Pinotus triquetrus Luederwaldt, 1923:11 (original description). Pintous triquetrus Blackwelder 1944: 208 Dichotomius triquetrus Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae) Montoya-Molina, Santiago Vaz-De, Fernando Z. European Journal of Taxonomy 2021 2021-02-11 734 1 64 35ND7 (Luederwaldt, 1923) Luederwaldt 1923 [416,1171,1046,1073] Insecta Scarabaeidae Dichotomius Animalia Coleoptera 46 47 Arthropoda species triquetrus Luederwaldtinia      Pinotus triquetrus Luederwaldt, 1923:11(original description).    Pintous triquetrus–  Blackwelder 1944: 208.   Dichotomius triquetrus– Schoolmeesters 2018( Catalogue of Life).    Diagnosis   Dichotomius triquetrusis separated from other species in the group by the following combination of characters: dorsal and ventral surface reddish to dark brown. Anterior clypeal margin with deep emargination separating two obtuse teeth. Head interocular space displaying flat punctures, resembling rough; distinct ocellate punctuation near eyes. Pronotum medial disc with fine punctures, anterolateral edges with well-impressed punctures, ocellate punctuation along margins. Elytra striae strongly marked, interstriae microsculpture shagreened    Material examined   Lectotype(here designated)   TRINIDAD AND TOBAGO• ♀; “Caparo Trinidad Heyne, Berlin-Wilm.” [white, p]/ “  Pinotus triquetrus n. sp.L̹d.det.23” [white, hw]/ “18188” [white, p] /[blue rounded label]; MZSP.  Additional material   TRINIDAD AND TOBAGO• 1 ♂, 1 ♀; St. George Co., Range, Simla Research Station, Arima Valley, N. Range;  800 fta.s.l.; 10º41′34″ N, 61º17′22″ W; FIT [ flight interception trap]; premontane tropical rainforest;  3 Jun. 2000; S. Alistairand Ramsdaleleg.; CEMT•  1 ♀; Trinidad; 1904; F. Birchleg.; BMNH•  1 ♂; Antilles Trinidad; Fryleg.; BMNH 1905-100•  1 ♂; Porto Spain,  Dec 1912–  May 1913; R. Thaiterleg.; MZSP.   VENEZUELA• 1 ♂; BMNH. COUNTRY UNKNOWN •  1 ♀; Nevinsonleg.; BMNH 1918-14.    Redescription  Female MEASUREMENTS. Length: 17.89 mm. Dorsal and ventral surfaces reddish to dark brown. HEAD. Anterior edge surface horizontally rugose, interocular space displaying flat punctures, resembling rough; distinct ocellate punctuation near eyes. Clypeus with central deep emargination separating two acute clypeal teeth ( Fig. 16B–C). Cephalic carina bituberculate, wider than high, tubercles obtuse apically ( Fig. 16C). Ventral clypeal process truncate (see Fig. 2B). PRONOTUM. Gibbous, wider than long. Pronotal anterior disc shiny and smooth with fine, dense and wellimpressed punctures near antero-lateral sides. Uninterrupted ocellate punctures along pronotal margins ( Fig. 16B, D). Hypomeral disc punctate; external margin bearing single row of organized long setae, approximately of same length (see Fig. 2F). ABDOMEN. Prosternum surface with fine microsculptured, glabrous. Mesosternum and metepisternum surface microsculpture shagreened with ocellate setigerous punctures. Metasternum setigerous punctures restricted mainly to anterior and lateral edges of mestasternal lobe, puncture diameter variable. Punctuation of antero-median edge connecting with lateral ones through margin of mesocoxa (see Fig. 2G). Lateral edges of metasternum with ocellate, setigerous punctures separated by width of a puncture, pilosity sparse, setae short, not reaching metepisternum suture. Longitudinal sulcus clearly visible. Ocellate punctures along anterior margins of sternites. Pygidium ocellate punctation on basal edge and through lateral margins. ELYTRA. Interstriae microsculpture shagreened (see Fig. 2J), fine punctures randomly distributed. Striae strongly marked, ocellate punctures separated by diameter of a puncture on elytral disc ( Fig. 16D). LEGS. Metacoxa posterior edge bearing single row of organized ocellate punctures, separated by diameter of a puncture. Posterior margin of meta-trochanter with long setae.   Morphological variation  Males( Fig. 16A) are distinguished from females as follows: HEAD. Cephalic carina higher than wide, conical in shape. Median tubercle twice as high as lateral tubercles, rounded, weakly bifurcate apically and curved posteriorly. Apex of lateral tubercles acute. PRONOTUM. Surface covered by fine punctures, denser at antero-lateral areas; anterior disc with weak declivity. PARAMERES AND LAMELLA COPULATRIX. As in Fig. 16E–G.    Remarks This is an endemic species from Trinidad.The redescription was based on the well-developed female of the MZSP and the underdeveloped female at the CEMT. The male redescription was based on the specimen in the MZSP wrongly determined by Pereira as “  Pinotus belus” and an underdeveloped male from the CEMT. Male genitalia were extracted from the underdeveloped male at CEMT. Luederwaldt (1923)also mentioned a male specimen in the typeseries that could not be traced. 3037418471 MZSP Trinidad and Tobago TRINIDAD AND TOBAGO 46 47 1 1 lectotype 3037418559 2000-06-03 flight interception trap CEMT S. Alistair & Ramsdale Trinidad and Tobago St. George Co. 244 10.692778 Simla Research Station 21 -61.289444 Range 46 47 2 1 Arima 3037418340 BMNH F. Birch Trinidad and Tobago Trinidad 46 47 1 1 3037418593 [304,898,1959,1988] BMNH Fry Trinidad and Tobago Antilles Trinidad 46 47 BMNH 1905-100 1 1 3037418338 1912-12 1913-05 1912-12 MZSP R. Thaiter Trinidad and Tobago Porto Spain 46 47 1 1 3037418510 [189,538,263,291] BMNH Venezuela VENEZUELA 47 48 1 1 3037418564 [515,950,332,359] BMNH Nevinson 47 48 BMNH 1918-14 1 1