Scipopus (Parascipopus) savegre, Lindsay & Marshall, 2023

Lindsay, Kate & Marshall, Stephen A., 2023, A revision of Scipopus Enderlein including the subgenera Scipopus s. str., Phaeopterina Frey and Parascipopus subgen. nov. (Diptera, Micropezidae, Taeniapterinae), European Journal of Taxonomy 904, pp. 1-189 : 107-109

Scipopus (Parascipopus) savegre subgen. et sp. nov.

urn:lsid:zoobank.org:act:4A53A1DC-7234-46C8-B8D7-EE14C43966AA

Fig. 44

Differential diagnosis

Scipopus (Parascipopus) savegre sp. nov. resembles S. (Pa.) manifestus in having a primarily black head and body, but it differs in having a black first fore tarsomere and three indistinct subapical hyaline spots on the wing (often hard to see, as in Fig. 44F ).

Etymology

The name is a noun in apposition inspired by the type locality.

Type material examined

Holotype

COSTA RICA • 1 ♂; San José, San Gerardo de Dota, Savegre Lodge ; 9°33′14″ N, 83°47′48″ W; 4–8 Aug. 2010; S.A. Marshall leg.; MYCRO091-15/UGIC190-15 sequenced for CO1; MNCR (dissected and photographed, Fig. 44B, D ).

Paratypes

COSTA RICA • 1 ♀; San José; Mirador Quetzal , 19 km S of Empalme; 2600 m a.s.l.; May 2000; P. Hanson leg.; MYCRO090-15/UGIC189-15 sequenced for CO1; DEBU • 2 ♀♀; same collection data as for preceding; DEBU (dissected and photographed, Fig. 44C ) • 1 ♀; Prov. San José, Est. Cuerici, Sendero al Mirador , 4.6 km al E. de Villa Mills; 2640–2700 m a.s.l.; 9 Jun. 1996; A. Picado leg.; L_S_389700_499600; MNCR • 1 ♀; same collection data as for preceding; Canto de la Alves Trail; 19–21 Feb. 2008; MYCRO158-15/UGIC257-15 sequenced for CO1; DEBU (photographed, Fig. 44A ) • 1 ♂, 1 ♀; Prov. Puntarenas, Coto Brus, Zona Prot Las Tablas , Sura ; 2100 m a.s.l.; 29 Mar. 2000; M. Alfaro leg.; manual; L_S_323300_601500; MNCR (♀ photographed, Fig. 44E ) .

Description

LENGTH. 12–14 mm.

HEAD. Antenna black. Palpus black, pale microtrichose and setulose, wide (length 3.5 × height). Clypeus black, width ~1.9× height, silvery microtrichose on entire surface. Frontal vitta dark brown or black, entirely silvery microtrichose, posterior apex strongly tapered to a point. Orbital plate black, shiny, bare, clearly delineated. Epicephalon black, shiny, wide (width>⅔ of upper frontal vitta width at inner verticals), clearly delineated from upper frontal vitta. Paracephalon black. One lower fronto-orbital seta, outer vertical seta present or absent, all other head chaetotaxy well-developed.

THORAX. Scutum black, dark brown microtrichose. Female cervical sclerite convex. Postpronotal lobe black, densely setulose. Notopleuron black. Pleuron black, entirely white microtrichose, denser posteriorly, forming silvery transverse band only visible at some angles. Legs black, fore and hind tarsomere 1 black with very slight ventral golden fringe. Wing mostly dark black-brown infuscate with three indistinct subapical, slightly hyaline spots. Apical scutellar seta present.

ABDOMEN (J+ ♀). T1 with fine, long, dark brown or black setae.

FEMALE ABDOMEN. Pleuron mostly dark brown or black, microtrichose, pale grey or with white medial spots ventral to posterior edge of P1+2 and P4 ( Fig. 44F ). T1+2 ~2.3 × length of T3. Oviscape black, setulose, white microtrichose on anterior ventral and lateral ½, anteromedian ⅛; ~4.0–5.0× length of T6. Common duct short, ¼ of entire duct length. Paired spermathecal duct long (9.0 × length of paired spermathecae), narrow basally, gradually becoming wider distally. Paired spermathecal stems divided into very short, straight basal section and long, coiled and sinuate distal portion with sparsely distributed tubercles. Paired spermathecae cup-like (collapsed in Fig. 44C ) with prominent apical indentation. Single spermathecal duct arising from base of paired duct, narrow, <½ diameter and length of paired duct. Single spermatheca elongate, wider distally, inornate.

MALE ABDOMEN. Pleuron dark brown or black, microtrichose, anterior half of P3 light grey or white. T1+2 ~2.3 × length of T3. T6 short, genital fork 2.3× as long as T6, with arms converging, inner basal process angled inward, ⅓ length of arm. Epandrium elongate (length ~2.0 × height), short setose on posteroventral margin. Basiphallus small, crescent-shaped. Basal distiphallus short (~⅓ length of epandrium). Phallic bulb with upper chamber with rounded posterior projection and rounded lower chamber. Distal distiphallus long, narrow (≈ in length to epandrium), very slightly expanded apically. Phallapodeme broad and expanded apically. Anterior hypandrium expanded and broad.

Remarks

Scipopus (Parascipopus) savegre sp. nov. is known only from relatively high elevation forests, from 2100–2700 m.

Distribution

Costa Rica.

Figures

Fig. 44. Scipopus (Parascipopus) savegre subgen. et sp. nov. A. Head, ♀, lateral view, Costa Rica, paratype (DEBU). B. Male genital fork, ventral view, Costa Rica, holotype (MNCR). C. Female spermathecae and associated structures, Costa Rica, paratype (DEBU). D. Male terminalia, lateral view, Costa Rica, holotype (MNCR). E. Head and thorax, ♀, dorsal view, Costa Rica, paratype (MNCR). F. Living, J, Costa Rica, holotype (MNCR). Abbreviations: b = basiphallus; bdp = basal distiphallus; cd = common duct; ddp = distal distiphallus; e = epandrium; h = hypandrium; pb = phallic bulb; pd = paired spermathecal duct; ps = paired spermatheca; sd = single spermathecal duct; ss = single spermatheca.