Fairmaire 1898b : 468
Hydraenidae of Madagascar (Insecta: Coleoptera)
Perkins, Philip D.
Zootaxa
2017
2017-11-03
4342
1
1
264
3N394
Fairmaire
Fairmaire
[151,514,1837,1863]
Insecta
Hydraenidae
Hydraena
GBIF
Animalia
Coleoptera
108
109
Arthropoda
species
dilutipes
Hydraena dilutipes Fairmaire 1898b: 468.
Type Material. Lectotype(male): Labels on lectotype: "Madag [illegible]; TYPE [red]; MUSEUM PARIS MADAGASCAR PERRIER DE LA BATHIE COLLECTION LEON FAIRMAIRE 1906; LECTOTYPE Hydraena dilutipesFairmaire 1898, design. P. D. Perkins 2015[red]". Two female paralectotypes: 1) " Hydraena dilutipes Farm Madag; MUSEUM PARIS MADAGASCAR PERRIER DE LA BATHIE COLLECTION LEON FAIRMAIRE 1906; PARALECTOTYPE Hydraena dilutipesFairmaire 1898, design. P. D. Perkins 2015[red]". 2) "Madag Perrier; MUSEUM PARIS MADAGASCAR PERRIER DE LA BATHIE COLLECTION LEON FAIRMAIRE 1906; PARALECTOTYPE Hydraena dilutipesFairmaire 1898, design. P. D. Perkins 2015[red]" (MNHN). FIGURE 98. Hydraena dilutipes, lectotype, habitus dorsal, lateral, and ventral. Specimensexamined (883): Antananarivo, 18.158S 47.2104E Analamanga, Manankazo riverby the bridge of RN4, medium size river over bedrock, elev. 1450 m, 18° 9' S, 47° 12' E, 21 xi 2014, J. Bergsten, R. Bukontaite, J.H. Randriamihaja, T. Ranarilalatiana& S. Holmgren( MAD14-75) (4 NHRS); Manakambahiny, in seepage over exposed granite cliff face, 18° 55' S, 47° 32' E, 12 i 1990, W. E. Steiner(28 USNM); Antsiranana, 14.4345S 49.7606E Sava Marojejy NP Camp II, wide river forming hygropetric rocks, wide river forming hygropetric rocks and rockpools, elev. 710 m, 14° 26' S, 49° 45' E, 9 xi 2014, J. Bergsten, R. Bukontaite, J.H. Randriamihaja, T. Ranarilalatiana& S. Holmgren( MAD14-54) (3 NHRS); 14.4369S 49.7749E Sava, Marojejy NP, 300mN from Camp I, upstream, waterhole with dead leaves next to the stream, elev. 490 m, 14° 44' S, 49° 46' E, 8 xi 2014, P. D. Perkins( MAD14-50) (10 MCZ); 14.4369S 49.7749E Sava MarojejyNP 300mN from CampI, upstream, waterhole with dead leaves next to the stream, elev. 490 m, 14° 26' S, 49° 46' E, 8 xi 2014, J. Bergsten, R. Bukontaite, J.H. Randriamihaja, T. Ranarilalatiana& S. Holmgren( MAD14-50) (22 NHRS); 14.5221S 49.7545E Sava, bridge 80kmfrom Sambava, where tributary river enters Lokoho river, rockpools and sandy waterpools at river margin, elev. 120 m, 14° 31' S, 49° 45' E, 17 xi 2014, J. Bergsten, R. Bukontaite, J.H. Randriamihaja, T. Ranarilalatiana& S. Holmgren( MAD14-73) (9 NHRS); Antsaba, Galoko Mountains, hygropetric cascade, elev. 303 m, 13° 39' S, 48° 45' E, 