Salcedia procera, Balkenohl, 2020

Balkenohl, Michael, 2020, A genus in disguise. Revision of the genus Salcedia Fairmaire, 1899 with descriptions of nine new species (Coleoptera, Carabidae, Scaritinae, Salcediini), ZooKeys 901, pp. 1-81 : 1

publication ID	https://dx.doi.org/10.3897/zookeys.901.39432
publication LSID	lsid:zoobank.org:pub:843F6065-D00E-48E6-9A26-9CEB044BF102
persistent identifier	https://treatment.plazi.org/id/AD23EE1E-DDCE-44BB-B08F-65A80A794867
taxon LSID	lsid:zoobank.org:act:AD23EE1E-DDCE-44BB-B08F-65A80A794867
treatment provided by	ZooKeys by Pensoft
scientific name	Salcedia procera
status	sp. nov.

Treatment

Salcedia procera View in CoL sp. nov. Figs 18 View Figures 14–19 , 36 View Figures 36–43 , 62 View Figures 58–63 , 77 View Figures 77–83 , 84 View Figure 84

Type material.

Holotype: ♂, with labels and data: white, printed and handwritten in black ink "Angola: Lac Calundo 105 km E. Vila Luso 4492.8 XII-1954 E. Luna de Carvalho" / green, black framed “Récolté sous des pierres" / white, handwritten in pencil “ANG:4592-8” (MRACT). Paratypes: 1 spec., same data as holotype; 1 ♂, 1 ♀, same data as holotype but “4724-2”; 1 ♀, 3 specs., same data as holotype but with numbers 4510-18, 4538-11, 4643-5, 4623-3 and additional label " A la lumière” (MRACT, CBB).

Remark.

In some of the paratypes, parts of the tarsomeres and antennae are missing.

Diagnosis.

A medium sized species, with super-elongate outline of the elytra with maximum width anterior and posterior middle and the pronotum with three additional complete carinae. The pseudohumerus is obtuse angular and dentate. The antennomeres are sub-moniliform. Distinguished most clearly from the similar species S. africana by the smaller size, the sub-moniliform antennomeres and the front tibia with four lateral teeth. The second similar species, S. elongata Alluaud is much larger, its antennomeres are moniliform, the elytra are oblong-elongate and the anterior angles of the pronotum are acutely produced anteriorly. Moreover, it is the only species of the genus with slightly concave outline of the elytra, so that the maximum width is anterior and posterior to the middle.

Description.

Measurements in Table 1 View Table 1 .

Colour and surface: Piceous, areas between carinae and pits shiny; legs, and antennae fuscous, palpi leoninous.

Head: Three-quarters of pronotum width. Outline pentagonal shaped. Clypeus wide, straight anteriorly, fused with clypeal wings, separated from supra-antennal plates by distinct notches, with convex transverse field at middle, separated from frons by deep transverse furrow. Frons with two raised paramedian carinae, joining anteriorly into a central erected tubercle, with two minute acute glossy teeth bilaterally anterior to the central tubercle, with two parallel carinae paralaterally near base; frons and clypeus separated from supra-antennal and supraorbital plates by deep broad furrows, each furrow with deep slightly longitudinal pit between supra-antennal plate and clypeus; supra-antennal and supraorbital plates distinctly margined, margin raised, carina-like, nearly smooth, supra-antennal plates slightly vaulted. Base emarginated at middle, nearly right-angled laterally (angle 105°-107°). Eyes large, convex, with transverse shape in lateral view, with small part visible in dorsal view; genae slightly convex. Antenna with segments five to ten sub-moniliform (L/W 0.95-1.02), densely pubescent, segments two to four scarcely pubescent, scapus and pedicellus with longitudinal reticulation. Labrum convex anteriorly. Mandible moderately short, wide, slightly arcuate at apex. Apical segment of maxillary palpomere moderately long. Mentum small, elongated, epilobes elongated, projecting and distinctly angled anteriorly, margined medially, surface covered with flattened irregular pits.

