VERMETIDAE Rafinesque, 1815

Coan, Eugene V., Pacaud, Jean-Michel & Kabat, Alan R., 2024, Gérard-Paul Deshayes (1796 - 1875) and his taxa, Geodiversitas 46 (16), pp. 553-930 : 620-621

Family VERMETIDAE Rafinesque, 1815

Tubispiracea – Deshayes, 1832-EncyMeth: table facing p. 553, as “les Tubispirés”, later latinized by Reeve (1882), but unavailable because not based on a generic name (ICZN Code Art. 11.7.1.1).

Tubispirata – Deshayes, 1861-DescrCoqFoss: 279-290. Emendation or spelling error for Tubispiracea Deshayes, 1832, q.v. Listed by Bieler & Petit (2011: 12).

anguillinus, Serpulorbis – Deshayes, 1861-DescrCoqFoss: 289-290, 9, pl. 9, figs 16-17, pl. 10, figs 5-6. — Type localities: Mercin-et-Vaux, Laon, Cuise-la-Motte, Mons-en-Laonnois & Hérouval. — Type age: Eocene (Ypresian). — Type species (OD) of Anguillospira Cossmann, 1912 . Vermicularia (Anguillospira) anguillina (Deshayes, 1861) ( Le Renard & Pacaud 1995: 105) . Listed by Bieler & Petit (2011: 24). Anderson & Allmon (2023: 8) referred this taxon to the Siliquariidae .– Current status: Anguillospira anguillina (Deshayes, 1861) (Pacaud herein).

cancellatus , Serpulorbis – Deshayes, 1861-DescrCoqFoss: 284-285, 9, pl. 9, fig. 8. — Type localities: Auvers-sur-Oise, Valmondois, Mary-sur-Marne, Jaignes, Crouy & Vendrest. — Type age: Eocene (Bartonian). — Current status: Serpulorbis cancellatus Deshayes, 1861 ( Le Renard & Pacaud 1995: 105). Listed by Bieler & Petit (2011: 28).

clathratus , Serpulorbis –Deshayes, 1861-DescrCoqFoss: 286-287, 9, pl. 9, figs 9-10. — Type localities: Auvers-sur-Oise, Vendrest, Crouy, Acy-en-Multien, Betz, Caumont & Mary-sur-Marne. — Type age: Eocene (Bartonian). Listed by Bieler & Petit (2011: 30). — Current status: Serpulorbis clathratus Deshayes, 1861 ( Le Renard & Pacaud 1995: 105; Pacaud 2008: 57).

cristatus , Serpulorbis – Deshayes, 1861-DescrCoqFoss: 287, 9, pl. 9, fig. 12, non Vermetus cristatus Sandberger, 1860 , nec Biondi, 1859. — Type localities: Auvers-sur-Oise & Mary-sur-Marne. — Type age: Eocene (Bartonian). — Type material: UCBL-EM 32398, syntype from Auvers-sur-Oise. Vermetus excristatus Sacco, 1896 ( Sacco 1896: 7), replacement name ( Bieler & Petit, 2011: 36). Serpulorbis cristatus Deshayes, 1861 ( Le Renard & Pacaud 1995: 105; Pacaud 2008: 57). Listed by Bieler & Petit (2011: 33). — Current status: synonym of Thylacodes excristatus ( Sacco, 1896) .

laxatus , Serpulorbis – Deshayes, 1861-DescrCoqFoss: 287-288, 9, pl. 9, fig. 15. — Type locality: Mary-sur-Marne. — Type age: Eocene (Bartonian). Listed by Bieler & Petit (2011: 44). — Current status: Serpulorbis laxatus Deshayes, 1861 ( Le Renard & Pacaud 1995: 105; Pacaud 2008: 57).

masier , Vermetus – Deshayes, 1843-HistNatAnim2: 65, ex Adanson ms; 1843-HistNatAnim3: 526. In the synonymy of Serpula sipho Lamarck, 1818 . West Africa. — Type age: Recent. Listed by Bieler & Petit (2011: 44, 46). — Current status: Thylacodes masier (Deshayes, 1843) (MolluscaBase) .

moerchi , Serpulorbis – Deshayes, 1861-DescrCoqFoss: 286, 9, as “ morchi ”, pl. 9, figs 21-22. — Type localities: Mercin-et-Vaux, Cuise-la-Motte & Laon. — Type age: Eocene (Ypresian). Stated to be named for Mörch, so automatically corrected (ICZN Code Art. 32.5). Serpulorbis moerchi (Deshayes, 1861) ( Le Renard & Pacaud 1995: 105, as “ S. morchi ”). Listed by Bieler & Petit (2011: 30). — Current status: Thylacodes moerchi (Deshayes, 1861) ( Leroy et al. 2014: 25, pl. 21, figs 2-5).