1 xi–29 xii 2004, M. Balke(MD 016) (124 ZSMC; one male " MNCN AI-547 DNAspecimen"); Diana: Antsaba: Ambolamena mountain, 4kmSE Antsaba village,“Antohatea be Cascade”, S 13.67593 E48.75589, GB nets, sieves, aspirator, forceps: waterfall with hygropetric rocks, elev. 381 m, 13° 41' S, 48° 45' E, 27 xi 2012, J. Bergsten, R. Bukontaite, J.H. Randriamihaja& T. Ranarilalatiana( MAD12-29) (1 NHRS); Diana: Antsaba: Galoko mountains, S 13.60974 E48.72175, sieves and aspirator: hygropetric rocks and water pools, elev. 263 m, 13° 37' S, 48° 43' E, 25 xi 2012, J. Bergsten, R. Bukontaite, J.H. Randriamihaja& T. Ranarilalatiana( MAD12-26) (1 NHRS); Forêtd' Antsahabe, 11.4 km275° W Daraina, malaise trap, tropical dry forest, elev. 550 m, 13° 12' S, 49° 33' E, 12 xii 2003, B.L.Fisher(BLF 10117) (1 CAS); Forêt d'Anabohazo, 21.6 km247° WSW Maromandia, malaise trap- in tropical dry forest, elev. 120 m, 14° 18' S, 47° 54' E, 11–16 iii 2001, Fisher, Griswold et al.(BLF 3336) (1 CAS); MarojejyNP, camp 2, hygropetric habitat, 14° 26' S, 49° 46' E, 30 x 2007, J. Stastny(1 NHRS); Montagne d'Ambre, Grande Cascade, elev. 740 m, 13° 34' S, 49° 10' E, 1 xi–30 xii 2004, M. Balke(MD 004) (20 ZSMC); Sambava: MarojejyNP, Camp2 long waterfall along smooth rock area (hygropetric), N:-14.43400, E:49.75633, elev. 717 m, 14° 26' S, 49° 45' E, 7 xii 2006, Isambert et al.(P57 Bi12) (8 NHRS); Sava, Marojejy NP, 800mN from Camp I, Humbertwaterfall; 14.4333S 49.773E, waterfall, elev. 550 m, 14° 26' S, 49° 46' E, 8–12 xi 2014, J. Bergsten, R. Bukontaite, J.H. Randriamihaja, T. Ranarilalatiana & S. Holmgren (MAD14-48) (12 NHRS); Marojejy NP, 800mN from Camp I, Humbert waterfall; 14.4333S 49.773E, ex. bedrock pools at side of waterfall, elev. 550 m, 14° 26' S, 49° 46' E, 8 xi 2014, P. D. Perkins (MAD14-48) (256 MCZ); Atsinanana, NP Midongy, 23° 30' S, 46° 55' E, 17 viii 2011, (collector not given on label) (4 NHRS); Fianarantsoa, 20.1711S 47.091E, Amoron'i, ManiaUpstream Talaviana riverfrom the crossing with RN7, 37km S of Antsirabe, rocky river with sandy bottom and some hygropetric rocks, elev. 1360 m, 20° 10' S, 47° 5' E, 1 xi 2014, P. D. Perkins( MAD14-01) (1 MCZ); 21.2581S 47.4213E, Vatovavy-Fitovinany, Ranomafana NP, Namorona riverby bridge at park entrance, rockpools and hygropetric sites by river, elev. 890 m, 21° 15' S, 47° 25' E, P. D. Perkins( MAD14-09) (37 MCZ); 7 kmW Ranomafana, from seepage over sunlit rock at river, montane rainforest, elev. 900 m, 21° 16' S, 47° 25' E, 6 ix 1993, W. E. Steiner& M. Stebbins(32 USNM); same