Afroscatopse haennii, Szadziewski & Bojarski & Sontag & Szwedo, 2022

Szadziewski, Ryszard, Bojarski, Błażej, Sontag, Elżbieta & Szwedo, Jacek, 2022, A new giant dung midge from Miocene Ethiopian amber (Diptera: Scatopsidae), Zootaxa 5099 (5), pp. 549-562 : 552-554

Afroscatopse haennii sp. nov.

( Figs 1–3 )

urn:lsid:zoobank.org:act:EF05224A-51D4-408B-B4B7-FE389A786D12

Diagnosis. As for genus, as it is only known species so far.

Description. Male ( Fig. 1 ) large, with total length about 7.0 mm, whole body including wings dull, not shining, mostly black; pleurae more pale, scutum at scutellum with 3 yellowish elliptic spots covering also scutellum (2 lateral and 1 median), anterior shoulders also yellowish. Head rounded, as wide as high. Eyes emarginated at antennal bases, separated above by distance of an ocellus; hairs between ommatidia not visible ( Figs 2A, B ). Three well developed ocelli present ( Fig. 2B ). Antennae much longer than head, a little bit longer then scutum ( Fig. 2A ). Scape and pedicel of antenna short, each with irregularly distributed setulae ( Fig. 2C ). Flagellum 1.50 mm, much longer than head and a little bit longer than scutum, composed of 9 uncompressed flagellomeres; flagellomere 1 longest, cylindrical, almost 2 times longer than next flagellomere; flagellomeres 2-5 cylindrical as long as broad, flagellomeres 6-8 more or less spherical, flagellomere 9 cylindrical 1.5 times longer than broad; each flagellomere with more or less distinct rings of short spine-like setae, flagellomere 1 with 4-5 irregular rings, flagellomeres 2-7 and 9 with 2 distinct rings, and flagellomere 8 with 1 ring ( Fig. 2C ). Frons bare, separated from clypeus bearing setae ( Fig. 2B ). Maxillary palpus one-segmented, very short.

Thorax of holotype with proportions of length/width/height: 26/22/27. Scutum almost as long (26) as wide (22); widest at wing bases, narrower at head ( Fig. 1 ). Scutum bare except for lateral margins bearing irregular rows of short setulae; scutellum bare, without marginal setae. Spiracular sclerite distinct and setose, almost rectangular, with central opening; postpronotum bearing short setae, katepisternum bare.

Wing very broad, almost 2 times as long as broad, length 4.68-5.14 mm. Subcosta very short, not reaching costa at wing margin ( Fig. 2D ). Costal vein swollen on proximal 1/3, R 2+3 lost. R 4+5 long, reaching margin near wing apex. Medial vein M 1 angled, reduced on proximal part; a short spur at angle present; M 2 well developed from transverse vein r-m to wing apex; M 3+4 or M 4 and CuA well visible, both gradually C-curved; all veins reaching wing margin. Wing membrane with distinct microtrichia; macrotrichia present only on costal vein and on dorsal surface of R 1, absent on wing membrane and other veins. Stem of haltere with a row of spine-like setae.

Legs relatively stout, without modifications. Tibial spurs absent. Tarsi short, 5-segmented, tarsomere 1 as long as 2-4 together, claws equal, simple, empodium well developed; tarsus of hind leg about 0.9 mm long, proportions of tarsomeres as follows: 1—37, 2-4—35, 5—27.

Genitalia barely visible, not rotated, symmetrical. Sternite 7 with 4 caudal projections ( Fig. 3B ), tergite 7 divided into 2 long and blunt, bare, divergent lobes ( Figs 3A–D ); gonoxite broad with pointed apical projection, covered with long setae, gonostyli reduced ( Fig. 3D ).

Female unknown.

Material examined. Holotype male (complete), paratype male (incomplete) , and head of third specimen not included to the type series , MAIG 6013. Syninclusions: Coleoptera : Staphylinidae 1, Hemiptera : Miridae 1, Hymenoptera: Ceraphronidae 1, Isoptera 1 winged, Acari at least 2; Scatopsidae 1; Lepidoptera 1. Abdomen of the holotype is covered by milky veil. Till now such a veil was observed only in not heated Baltic amber inclusions ( Szadziewski 2006).

Etymology. The new species is dedicated to Jean-Paul Haenni of the Muséum d’histoire naturelle, Neuchâtel, Switzerland, in recognition of his important contributions to the taxonomy of Scatopsidae .

Figures

FIGURE 1. Afroscatopse haennii gen. et sp. nov., holotype male, dorsal view.

FIGURE 2. Afroscatopse haennii gen. et sp. nov., holotype male, head and wing.A—head with antennae, B—vertex and frons, C—antenna, D—wing.

FIGURE 3. Afroscatopse haennii gen. et sp. nov., male genitalia. A—ventral view, phot., B—ventral aspect, drawing, C—lat- erodorsal view, phot., D—laterodorsal aspect of tergite 7, and gonocoxite. Abbreviations: st7—abdominal sternite 7, tg7—ab- dominal tergite 7, gx—gonocoxite.