Metasarcus Roewer, 1913

Benedetti, Alipio Rezende & Pinto-da-Rocha, Ricardo, 2022, Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species, Arthropod Systematics & amp; Phylogeny 80, pp. 309-388 : 309

publication ID	https://dx.doi.org/10.3897/asp.80.e73829
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scientific name	Metasarcus Roewer, 1913
status

Treatment

3.39. Metasarcus Roewer, 1913

Figs 5 View Figure 5 , 10 View Figure 10 , 14C-H View Figure 14 , 19 View Figure 19 , 20 View Figure 20 , 25 View Figure 25 , 26 View Figure 26 , 31 View Figure 31

Metasarcus Roewer, 1913: 304 (desc); Roewer 1923: 508 (key), 517 (rdesc); Mello-leitão 1926: 34 (key); 1932: 408 (rdesc); 1935: 108 (cit.); Soares and Soares 1949: 234 (rdesc); Kury 1994: 349 (syst); Kury and Maury 1998: 145 (cit); Kury 2003: 145 (cat): Kury & Villarreal, 2015: 5, 14 (syst). Type-species: Metasarcus bolivianus Roewer, 1913 (by monotypy).

Metarascus [lapsus calami]: Roewer 1931: 107 (key).

Chaconatus Roewer, 1929: 275 (key), 276 (desc); Mello-leitão 1932: 108 (rdesc); Juberthie 1970: 142 (cit); Soares, Soares and Jim 1992: 5 (cat); Kury 1994: 349 (syst); Kury 2003: 145 (cat). Type-species: Chaconatus armatipalpus Roewer, 1929 (by monotypy). Synonymy established by Kury 1994: 349.

Chacoikeontus Roewer, 1929: 275 (key), 278 (desc); Mello-leitão 1932: 104 (key), 108 (rdesc); Juberthie 1970: 142 (cit); Soares, Soares and Jim 1992: 3 (key), 5 (rdesc, cat); Kury and Maury 1998: 145 (syst); Kury 2003: 144 (cat); Yamaguti and Pinto-da-Rocha 2009: 321 (syst); Kury & Villarreal, 2015: 23, 26 (cit). Type-species: Chacoikeontus clavifemur Roewer, 1929 (by monotypy). Synonymy established by Townsend et al. 2019: 102

Diagnosis.

Metasarcus can be diagnosed from other Metasarcidae genera with long legs (except Ayacucho ) by the combination of the following features: gamma-type; gamma-P-type or kappa-type DSS; ocularium unarmed; area III with two spines. Half of Metasarcus species present a spine, long projection or trifid projection in free tergite III, present in no other Metasarcidae species; coxa III with two apophyses; coxa IV reaching posterior margin of dorsal scutum or surpassing it; femur IV smooth or slightly granulate; penis stylus and VP thin thickness, with less than 10 MS C

Redescription.

Gamma-type DSS ( M. bolivianus and M. fellinii sp. nov.; Fig. 6E View Figure 6 ), gamma-P-type DSS ( M. beni sp. nov., M. bergmani sp. nov., M. limachii sp. nov. and M. trispinosus sp. nov.; Fig. 6A, B, G, H View Figure 6 ) or kappa-type DSS ( M. clavifemur , M. kurosawai sp. nov. and M. vacafloresae sp. nov.; Fig. 6D, F, I View Figure 6 ). Ocularium low, medially depressed. Ocularium unarmed, small tuberculate or smooth (Fig. 6 View Figure 6 ). Areas of dorsal scutum moderately or densely tuberculate ( M. fellinii sp. nov.). Area I undivided (Fig. 6G-I View Figure 6 ) or divided in two halves (Fig. 6A-F View Figure 6 ). Area III armed with two high spines in most species (e.g. Fig. 6D View Figure 6 ), with two tubercles ( M. trispinosus sp. nov.; Fig. 6H View Figure 6 ) or unarmed ( M. vacafloresae sp. nov.; Fig. 6I View Figure 6 ). Posterior margin unarmed. Coxa IV reaching area III-IV or posterior margin. Coxa IV unarmed or with retrolateral armature ( M. bergmani sp. nov., A. limachii sp. nov.; Fig. 6B, G View Figure 6 ). Femur IV much longer than dorsal scutum length. (Figs 10 View Figure 10 , 25 View Figure 25 , 26 View Figure 26 ) Less than 10 MS C. Penial stylus thin. Penial ventral plate thin (Figs 14C-H View Figure 14 , 19 View Figure 19 , 20 View Figure 20 ).

Distribution.

(Fig. 31 View Figure 31 ) BOLIVIA. Beni, Cochabamba, La Paz and Tarija.

Remarks.

Townsend et al. (2019: pp. 102-103, 105-106), in a non-taxonomic or systematic study, cited Chacoikeontus clavifemur as Metasarcus clavifemur (Roewer, 1929). The authors did not provide justification for or specify it was a new combination or nomenclatural act, did not mention the genus Chacoikeontus (and its consequent status as a junior subjective synonym) and did not discuss the genus Metasarcus with the inclusion of the species. Once the new combination has been published, the nomenclatural act is valid and herein confirmed (see below). As Chacoikeontus clavifemur is the type-species of the genus, by implication Chacoikeontus was considered a junior subjective synonym of Metasarcus .

Species composition.

Metasarcus beni sp. nov.; Metasarcus bergmani sp. nov.; Metasarcus bolivianus Roewer, 1913; Metasarcus clavifemur (Roewer, 1929); Metasarcus fellinii sp. nov.; Metasarcus kurosawai sp. nov.; Metasarcus limachii sp. nov.; Metasarcus trispinosus sp. nov. and Metasarcus vacafloresae sp. nov.