locality, W. E. Steiner& J. E. Cadle(41 USNM); 7 kmW Ranomafana, on seepage over exposed rocks at bank of Namorona River, elev. 900 m, 21° 16' S, 47° 25' E, 1–7 iii 1990, W. E. Steiner(44 USNM); Ionilahy( Fianarantsoa), first left affl. of Riv. Ionilahyupstr. railroad bridge to Manakara, 200 m, 23.5°C, 0.088 mS/cm; hygropetric, elev. 200 m, 21° 43' S, 47° 38' E, 11 viii 2001, Gerecke & Goldschmidt (MD 20c) (4 NMW); Ionilahy( Fianarantsoa) Mahavavona, spring 3 Exp.NW, (hygropetric on rocks exposed to sun in rice field), elev. 284 m, 21° 42' S, 47° 38' E, 20 iv 2011, R. Gerecke(MD 218) (1 NMW); Namorona River, 7 kmW Ranomafana, from semi-shaded temporary pond in montane rain forest, elev. 1100 m, 21° 15' S, 47° 25' E, 1–7 xi 1988, W. E. Steiner(55 USNM); Namorona River, 7 kmW Ranomafana, on seepage over exposed rock at bank of Namorona River, elev. 900 m, 21° 16' S, 47° 25' E, 8– 21 x 1988, W. E. Steiner(38 USNM); Ranomafana, Namorona main river, rockpools, elev. 927 m, 21° 15' S, 47° 25' E, 1 xi–30 xii 2004, M. Balke (MD 029) (93 ZSMC); Vatovavy Fitovinany: RanomafanaNP: Namorona riverby the bridge below park entrance: S 21.25809 E0 47.42165, GB Nets& sieves, rockpools, elev. 920 m, 21° 15' S, 47° 25' E, 1 xi 2011, J. Bergsten, R. Bukontaite, T. Ranarilalatiana, & J.H. Randriamihaja( MAD11-09) (9 NHRS); Melaky: Tsingy de BemarahaNP. S 18.75875 E0 44.71648, elev. 112 m, 18° 46' S, 44° 43' E, 17 xii 2009, J. Bergsten, N. Jönsson, T. Ranarilalatiana, HJ. Randriamihaja( MAD09-68) (1 NHRS); Toliara, Andhahela( Tulear), Isaka, stream exp. W 1 kmN from the vill., elev. 250 m, 24° 48' S, 46° 51' E, 7 ix 2001, Gerecke & Goldschmidt (MD 63) (1 NMW); Ifaty, under seaweed drift on sand beach, 23° 9' S, 43° 37' E, 19 ix 1993, W. E. Steiner& R. Andriamasimanana(1 USNM); Tsimelahy( Tulear), Riv. Antarantsa, ca. 1 kmupstream from village, elev. 300 m, 24° 56' S, 46° 38' E, 4 ix 2001, Gerecke & Goldschmidt (MD 58) (20 NMW).
Differential Diagnosis.Closely related to H. dilutipoides, differing therefrom by the larger size (ca. 1.81 mmvs. 1.73), the more coarsely densely punctate pronotum, and having the elytral intervals strongly raised and forming unbroken lines. The plaques of H. dilutipoidesare more widely separated, especially posteriorly, and more sharply rounded at the posterior edge. Males of both species have large metatibial brushes. The aedeagi also show a relationship, having a large, flat distal extension of the main piece; the complex gonopore bearing distal piece differs significantly in the two species ( Figs. 100–102).
