Enicospilus yonezawanus (Uchida, 1928)

Shimizu, So, Broad, Gavin R. & Maeto, Kaoru, 2020, Integrative taxonomy and analysis of species richness patterns of nocturnal Darwin wasps of the genus Enicospilus Stephens (Hymenoptera, Ichneumonidae, Ophioninae) in Japan, ZooKeys 990, pp. 1-144 : 1

Enicospilus yonezawanus (Uchida, 1928) Figure 54

Henicospilus yonezawanus Uchida, 1928: 218; LCT ♀ from Japan, designated by Townes et al. (1965: 337), SEHU, examined.

Enicospilus microstriatellus Uchida, 1956: 95; HT ♂ from Ryûkyûs, SEHU, examined; synonymised by Gauld and Mitchell (1981: 337).

Specimens examined.

Total of 303 specimens (196♀♀103♂♂ and 4 unsexed): India (12♀♀ and 2 unsexed), Indonesia (1♀), Japan (166♀♀101♂♂), Laos (11♀♀1♂), Malaysia (5♀♀ and 2 unsexed), Papua New Guinea (1♀), Taiwan (1♂).

Type series: LCT ♀ of Henicospilus yonezawanus Uchida, 1928, Yonezawa, Yamagata Pref., Tôhoku, JAPAN, 23.VII.1919, S. Matsumura leg. (SEHU); HT ♂ of Enicospilus microstriatellus Uchida, 1956, Sinmura, Amami-ôshima, Kagoshima Pref., Ryûkyûs, JAPAN, 7.IV.1954, T. Kumata leg. (SEHU).

Distribution.

Australasian, Eastern Palaearctic, and Oriental regions ( Yu et al. 2016).

JAPAN: [ Tôhoku] Akita*, Yamagata ( Uchida 1928; Shimizu 2020; present study), and Fukushima*; [Hokuriku] Niigata*; [ Kantô-Kôshin] Tochigi ( Uchida 1928) and Tôkyô ( Konishi and Maeto 2000; present study); [ Tôkai] Shizuoka*; [Kinki] Shiga*, Kyôto *, Hyôgo *, and Wakayama*; [ Chûgoku] Shimane* and Hiroshima ( Konishi and Nakamura 2002, 2005, 2010; Maeto and Shimizu 2019; present study); [ Kyûshû] Fukuoka*, Nagasaki*, and Kagoshima ( Shimizu 2020); [ Ryûkyûs] Kagoshima ( Uchida 1956; Momoi 1970; Shimizu 2020; present study) and Okinawa ( Momoi 1970; Shimizu 2020; present study). *New records.

Bionomics.

Unknown.

Differential diagnosis.

Enicospilus yonezawanus is one of the most common Enicospilus species in Japan and easily distinguished from all other Enicospilus species by the following combination of character states: ventral margin of clypeus impressed (Fig. 54B, D ); fore wing fenestra with triangular proximal sclerite and without central sclerite (Fig. 54F ); and meso- and metapleuron closely punctostriate (Fig. 54E ). Enicospilus yonezawanus is also sometimes confused with E. jilinensis but can easily be separated (cf. Differential diagnosis of E. jilinensis ).

Remarks.

There is some variation in the shape of the proximal sclerite, but in Japanese specimens it is usually very stable.

Figures

Figure 54. Enicospilus yonezawanus (Uchida, 1928) ♀ from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing.