Healyella boluana Omodeo & Rota, 1989

Szederjesi, Tímea, Dányi, László, Kaydan, Mehmet Bora & Csuzdi, Csaba, 2018, Contribution to the knowledge of the earthworm fauna of Turkey with description of three new species (Clitellata: Megadrili), Zootaxa 4496 (1), pp. 160-172 : 167-168

Healyella boluana Omodeo & Rota, 1989

Healyella boluana Omodeo & Rota, 1989: 176 . Csuzdi et al. 2006: 20.

Material examined. HNHM /17095 3 (2 truncated and 1 intact) ex., No. 15, Akçaören , Düzce, above the lake S of the village, open forest with deciduous oak and Platanus , N40°55.721' E31°17.392', 296 m, leg. L. Dányi, M.B. Kaydan, 21.0 4.2015 . HNHM /17097 1 truncated ex., No. 19, Kazpınarı , Bartın, N of the village, rocky slope with bushes, N41°42.634' E32°21.811', 319 m, leg. L. Dányi, M.B. Kaydan, 21.0 4.2015.

External characters. 62 mm long and 4 mm wide. Number of segments 116. Slight purplish pigmentation dorsally. Prostomium tanylobous. First dorsal pore at intersegmental furrow 6/7. Setae distant. Setal arrangement behind clitellum: aa: ab: bc: cd: dd = 1.27:1.8:1:1.13:1.8. Male pores on 15 between a and b, spherical, slightly invaginated, with large circular, glandular tumescences. Female pore not visible, covered by flat spermatophores. Nephridial pores irregularly alternate between setal line b and above d. Clitellum annular on 26–30. Tubercula pubertatis lacking.

Internal characters (based on one specimen). Septa 6/7–14/15 slightly thickened. Testes and funnels paired in 10–11, covered by perioesophageal testic sacs. Two pairs of seminal vesicles in 11 and 12. Spermathecae lacking. Calciferous glands in 10–12 with small diverticula in the posterior part of 10. Last heart in 11, with a pair of extraoesophageal vessels in 12. Nephridial bladders simple, sausage-shaped. Crop in 15–16, gizzard in 17–18, occupying the cavity of 19 as well. Typhlosolis large, anchor-shaped. Longitudinal musculature of pinnate type.

Remarks. He. boluana was described on a single specimen from the surroundings of Bolu city (Boludagi Geçidi). Our specimens are from some 60 (No. 15) and 120 (No. 19) km North-East of the type locality.

Our specimens completely agree with the original description, except the position of the nephridial pores: Omodeo & Rota (1989) state that they are aligned just above setal line b, in our specimens they alternate irregularly between b and above d.