simple and slightly shorter than in male.   Remarks.Fairmaire (1898) in the original description notes the differences in the shape of the abdomen of the male and female. It is therefore unquestionable that he had both sexes; however, he does not indicate how many specimens total he had. The locality given is "Dans l'Ikopa"; Ikopa is the river that flows through Antananarivo. Although the river is quite long, ca. 300 km, the typespecimens were most likely collected in or very near the capital. Specimens were collected in the capital as late as 1990, by W. E. Steiner. However, these were not collected in the river, which is now quite polluted.  Trizzino et al.(2012)presented the DNA sequence information of one male specimen ("MNCN AI-547 DNA specimen") with this locality data: Antsaba, Galoko Mountains, hygropetric cascade, elev. 303 m, 13° 39' S, 48° 45' E, 1 xi–29 xii 2004, M. Balke (MD 016). 1697157591 1990-01-12 2014-11-21 1990-01-12 NHRS, USNM J. Bergsten & R. Bukontaite & J. H. Randriamihaja & T. Ranarilalatiana & S. Holmgren & W. E. Steiner Specimens 1450 -18.916666 Manankazo river 1273 47.533333 Analamanga 109 110 MAD14-75 1 Antananarivo 1697157633 2014-11-09 NHRS NP Camp II & J. Bergsten & R. Bukontaite & J. H. Randriamihaja & T. Ranarilalatiana & S. Holmgren 710 -14.433333 Sava Marojejy 1288 49.75 109 110 MAD14-54 1 Antsiranana 1697157607 2014-11-08 Marojejy NP & Camp I & Philip D. Perkins 300 -14.733334 Sava 1287 49.766666 109 110 1 Antsiranana 1697157589 [151,537,151,176] MCZ Perkins 110 111 MAD14-50 1 1697157604 2014-11-08 NHRS J. H. Randriamihaja & Ranarilalatiana & Holmgren 300 -14.433333 Camp 1288 49.766666 Sava Marojejy 110 111 MAD14-50 1 1697157580 2014-11-17 NHRS T. Ranarilalatiana & Holmgren Sava 120 -14.516666 Lokoho river 1287 49.75 Sambava 110 111 MAD14-73 1 1697157583 2004-11-01 2012-11-27 2004-11-01 ZSMC, MNCN, DNA, NHRS M. Balke & J. Bergsten & R. Bukontaite & J. H. Randriamihaja & T. Ranarilalatiana Antsaba 303 -13.683333 Ambolamena mountain 1290 48.75 Galoko Mountains 110 111 AI-547, MAD12-29 1 Cascade 1697157605 2012-11-25 NHRS J. H. Randriamihaja & Ranarilalatiana Diana 263 -13.616667 Galoko mountains 1290 48.716667 Antsaba 110 111 MAD12-26 1 Cascade 1697157610 2003-12-12 CAS B. L. Fisher Foret 550 -13.2 Daraina 1291 49.55 Antsahabe 110 111 1 Cascade 1697157599 2001-03-11 2001-03-16 2001-03-11 CAS 120 -14.3 Maromandia 1288 47.9 Foret d'Anabohazo 110 111 1 Cascade 1697157612 2007-10-30 NHRS J. Stastny -14.433333 Marojejy 1288 49.766666 110 111 1 Cascade 1697157624 2004-11-01 2004-12-30 2004-11-01 ZSMC M. Balke 740 -13.566667 Montagne d'Ambre 1290 49.166668 110 111 1 Cascade 1697157645 2006-12-07 NHRS Isambert & Bi Sambava 717 -14.433333 Camp 1288 49.75 Marojejy 110 111 1 Cascade 1697157623 [250,948,836,861] Marojejy NP & Camp I & Humbert 800 Sava 110 111 1 Cascade 1697157630 [238,1432,980,1005] 2011-08-17 NHRS NP Midongy -23.5 Atsinanana 1255 46.916668 110 111 1 Cascade 1697157632 2014-11-01 MCZ P. D. Perkins Amoron'i 1360 -20.166666 Antsirabe 1268 47.083332 ManiaUpstream Talaviana river 110 111 MAD14-01 1 Fianarantsoa 1697157596 MCZ Ranomafana NP & Perkins 890 -21.25 Namorona river 1264 47.416668 Vatovavy-Fitovinany 110 111 MAD14-09 1 Fianarantsoa 1697157640 1993-09-06 USNM W. E. Steiner & M. Stebbins 900 -21.266666 Ranomafana 1264 47.416668 110 111 1 Fianarantsoa 1697157603 1993-09-06 USNM W. E. Steiner & J. E. Cadle 900 -21.266666 Ranomafana 1264 47.416668 110 111 1 Fianarantsoa 1697157619 1990-03-01 1990-03-07 1990-03-01 USNM W. E. Steiner 900 -21.266666 Namorona River 1264 47.416668 Ranomafana 110 111 1 Fianarantsoa 1697157613 2001-08-11 NMW Ionilahy 200 -21.716667 Manakara 1262 47.633335 Riv. Ionilahy 110 111 1 Fianarantsoa 1697157597 2011-04-20 NMW R. Gerecke Ionilahy 284 -21.7 Exp. 1262 47.633335 Mahavavona 110 111 1 Fianarantsoa 1697157622 1988-11-01 1988-11-07 1988-11-01 USNM W. E. Steiner 1100 -21.25 Ranomafana 1264 47.416668 Namorona River 110 111 1 Fianarantsoa 1697157627 1988-10-08 1988-10-21 1988-10-08 USNM W. E. Steiner Namorona River 900 -21.266666 Namorona River 1264 47.416668 Ranomafana 110 111 1 Fianarantsoa 1697157617 2011-11-01 NHRS J. H. Randriamihaja Vatovavy Fitovinany 920 -21.25 Namorona river 1264 47.416668 Ranomafana 110 111 MAD11-09 1 Fianarantsoa 1697157616 2009-12-17 NHRS J. Bergsten & N. Jonsson & Ranarilalatiana & Randriamihaja 112 -18.766666 Tsingy de Bemaraha 1274 44.716667 Melaky 110 111 MAD09-68 1 Fianarantsoa 1697157660 2001-09-07 NMW Andhahela 250 -24.8 Isaka 1249 46.85 Tulear 110 111 1 Toliara 1697157643 1993-09-19 USNM W. E. Steiner & R. Andriamasimanana -23.15 Ifaty 1257 43.616665 110 111 1 Toliara 1697157615 2001-09-04 NMW Tsimelahy 300 -24.933332 Riv. Antarantsa 1249 46.633335 Tulear 110 111 1 Toliara