Description. Size: Lectotype(length/width, mm): body (length to elytral apices) 1.81/0.76; head width 0.43; pronotum 0.42/0.60, PA 0.44, PB 0.53; elytra 1.11/0.76. Dorsum dark brown, head darkest; legs brown; maxillary palpi brown to light brown, distal ½ of last palpomere darker. Mentum sparsely finely punctulate, shining; postmentum rugulose, dull. Genae raised, weakly rugulose, dull, without distinct posterior ridge. Head and pronotum coarsely densely punctate, punctures of pronotum slightly larger than those of head; interstices narrow ridges, shining. Pronotum slightly sinuate laterally; anterior margin straight behind eyes, emarginate behind frons; PF1 very shallow; PF2 shallow, very shallowly confluent medially forming weak U-shaped impression; PF3 moderately deep; PF4 absent. Elytra weakly arcuate laterally; summit of posterior declivity near distal 2/3; lateral explanate margins moderately narrow; serial punctures deeply impressed, round, and closely spaced, slightly larger than pronotal punctures. Intervals raised, narrow, shining, ca. 0.5xpd. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming very shallow angle with one another. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/2/1.3. P1 laminate; median carina sinuate in profile. P2 l/w ca. 2/1, sides parallel, apex blunt, apex raised slightly above mesoventral intercoxal process. Plaques large, anteriorly and posteriorly rounded, parallel, anterior at level of surrounding cuticle, gradually elevating posteriorly such that posterior border is slightly raised above surrounding cuticle, located at sides of median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at slightly arcuate posterior margin ca. 1.5x P2. Protibia moderately wide, slightly arcuate on medial margin, slightly more so on lateral margin. Mesotibia straight; medial margin with 7–8 very short spines (microslide mount), margin not notched between spines. Metatibia slightly arcuate, distal ½ of medial margin slightly emarginate and with large brush of setae. Abdominal apex slightly asymmetrical; last tergite with apicomedian notch. Females average slightly smaller than males; male metafemur longer and more sinuately shaped than in female; female metatibia simple and slightly shorter than in male.
Remarks.Fairmaire (1898) in the original description notes the differences in the shape of the abdomen of the male and female. It is therefore unquestionable that he had both sexes; however, he does not indicate how many specimens total he had. The locality given is "Dans l'Ikopa"; Ikopa is the river that flows through Antananarivo. Although the river is quite long, ca. 300 km, the typespecimens were most likely collected in or very near the capital. Specimens were collected in the capital as late as 1990, by W. E. Steiner. However, these were not collected in the river, which is now quite polluted. Trizzino et al.(2012)presented the DNA sequence information of one male specimen ("MNCN AI-547 DNA specimen") with this locality data: Antsaba, Galoko Mountains, hygropetric cascade, elev. 303 m, 13° 39' S, 48° 45' E, 1 xi–29 xii 2004, M. Balke (MD 016).
1697157591
1990-01-12
2014-11-21
1990-01-12
NHRS, USNM
J. Bergsten & R. Bukontaite & J. H. Randriamihaja & T. Ranarilalatiana & S. Holmgren & W. E. Steiner
Specimens
1450
-18.916666
Manankazo river
1273
47.533333
Analamanga
109
110
MAD14-75
1
Antananarivo
1697157633
2014-11-09
NHRS
NP Camp II & J. Bergsten & R. Bukontaite & J. H. Randriamihaja & T. Ranarilalatiana & S. Holmgren
