Enghoffosoma

Likhitrakarn, Natdanai, Golovatch, Sergei I. & Panha, Somsak, 2014, Review of the Southeast Asian millipede genus Enghoffosoma Golovatch, 1993 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of new species, Zootaxa 3811 (4), pp. 491-514 : 511

Key to species of Enghoffosoma , chiefly based on male characters

1. Paraterga virtually absent; midbody legs very long, more than 2 times as long as body height. Southern China.... E. longipes

- Paraterga present, up to very strongly developed ( Figs 2 C, J, K, 8D, 11D, 14D & 17D); midbody legs rather short, less than 1.5 times as long as body height............................................................................. 2

2. Paraterga very strongly developed ( Figs 8 B–F, H, 17B–F & H)................................................. 3

- Paraterga rather well to poorly developed ( Figs 2 A–F, J–L, 11B–F, H, 14B-F & H)..................................4

3. Gonopod process e spear-shaped, pointed and short, directed ventrad ( Figs 9 & 10 ).................. E. lanceolatum sp. n.

- Gonopod process e long and spiniform, directed forward ( Figs 18 & 19 )............................ E. bispinum sp. n.

4. Solenomere subspiniform, twisted and bifid ( Figs 15 & 16 ).................................................... 5

- Solenomere finger-shaped, expanded apically ( Figs 3 A–C, 4–7, 12 & 13)......................................... 6

5. Solenomere long and clearly coiled, process e stout and rather short. Myanmar........................ .. E. spinipleurum

- Solenomere not coiled, process e slender, spiniform, and pointed ( Figs 15 & 16 )........................ .. E. funda sp. n.

6. Gonopod process p large, stout, anchor-shaped ( Figs 12 & 13 )................................. E. anchoriforme sp. n.

- Gonopod process p pointed, rather short or long ( Figs 3 A–C, 4A, B, 5, 6C & D), up to to broadly lobe- ( Figs 4 C, D & 7) or hook-shaped ( Figs 4 A, B & 6A, B)............................................................. E. zebra sp. n.

Figures

FIGURE 2. Enghoffosoma zebra sp. n., holotype (A – I), male paratype from Numtip Temple (J, K), male paratype from Ban Xanapho (L). A, B: anterior part of body, dorsal and lateral views, respectively. C: segments 10 and 11, dorsal view. D: segments 9 – 11, lateral view. E – G: posterior part of body, lateral, dorsal and ventral views, respectively. H, I: sternal cones between coxae 4, subcaudal and sublateral views, respectively. J, K: segments 10 and 11, dorsal view. L: anterior part of body, dorsal view.

FIGURE 3. Enghoffosoma zebra sp. n., holotype. A – C: right gonopod, lateral, mesal and submesal views, respectively. Scale bar: 0.5 mm. C: sterna of segment 10. E: leg of segment 10. Drawn not to scale. Designations explained in text.

FIGURE 4. Enghoffosoma zebra sp. n., male paratype from Numtip Temple (A, B), male paratype from Ban Xanapho (C, D). A, B: right gonopod, mesal and submesal views, respectively. C, D: right gonopod, mesal and submesal views, respectively. Scale bar: 0.5 mm. Designations explained in text.

FIGURE 8. Enghoffosoma lanceolatum sp. n., holotype. A: habitus, live coloration. B, C: anterior part of body, dorsal and lateral views, respectively. D, E: segments 10 and 11, dorsal and lateral views, respectively. F – H: posterior part of body, dorsal, ventral and lateral views, respectively. I, J: sternal cones between coxae 4, subcaudal and sublateral views, respectively.

FIGURE 9. Enghoffosoma lanceolatum sp. n., holotype. A – C: right gonopod, lateral, mesal and subcaudal views, respectively. Scale bar: 0.5 mm. Designations explained in text.

FIGURE 10. Enghoffosoma lanceolatum sp. n., holotype, left gonopod. A, B: lateral and mesal views, respectively. C – F: distal part, lateral, mesal, subcaudal and suboral views, respectively. Scale bar: 0.2 mm.

FIGURE 12. Enghoffosoma anchoriforme sp. n., holotype. A – C: right gonopod, lateral, mesal and subcaudal views, respectively. Scale bar: 0.5 mm. Designations explained in text.

FIGURE 13. Enghoffosoma anchoriforme sp. n., holotype, left gonopod. A, B: lateral and mesal views, respectively. C – F: distal part, lateral, mesal, subcaudal and suboral views, respectively. Scale bar: 0.2 mm.

FIGURE 15. Enghoffosoma funda sp. n., male holotype. A, B: right gonopod, lateral, mesal and caudal views, respectively. Scale bar: 0.5 mm. Designations explained in text.

FIGURE 16. Enghoffosoma funda sp. n., holotype, right gonopod. A, B: mesal and lateral views, respectively; C – F: distal part, mesal, lateral, subcaudal and suboral views, respectively. Scale bar: 0.2 mm.

FIGURE 18. Enghoffosoma bispinum sp. n., holotype. A – C: right gonopod, lateral, mesal and caudal views, respectively. Scale bar: 0.5 mm. Designations explained in text.

FIGURE 19. Enghoffosoma bispinum sp. n., holotype, right gonopod. A, B: mesal and lateral views, respectively. C – F: distal part, mesal, lateral, subcaudal and suboral views, respectively. Scale bar: 0.2 mm.