Hypherpes (Cryobius), Chaudoir, 1838

Bousquet, Yves, 2012, Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico, ZooKeys 245, pp. 1-1722 : 543-544

Subgenus Cryobius Chaudoir, 1838

Cryobius Chaudoir, 1838: 11. Type species: Poecilus ventricosus Eschscholtz, 1823 by original designation. Etymology (original). From the Greek cryos (cold) and bios (life), alluding to the cold habitat where these species live [masculine].

Haptoderus Chaudoir, 1838: 10. Type species: Feronia spadicea Dejean, 1828 (= Feronia pumilio Dejean, 1828) by original designation. Synonymy established by Bousquet (1999: 183). Etymology (original). From the Greek hapto (fasten to, by extension to apply) and dere (neck, by extension pronotum) [masculine].

Pseudorthomus Chaudoir, 1838: 12, 19. Type species: Feronia amaroides Dejean, 1828 by original designation. Synonymy established by Bousquet (1999: 183). Etymology (original). From the Greek pseudos (fallacy, lie) and the generic name Orthomus [masculine].

Pseudocryobius Motschulsky, 1850a: ix. Type species: Feronia nivalis Sahlberg, 1844 designated by Bousquet (1984a: 4). Synonymy established by Poppius (1906b: 7). Etymology (original). From the Greek pseudos (fallacy, lie) and the generic name Cryobius [q.v.] [masculine].

Abaxodes Gistel, 1857: 31. Type species: Feronia abaxoides Dejean, 1828 designated by Strand (1917: 76). Etymology. From the generic name Abax [q.v.] and the Greek - odes (likeness) [masculine].

Orites Schaum, 1858: 442, 463. Type species: Platysma negligens Sturm, 1824 by monotypy. Synonymy established by Schatzmayr (1942: 77). Etymology. From the Greek oreites (mountaineer) [masculine].

Pyreneorites Jeannel, 1937b: 11, 16. Type species: Feronia pusilla Dejean, 1828 by original designation. Synonymy established by Bousquet (1999: 183). Etymology. From the geographical name Pyrenees and the generic name Orites [q.v.] [masculine].

Euhaptoderus Jeanne, 1969: 33. Type species: Carabus unctulatus Duftschmid, 1812 by original designation. Synonymy established by Bousquet (1999: 183). Etymology. From the Greek eu (agreeable, original, primitive) and the generic name Haptoderus [q.v.] [masculine].

Iberoderus Jeanne, 1969: 34. Type species: Feronia nemoralis Graells, 1851 by original designation. Synonymy established by Bousquet (1999: 183). Etymology. From the geographic name Iberia (the Iberian Peninsula) and the Greek dere (neck, by extension pronotum) [masculine].

Cryobiopterus O. Berlov, 1997: 36. Type species: Argutor brevicornis Kirby, 1837 by original designation. Synonymy established by Bousquet (1999: 183). Etymology. From the Greek cryos (cold), bios (life), and pteron (wing, by extension elytron) [masculine].

Diversity.

About 115 species in the arctic, subarctic, and boreal regions of North America (23 species) and the Palaearctic Region (about 100 species). Several species are Holarctic.

Identification.

Ball (1966a) revised the North American species which he arrayed in three groups. Since then, two new species have been described, one ( Pterostichus haftorni ) by Lindroth (1969a) which has been subsequently synonymized, the other one ( Pterostichus woodi ) by Ball and Currie (1997), and several species related to Pterostichus planus have been downgraded to subspecies of Pterostichus bryanti by Ball and Currie (1997). Species identifications are difficult in part because of the important intraspecific variation observed in most species.

Taxonomic Note.

Lorenz (2005: 275) listed Parahaptoderus Jeanne as a junior synonym of Cryobius . However, the taxon, which is composed of three European species of which one extends to Turkey, is probably not closely related to Cryobius (see Bousquet 1999: 187).