710
-14.433333
Sava Marojejy
1288
49.75
109
110
MAD14-54
1
Antsiranana
1697157607
2014-11-08
Marojejy NP & Camp I & Philip D. Perkins
300
-14.733334
Sava
1287
49.766666
109
110
1
Antsiranana
1697157589
[151,537,151,176]
MCZ
Perkins
110
111
MAD14-50
1
1697157604
2014-11-08
NHRS
J. H. Randriamihaja & Ranarilalatiana & Holmgren
300
-14.433333
Camp
1288
49.766666
Sava Marojejy
110
111
MAD14-50
1
1697157580
2014-11-17
NHRS
T. Ranarilalatiana & Holmgren
Sava
120
-14.516666
Lokoho river
1287
49.75
Sambava
110
111
MAD14-73
1
1697157583
2004-11-01
2012-11-27
2004-11-01
ZSMC, MNCN, DNA, NHRS
M. Balke & J. Bergsten & R. Bukontaite & J. H. Randriamihaja & T. Ranarilalatiana
Antsaba
303
-13.683333
Ambolamena mountain
1290
48.75
Galoko Mountains
110
111
AI-547, MAD12-29
1
Cascade
1697157605
2012-11-25
NHRS
J. H. Randriamihaja & Ranarilalatiana
Diana
263
-13.616667
Galoko mountains
1290
48.716667
Antsaba
110
111
MAD12-26
1
Cascade
1697157610
2003-12-12
CAS
B. L. Fisher
Foret
550
-13.2
Daraina
1291
49.55
Antsahabe
110
111
1
Cascade
1697157599
2001-03-11
2001-03-16
2001-03-11
CAS
120
-14.3
Maromandia
1288
47.9
Foret d'Anabohazo
110
111
1
Cascade
1697157612
2007-10-30
NHRS
J. Stastny
-14.433333
Marojejy
1288
49.766666
110
111
1
Cascade
1697157624
2004-11-01
2004-12-30
2004-11-01
ZSMC
M. Balke
740
-13.566667
Montagne d'Ambre
1290
49.166668
110
111
1
Cascade
1697157645
2006-12-07
NHRS
Isambert & Bi
Sambava
717
-14.433333
Camp
1288
49.75
Marojejy
110
111
1
Cascade
1697157623
[250,948,836,861]
Marojejy NP & Camp I & Humbert
800
Sava
110
111
1
Cascade
1697157630
[238,1432,980,1005]
2011-08-17
NHRS
NP Midongy
-23.5
Atsinanana
1255
46.916668
110
111
1
Cascade
1697157632
2014-11-01
MCZ
P. D. Perkins
Amoron'i
1360
-20.166666
Antsirabe
1268
47.083332
ManiaUpstream Talaviana river
110
111
MAD14-01
1
Fianarantsoa
1697157596
MCZ
Ranomafana NP & Perkins
890
-21.25
Namorona river
1264
47.416668
Vatovavy-Fitovinany
110
111
MAD14-09
1
Fianarantsoa
1697157640
1993-09-06
USNM
W. E. Steiner & M. Stebbins
900
-21.266666
Ranomafana
1264
47.416668
110
111
1
Fianarantsoa
1697157603
1993-09-06
USNM
W. E. Steiner & J. E. Cadle
900
-21.266666
Ranomafana
1264
47.416668
110
111
1
Fianarantsoa
1697157619
1990-03-01
1990-03-07
1990-03-01
USNM
W. E. Steiner
900
-21.266666
Namorona River
1264
47.416668
Ranomafana
110
111
1
Fianarantsoa
1697157613
2001-08-11
NMW
Ionilahy
200
-21.716667
Manakara
1262
47.633335
Riv. Ionilahy
110
111
1
Fianarantsoa
1697157597
2011-04-20
NMW
R. Gerecke
Ionilahy
284
-21.7
Exp.
1262
47.633335
Mahavavona
110
111
1
Fianarantsoa
1697157622
1988-11-01
1988-11-07
1988-11-01
USNM
W. E. Steiner
1100
-21.25
Ranomafana
1264
47.416668
Namorona River
110
111
1
Fianarantsoa
1697157627
1988-10-08
1988-10-21
1988-10-08
USNM
W. E. Steiner
Namorona River
900
-21.266666
Namorona River
1264
47.416668
Ranomafana
110
111
1
Fianarantsoa
1697157617
2011-11-01
NHRS
J. H. Randriamihaja
Vatovavy Fitovinany
920
-21.25
Namorona river
1264
47.416668
Ranomafana
110
111
MAD11-09
1
Fianarantsoa
1697157616
2009-12-17
NHRS
J. Bergsten & N. Jonsson & Ranarilalatiana & Randriamihaja
112
-18.766666
Tsingy de Bemaraha
1274
44.716667
Melaky
110
111
MAD09-68
1
Fianarantsoa
1697157660
2001-09-07
NMW
Andhahela
250
-24.8
Isaka
1249
46.85
Tulear
110
111
1
Toliara
1697157643
1993-09-19
USNM
W. E. Steiner & R. Andriamasimanana
-23.15
Ifaty
1257
43.616665
110
111
1
Toliara
1697157615
2001-09-04
NMW
Tsimelahy
300
-24.933332
Riv. Antarantsa
1249
46.633335
Tulear
110
111
1
Toliara