ornatus , Serpulorbis – Deshayes, 1861-DescrCoqFoss: 285, 9, pl. 9, fig. 23, non S. ornata (J. de C. Sowerby, 1850) [ Serpula ]. — Type localities: Montmirail & Hautteville-Bocage. — Type age: Eocene (Lutetian). — Type material: UCBL-EM 32400, syntypes from Montmirail. Replaced as Thylacodes deshayesi Newton, 1891 ( Newton 1891: 217). However, the latter is a secondary homonym of Serpulorbis deshayesi Mayer-Eymar, 1889 , which is also a Thylacodes . Sacco (1896: 55) proposed as a replacement name Vermetus deshayesianus . A third replacement name, Serpulorbis newtoni Glibert, 1962b ( Glibert 1962b: 130), for Serpulorbis ornatus Deshayes, 1861 , was unnecessary. All these taxa are Thylacodes . The name Serpula ornatus J. de C. Sowerby, 1850 , junior primary homonym of Serpula ornata I. Lea , 1833, is now Vermetus compsus Cossmann, 1892 ( Cossmann 1892: 60-61), a replacement name. It is not a nomen nudum, as Bieler & Petit (2011: 30) asserted. Serpulorbis ornatus Deshayes , listed by Bieler & Petit (2011: 50). — Current status: Thylacodes deshayesianus ( Sacco, 1896) (Pacaud herein).

polygonus, Serpulorbis – Deshayes, 1861-DescrCoqFoss: 290, 9, pl. 9, fig. 14. — Type localities: Chambors, Hérouval, Thiverval-Grignon & Valmondois. — Type age: Eocene (Lutetian-Bartonian). Vermicularia (Anguillospira) polygona (Deshayes, 1861) ( Le Renard & Pacaud 1995: 105; Pacaud 2008: 49). Listed by Bieler & Petit (2011: 53). — Current status: Anguillospira polygona (Deshayes, 1861) (Pacaud herein).

porrectus, Serpulorbis – Deshayes, 1861-DescrCoqFoss: 288-289, 9, pl. 9, fig. 18. — Type localities: Acy-en-Multien, Crouy & La Ferté-sous-Jouarre. — Type age: Eocene (Bartonian). — Type material: UCBL-EM 32399, syntype from Acy-en-Multien. Vermicularia (Anguillospira) porrecta (Deshayes, 1861) ( Le Renard & Pacaud 1995: 105) . Listed by Bieler & Petit (2011: 53). — Current status: Anguillospira porrecta (Deshayes, 1861) (Pacaud herein).

semipedalis , Serpulorbis – Deshayes, 1861-DescrCoqFoss: 288, 9, pl. 9, fig. 11. — Type localities: Valmondois, Auvers-sur-Oise, Crouy & Le Fayel. — Type age: Eocene (Bartonian). — Type material: UCBL-EM 32397, syntype from Auvers-sur-Oise. — Current status: Serpulorbis semipedalis Deshayes, 1861 ( Le Renard & Pacaud 1995: 105). Listed by Bieler & Petit (2011: 58).

serpuloides , Serpulorbis – Deshayes, 1861-DescrCoqFoss: 289, 9, pl. 9, figs 19-20. — Type localities: Thiverval-Grignon, Mouchy-le-Châtel & Parnes. — Type age: Eocene (Lutetian). Vermicularia (Anguillospira) serpuloides (Deshayes, 1861) ( Le Renard & Pacaud 1995: 105; Pacaud 2008: 49). Listed by Bieler & Petit (2011: 58). — Current status: Thylacoides serpuloides (Deshayes, 1861) (Pacaud herein).

strictus , Serpulorbis – Deshayes, 1861-DescrCoqFoss: 285-286, 9, pl. 9, fig. 13. — Type locality: Auvers-sur-Oise. — Type age: Eocene (Bartonian). Listed by Bieler & Petit (2011: 61). — Current status: Serpulorbis strictus (Deshayes, 1861) ( Le Renard & Pacaud 1995: 105) .

turonensis , Vermetus –Deshayes, 1850-TraitElem: 43, pl. 70, figs 14- 15. — Type locality: none given [ France]. — Type age: Miocene (Aquitanian).Listed by Bieler & Petit (2011: 66). — Current status: Vermetus (Lemintina) arenarius [Linnaeus, 1758; Serpula ] turonensis Deshayes, 1850 ( Cossmann & Peyrot 1922: 81-82, pl. 3, figs 18- 20, as “1839”).

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arenarius , Vermetus – Deshayes, 1838-RegAnim: pl. 62, fig. 5; 1850-TraitElem: 43, pl. 70, fig. 16. — Type locality: none given. — Type age: probably Recent. Listed by Bieler & Petit (2011: 26) as a separate species. However, the plate explanation and caption with his original figure in his makes it clear that this name was labeled as “Desh” because it was a generic transfer of Serpula arenaria Linnaeus, 1758 , to Vermetus . Bieler & Petit (2011: 26, 70-71) regarded what they thought was Deshayes’s species as a nomen dubium . Deshayes’ figure in 1838 of an animal was copied from Philippi (1836: pl. 9, fig. 18a), which Philippi had had labeled Vermetus gigas Bivona Bernardi, 1832 . Deshayes evidently considered this taxon to be a junior synonym of Linnaeus’ species. His later figure in the Traité was only of a shell. This species is now listed as Thylacodes arenarius (Linnaeus, 1758) ; Linnaeus’ species is the senior synonym of the type species of Thylacodes Guettard, 1770 .

lispe, Vermet [us] – See under Annelida .

tricarinatus , Vermetus – Menke, 1844: 23, as “Deshayes, 1843”, nomen nudum.