Causeyella causeyae, Shear, William A., 2003

Shear, William A., 2003, The milliped family Trichopetalidae, Part 1: Introduction and Genera Trigenotyla Causey, Nannopetalum n. gen., and Causeyella n. gen. (Diplopoda: Chordeumatida, Cleidogonoidea), Zootaxa 321, pp. 1-36 : 30-31

Causeyella causeyae , n. sp.

Figs. 23, 24

Types: Male holotype, male paratype and 2 female paratypes from ARKANSAS, Independence Co., Foushee Cave, 6 miles west of Locust Grove, collected by unknown person or persons, May, 1978 (FSCA). The following are paratypes: ARKANSAS: Independence Co., Cushman Cave, 0.5 miles from Cushman, 8 July 1974, G. Harp, four males, 3 females; Dodd Cave, 14 December 2001, G. Greening, D. Fenolio, male (SBP). Izard Co., Clay Cave, 7 mi SW Melbourne, 11 October 1974, G. Harp, numerous males and females; Donovan Cave, 2 mi SW Calico Rock, 24 January 1964, D. Martin, male; Goat (Prison) Cave, 22 November 2002, B. Wagner, D. Kempwerth, male (SBP). Stone Co., Hell’s Creek Cave, 2.5 mi N Mountain View, 26 January 1974, V. R. McDaniel, 2 males; Blanchard Springs Cave, 27 September 1967, T. C. Barr, juveniles probably this species; Rowland Cave, 12 June 2001, G. Greening et al., male (SBP); Alexander Cave, 24 March 2001, M. Slay, L. Moritz; Gustafson Cave, 6 October 2002, G. Graening et al., male (SBP).

Diagnosis: A large (13.5 mm long) species of Causeyella with lamellate branches of the gonopod colpocoxites; frontal coxal setae somewhat thickened and blunt.

Etymology: For Nell Bevel Causey.

Male holotype: Length, 13.5 mm, width 0.9 mm. Unpigmented, eyeless. Segmental shoulders well developed, paranota­like; segmental setae 65–70% of body width. Legpairs 3–7 noticeably enlarged compared to legpairs 12 and posterior; no podomeres modified. Gonopods ( Fig. 23 ) with well developed sternal lobes. Anterior surfaces of coxae entirely covered with prominent scaly trichomes becoming larger distally; about 15 setae on each coxa, not separated into groups; mesal coxites in anterior view slightly spatulate, ectal coxites only about half their length, broad in lateral view. Lamellate branch of colpocoxite very large, curled at tip, roughened with rows of blunt trichomes, extending anterior of mesal coxites. Ninth legpair ( Fig. 24 ) with coxae laterally flattened, mesally excavated, bearing vestigial glands, articulation of prefemora offset laterally; prefemora longer than coxae, two or three vaguely indicated articles distally, claw short, twisted. Legpairs 10, 11 with glands, unmodified.

Female paratype: Length, 13.5 mm, width 1.05 mm. Nonsexual characters as in male.

Distribution: Independence, Stone, and Izard counties, ARKANSAS, as listed under “ Types ” above. Literature records: ARKANSAS: Izard Co., Bergren Cave, Needles Cave; Stone Co., Hell Creek Cave (as “ Scoterpes n. sp. ” in McDaniel and Smith, 1976). These records were not verified; the wherabouts of the specimens are unknown. It appears that this species occurs on both sides of the White River. Records of “ Scoterpes ” from Oregon Co., Missouri (see under C. dendropus above) may be C. causeyae , but are probably in error.

Causey labelled examples of this species as “ Scoterpes martini ” or “ Scoterpes dendropus ssp. martini . ”

Figures

FIGURES 22 – 26. Causeyella dendropus Loomis, 22, gonopods, anterior view. C. causeyae, n. sp., 23, 24. 23, gonopods, anterior view. 24, male ninth legs, anterior view. C. youngsteadtorum, n. sp., 25, 26. 25, gonopods, anterior view. 26, male ninth legs, anterior view. Scale lines = 0.1 mm (100 µm) for Figs. 22, 23, 25, 0.5 mm (500 µm) for Figs. 24, 26.