Pronotum (Fig. 36 View Figures 36–43 ): Outline rectangular, transverse, a fifth wider than long. Lateral margin nearly straight at middle, slightly converging anteriorly, maximum width behind middle, slightly converging at anterior and posterior angles. Lateral margin indistinctly crenulated, with nine to ten small tubercles, tubercles sometimes doubled (100 × magnification); with anterior angles right-angled, with distinct emargination at posterior angles. Base straight laterally, with distinct keel at middle. Disc with two raised paramedian carinae parallel to median line and diverging posteriorly, with two slight notches, with median line narrow and moderately short, ending in longitudinal pits anteriorly and posteriorly, with four additional shorter carinae bilaterally, the paralateral ones joining with the paramedian carinae at base and forming tooth-like tubercle pointing posteriorly, extended anteriorly up to anterior margin as less raised paralateral carina. With inner and outer lateral carinae well developed, the inner one connected with the extension of the paralateral carina, not connected with the outer lateral carina. All carinae sub-crenulate. Lateral margin and space between lateral margin and paralateral carina wing-like bent up, with six large transverse pits.

Elytron: Flattened to slightly depressed in anterior half (lateral view), moderately convex in frontal view. Super-elongate, margin nearly parallel but slightly concave at middle, maximum width anterior and posterior middle. Pseudohumerus obtuse angular (angle 111°), dentate. Apex rounded, indistinctly denticulate at suture. Disc with interneur six sub-crenulate, interneur three slightly carinate in anterior third, interneur two running up to apex as parallel line, distinctly raised; interneur four running in parallel at middle, convex towards apex, almost reaching apex, nearly reaching base. Interval five and six with two rows of serial pits. Interneur five indistinctly visible due to transversally connected pits.

Hind wings: Fully developed.

Lower surface: Antennal channel of pronotum with isodiametric reticulation. Pseudoepipleura with one row of pits, interrupted at middle, lateral margin of elytron smooth. Metepisternum distinctly elongated, with broad longitudinal groove, groove isodiametrically reticulated. Last visible sternum with longitudinal carina at middle.

Legs: Profemora dorsally with a small pit and irregular reticulation. Protibia of moderate length, with short curved and flattened terminal spine, laterally with four teeth of decreasing size, dorsally and ventrally with two carinae. Movable spur short, length a third of first tarsomere. Meso- and metafemora of moderate length, slender, slightly dilated basally. First tarsomere distinctly elongated, almost as long as tarsomere two to five together.

External sexual dimorphism: Not observed.

Male genitalia (Fig. 62 View Figures 58–63 ): Median lobe conspicuously slender, elongated, in dorsal view regularly arcuate from base to apex, in lateral view slightly bisinuate, with single fine pili at beginning of apical third, apex spatulate, in cross section convex at apex, convexity directed dorsally. Oroficium of moderate size. Endophallus with two groups of microtrichia, a broader one with spine-like trichia, a small one at base of the first one at the opposite side. Dorsal paramere elongated, bisinuate, with elongated narrow apophyses; ventral one conspicuously small; both parameres slightly distorted.

Female genitalia (Fig. 77 View Figures 77–83 ): Coxostylus slender, obtusely curved, acute at apex, at end of basal third with one larger nematiform seta, five slender and one small nematiform setae laterally, SSO with one microtrichium.

Variation: The number of small tubercles at the lateral margin of the pronotum varies from eight to ten. In some specimens the tubercles are doubled.

Etymology.

The species name refers to its very slender shape and is expressed as a Latinised adjective (procerus, a, um = slender).

Distribution.

Known from the type locality at the Lac Calundo in the west of Angola. The specimens were found under stones.

Remark.

There is one additional female specimen from a different collecting locality with label data "9.XI.2011 ANGOLA, Bié Province, 50 km NEE Kuito. Chissamba Mission P. Schüle leg." (CBP). This specimen shares some characters with S. procera sp. nov. but it could not be determined with certainty and is therefore not included in the series of paratypes.