CANCELLARIIDAE Forbes & Hanley, 1851

Coan, Eugene V., Pacaud, Jean-Michel & Kabat, Alan R., 2024, Gérard-Paul Deshayes (1796 - 1875) and his taxa, Geodiversitas 46 (16), pp. 553-930 : 638-639

Family CANCELLARIIDAE Forbes & Hanley, 1851

angulifera , Cancellaria – Deshayes, 1864-DescrCoqFoss: 107, 73, pl. 73, figs 13-15. — Type locality: Le Fayel. — Type age: Eocene (Bartonian). — Type material: UCBL-EM 32702, holotype. Listed by Petit & Harasewych (2005: 24). — Current status: Unitas angulifera (Deshayes, 1864) ( Le Renard & Pacaud 1995: 121) .

asperula , Cancellaria – Deshayes, 1830a-EncyMeth: 187; Deshayes in Guérin-Méneville 1835: pl. 17, fig. 3, 3a; 1843-HistNatAnim2: 411; 1843-HistNatAnim3: 646-647;Deshayes in Guérin-Méneville 1844: 33; Guérin-Méneville & Deshayes 1868: pl. 16, fig. 3, 3a, as C. asperella (Lamarck, 1822) . — Type locality: none given. — Type age: Recent. Listed by Petit & Harasewych (2005: 26). — Current status: synonym of the Indo-Pacific Scalptia obliquata (Lamarck, 1822) [ Cancellaria ], type species (OD) of Scalptia Jousseaume, 1887 .

basteroti , Cancellaria –Deshayes, 1864-DescrCoqFoss: 104. — Type locality: Bordeaux. — Type age: Miocene (Burdigalian). — Type material: MNHN.F.A70860, syntype. Synonym of Merica contorta ( Basterot, 1825) [ Cancellaria ] ( Peyrot 1928: 206-208, pl. 12, fig. 41 as “1860”). — Current status: Contortia contorta ( Basterot, 1825) ( Cahuzac et al. 2004: 235-236) .

bifasciata, Cancellaria –Deshayes, 1830a-EncyMeth: 181;1843-HistNatAnim2: 413; 1843-HistNatAnim3: 647. — Type locality: none given. —Type age: Recent. —Type material:MNHN-IM-2000-2120, holotype. — Current status: synonym of the Indo-Pacific Merica oblonga (G. B. Sowerby I, 1825) [ Cancellaria ] ( Petit & Harasewych 2005: 30).

canaliculata , Cancellaria – Deshayes, 1864-DescrCoqFoss: 97, 72, pl. 72, figs 26-28, non M. Hörnes, 1854. — Type locality: Parnes. — Type age: Eocene (Lutetian). — Type material: UCBL-EM 30567, syntype. Cancellaria rhabdota Bayan, 1873a ( Bayan 1873a: 103-104), replacement name. Listed byPetit & Harasewych (2005: 34, 90). — Current status: Unitas rhabdota (Bayan, 1873) ( Le Renard & Pacaud 1995: 121) .

crenulata , Cancellaria –Deshayes, 1835-DescrCoqFoss: 501-502, 42, pl. 79, figs 31-33; 1843-HistNatAnim2: 429; 1843-HistNatAnim3: 653. 1864-DescrCoqFoss: 99. — Type localities: Retheuil & Cuise-la-Motte. — Type age: Eocene (Ypresian). Preoccupies Cancellaria crenulata A. Adams, 1855 , which remains an unrenamed nomen dubium ( Petit & Harasewych 2005: 41) . — Current status: Unitas crenulata (Deshayes, 1835) ( Le Renard & Pacaud 1995: 121) .

dactyliosa , Rissoa – Deshayes, 1861-DescrCoqFoss: 403, 22, pl. 22, figs16-19.— Type locality:Aizy-Jouy.— Type age:Eocene (Ypresian).— Type species (OD) of Pezantia Cossmann, 1896 . — Current status: Pezantia dactyliosa (Deshayes, 1861) ( Le Renard & Pacaud 1995: 100) .

delecta , Cancellaria – Deshayes, 1864-DescrCoqFoss: 99. Replacement name for Cancellaria elegans Deshayes, 1835 (see below), non G. B. Sowerby I, 1821. Listed by Petit & Harasewych (2005: 43). — Current status: Unitas delecta (Deshayes, 1864) ( Le Renard & Pacaud 1995: 121; Pacaud 2008: 71).

dentifera , Cancellaria – Deshayes, 1864-DescrCoqFoss: 98, 73, pl. 73, figs 8-10. — Type localities: Aizy-Jouy & Cuise-la-Motte. — Type age: Eocene (Ypresian). — Type material: UCBL-EM 30576 syntype from Cuise-la-Motte. Listed by Petit & Harasewych (2005: 94). — Current status: Unitas dentifera (Deshayes, 1864) ( Le Renard & Pacaud 1995: 121; Pacaud 2008: 71).

dubia , Cancellaria – Deshayes, 1864-DescrCoqFoss: 105-106, 73, pl. 73, figs 25-27. — Type locality: Brasles. — Type age: Eocene (Lutetian). Bonellitia dubia (Deshayes, 1864) ( Le Renard & Pacaud 1995: 121) . Listed by Petit & Harasewych (2005: 47). — Current status: Admetula dubia (Deshayes, 1864) ( Pacaud 2008: 71) .

