Eutubercularium Brölemann, 1916

Wesener, Thomas, Akkari, Nesrine & Golovatch, Sergei I., 2025, Revision of the millipede family Dalodesmidae in Madagascar, with descriptions of two new Malagasy species of Dalodesmus Cook, 1896 (Diplopoda, Polydesmida), ZooKeys 1223, pp. 185-220 : 185-220

Genus Eutubercularium Brölemann, 1916

Eutubercularium Brölemann, 1916 ; type species Pterodesmus sakalava de Saussure & Zehntner, 1901 (currently Dalodesmus voeltzkowi Mesibov, Wesener & Hollier, 2018 ), by original designation, synonymised with Dalodesmus by Jeekel (1965), resurrected from synonymy following Hoffman (1980).

Diagnosis.

26 mm long, 4.5 mm wide, habitus identical to that of Dalodesmus , with five rows of transverse, often rather irregular rows of piligerous tuberculations or obliterated areations between always strongly developed and laterally similarly piligerous and crenulate / tuberculate paraterga.

The basic differences from Dalodesmus lie only in gonopodal conformation: femorites densely setose all over ventral and lateral sides, relatively stout, only ~ 2 × as long as acropodites and, much like in Phymatodesmus , clearly flattened dorsoventrally (vs less strongly setose to nearly bare, much longer, slender and subcylindrical); solenomere (sl) lateral, long, flagelliform, and non-sigmoid (vs medial, short, usually simple and rather rod- or lobe-shaped, often sigmoid), lateral branch (lb) remarkably tripartite and complex (vs lb unipartite and usually simple to rather simple).

Remarks.

Brölemann (1916) described Eutubercularium based on gonopod differences derived only from the drawings of the species published by de Saussure and Zehntner (1901: figs 8–10) for Polydesmus sakalava (now Dalodesmus voeltzkowi Mesibov, Wesener & Hollier, 2018 ). Apparently, the type specimens can no longer be located. Jeekel (1965) hesitantly decided to synonymise Eutubercularium with Dalodesmus , as the observed differences could be a simple drawing error. Hoffman (1980) listed Eutubercularium again as a valid genus, but without explanation, a move not followed in recent species lists ( Enghoff 2003; Wesener and Enghoff 2022). Here, we resurrect Eutubercularium from synonymy with Dalodesmus following Hoffman (1980).