Paraclavelia arabiae Shimizu, 2022

Shimizu, Akira, Broad, Gavin, Yoshimura, Jin & Pitts, James P., 2022, First records of the spider wasps Ctenocerus Dahlbom and Paraclavelia Haupt from Asia, with discussions on the systematics of Ctenocerinae (Hymenoptera: Pompilidae), European Journal of Taxonomy 845 (1), pp. 101-131 : 118-120

Paraclavelia arabiae Shimizu sp. nov.

urn:lsid:zoobank.org:act:C089DF38-E7E2-48D2-90BF-7549FF207C63

Fig. 7

Diagnosis

Female

Body and legs predominantly black ( Fig. 7A ); clypeus ( Fig. 7D ), antenna ( Fig. 7A, E ), mandible and legs except coxae ( Fig. 7G ) reddish brown; T6 and S6 ( Fig. 7I ) orange brown. FW bifasciate ( Fig. 7C ), outer fascia very broad. L-shaped groove on pronotum laterally obsolete ( Fig. 7H ). FW discal cell 1 without irregularity in membrane basally. T1–3 with posterior silvery bands of short, dense, appressed pubescence ( Fig. 7A ). Scape without longitudinal carina on its mesal face. T1 barely petiolate. S2 with transverse groove fine and almost obsolete ( Fig. 7I ).

Etymology

The species name is derived from its locality, the Arabian Peninsula.

Type material

Holotype OMAN • ♀; “ OMAN Ayn Razat 9.iv.85 H. Hamer ” [in handwriting]; NHMUK.

Description

Holotype female

MEASUREMENTS. Length: body 9.4 mm; FW 5.9 mm.

COLOURATION. Body black ( Fig. 7A ). Following reddish brown: lower frons ( Fig. 7D ), including supraantennal area below, clypeus, malar space, lower gena, antenna, mandible, pronotum anteriorly ( Fig. 7B, E ) and lateroventrally ( Fig. 7H ), apical margins of T3–5 and of S1–5 ( Fig. 7I ), and all legs excluding coxae (dark brown) ( Fig. 7G–H ). Labrum, labio-maxillary complex, T6 and S6 orange brown. FW transparent with two fasciae ( Fig. 7C ): inner fascia occupying on second abscissae of vein M and vein Rs (basal vein) and cross-vein cu-a, and apical half of clavus; outer fascia very broad, occupying marginal cell, SMC1 apically, SMCs 2 and 3, basal half of SMC4, distal ¾ of discal cell 2, discal cell 3, and subdiscal cell 2 except basally; outer wing margin slightly infuscate. HW transparent, broadly infuscate apical to cross-vein rs-m.

INTEGUMENT. Head and pro-, meso- and metanota with minute dense punctures (sparser on clypeus). Pronotal collar with several transverse rugulae ( Fig. 7E ). Mesopleuron posteriorly and metapleuron finely and obliquely rugulose. Side of metanotum with several oblique striae. Metapostnotum with few undulating striae, lacking median depression. Propodeum with transverse rugulae ( Fig. 7B ), those becoming stronger posteriorly, lacking median groove. Most of integument with short, appressed silvery pubescence, dense on lower frons, clypeus basally, gena ventrally, thorax laterally and ventrally, disc of metanotum, propodeum posterolaterally, coxae, posterior third of T1, posterior fourth of T2, and T3 narrowly along posterior margin ( Fig. 7A ). Setae on body scarce; frons along inner orbit, labrum apically, mandible, propleuron, T6 and S2–6 with sparse setae.

HEAD. 0.96 ×as broad as high. Vertex strongly convex above level of eye tops ( Fig. 7D ). Vertex and frons weakly polished. MID 0.65 ×TFD. Frons with median sulcus short and faint only below. Antennocular line slightly inclined from antennal base to eye ( Fig. 7E ). Inner orbits subparallel but barely emarginate above middle ( Fig. 7D ). UID:MID:LID = 8.5:10:9.7. POD:OOD = 1:0.63. Clypeus 1.5× as broad as long; anterior rim not depressed but smooth and polished; apical margin slightly convex. OOcD /POD = 0.94. Gena, in dorsal view, gradually narrowing posteriorly ( Fig. 7E ), in profile, 0.77 × as broad as eye, abruptly narrowing above ( Fig. 7H ). Scape flattened mesally ( Fig. 7E ). Scape:pedicel:fl1:fl2 = 12:3.2:10:8.5. Fl1 3.1× as long as broad, 0.49 × UID.

MESOSOMA. Pronotum with posterior margin subangulate medially ( Fig. 7B ). Metapostnotum 0.25 × as long as metanotum at midline, its posterior margin almost straight medially. Propodeum in dorsal view 1.1× as broad as long; dorsum barely convex laterally in dorsal view ( Fig. 7B ), in profile roundly merging into declivity ( Fig. 7H ).

LEGS. Mid tibia with few minute spines laterally. Hind tibia almost lacking spines dorsally ( Fig. 7J ) with spur 0.46 × as long as hind tarsomere 1.

WINGS ( Fig. 7C ). FW marginal cell from wing tip by 0.65× its own length. SMC2:SMC3 = 1:1.1 on vein M, 1:1.5 on vein Rs. SMC2 narrowed on vein Rs by 0.57 × its length on vein M, receiving cross-vein 1m-cu at basal 0.45. SMC3 narrowed on vein Rs by 0.75× its length on vein M, receiving cross-vein 2m-cu at basal 0.53. Cross-veins 2rs-m and 3rs-m gently curved outwardly. Cross-vein cu-a slightly apical to point of separation of vein M+CuA, oblique but abruptly recurved near its terminal. Cross-vein 2m-cu strongly curved near its origin. HW cross-vein cu-a originating at point of separation of vein M+CuA.

METASOMA. S6 somewhat compressed laterally with median carina only apically ( Fig. 7I ).

Male

Unknown.

Distribution

The southeastern Arabian Peninsula ( Oman).

Figures

Fig. 7. Paraclavelia arabiae Shimizu sp. nov., holotype, ♀ (NHMUK). A. Whole body, dorsal view. B. Head and mesosoma, dorsal view. C. Right wings. D. Head, anterior view. E. Head and pronotum, dorsal view. F. Apical part of right fore tibia, mesial view, arrow indicates apicomesial spine on fore tibia. G. Whole body, lateral view. H. Head and mesosoma, lateral view. I. Metasoma, ventrolateral view. J. Right hind tibia, dorsal view. Scale bars: A–E, G–J = 1 mm; F = 0.5 mm.