elegans , Cancellaria –Deshayes, 1835-DescrCoqFoss: 502, 42, pl. 79, figs 24-26, non G. B. Sowerby I, 1821;1843-HistNatAnim2: 429-430; 1843-HistNatAnim3: 653-654.— Type localities: Thiverval-Grignon, Parnes & Senlis. — Type age: Eocene (Lutetian-Bartonian). — Type material: UCBL-EM 30553-30566, syntypes. Oddly, Crosse (1861) renamed Sowerby’s species as Cancellaria reeveana on the grounds of it being preoccupied by that of Deshayes, “1824” ( Breure et al. 2022a: 189). Listed by Petit & Harasewych (2005: 48). Cancellaria delecta Deshayes, 1864 , replacement name (see above).

fusiformis , Cancellaria – Deshayes, 1864-DescrCoqFoss: 102-103, 72, pl. 72, figs 31-32, non Cantraine, 1835. — Type locality: Thiverval-Grignon. — Type age: Eocene (Lutetian). Listed by Petit & Harasewych (2005: 53). — Current status: Unitas beui Le Renard, 1994 ( Le Renard 1994: 38), replacement name ( Le Renard & Pacaud 1995: 121; Pacaud 2008: 71).

granifera , Cancellaria –Deshayes, 1830a-EncyMeth: 183;1835-DescrCoqFoss: 500-501, 42, pl. 79, figs 34-35. — Type localities: Thiverval-Grignon & Parnes. — Type age: Eocene (Lutetian). — Type material: UCBL-EM 30549-30552 & 30600-30603, syntypes from Thiverval-Grignon. Listed byPetit & Harasewych (2005: 55). — Current status: synonym of Unitas suturalis (G. B. Sowerby, I, 1822) (Pacaud herein). interrupta , Cancellaria – Deshayes, 1864-DescrCoqFoss: 100, 73, pl. 73, figs 5-7. — Type locality: Aizy-Jouy. — Type age: Eocene (Ypresian). — Type material: UCBL-EM 30577-30579, syntypes. Listed by Petit & Harasewych (2005: 59). — Current status: Unitas interrupta (Deshayes, 1864) ( Le Renard & Pacaud 1995: 121) .

lactea , Cancellaria –Deshayes, 1830a-EncyMeth: 180-181;1843-HistNatAnim2: 412; 1843-HistNatAnim3: 647. — Type locality: none given.— Type age:Recent.— Type material:MNHN-IM-2000-2150, syntype. — Current status: the Australian Sudaphera lactea (Deshayes, 1830) ( Wilson 1994: 174, pl. 37, fig. 23, as Cancellaria ; Petit & Harasewych 2005: 62).

milletii, Cancellaria – Deshayes, 1830a-EncyMeth: 181. — Type locality: Angers. — Type age: Miocene( Petit & Harasewych, 2005: 70). Does not preoccupy Cancellaria (Merica) milleti Brébion (1964: 522) , which is unavailable, having appeared in an unpublished thesis ( Landau et al. 2019: 142). — Current status: taxon inquirendum.

nana , Cancellaria – Deshayes, 1864-DescrCoqFoss: 106-107, 73, pl. 73, figs 11-12. — Type localities: Laversine, Retheuil & Mercin-et-Vaux. — Type age: Eocene (Ypresian). — Type material: UCBL-EM 30597-30599, syntypes from Laversine or Mercin-et-Vaux. Listed by Petit & Harasewych (2005: 74). — Current status: Sveltella nana (Deshayes, 1864) ( Le Renard & Pacaud 1995: 121; Pacaud 2008: 71).

obtusa , Cancellaria –Deshayes,1830a-EncyMeth: 187-188;1843-HistNatAnim2: 417;1843-HistNatAnim3: 649;1838-RegAnim: pl. 52, fig. 6, 6a; 1849-DictUnivAtlas: 6, pl. 21, fig. 4. — Type locality: none given. — Type age: Recent. — Type material: NHMUK 1964454, holotype. — Current status: the Panamic Cancellaria (Massyla) obtusa (Deshayes, 1830) ( Keen 1971: 654; Petit & Harasewych 2005: 76; Landau et al. 2012: 225).

ornata , Cancellaria – Deshayes, 1864-DescrCoqFoss: 101, 73, pl. 73, figs 19-20. — Type locality: Cuisy-en-Almont. — Type age: Eocene (Ypresian). Listed by Petit & Harasewych (2005: 77). — Current status: Unitas ornata (Deshayes, 1864) ( Le Renard & Pacaud 1995: 121) .

quantula , Cancellaria – Deshayes, 1864-DescrCoqFoss: 106, 72, pl. 72, figs 29-30. — Type localities: Thiverval-Grignon, Parnes & Chaussy. — Type age: Eocene (Lutetian). — Type material: UCBL-EM 30582-30596, syntypes. — Type species (OD) of Sveltella Cossmann, 1889 . Listed by Petit & Harasewych (2005: 18, 88). — Current status: synonym of Sveltella biplicata (Lamarck, 1803) ( Le Renard & Pacaud 1995: 121; Pacaud 2008: 71).

scabra, Cancellaria – Deshayes, 1830a-EncyMeth: 190; 1843-HistNatAnim2: 425; 1843-HistNatAnim3: 652. — Type locality: Plaisantin. — Type age: Pliocene (Piacenzian-Zanclean). Listed by Petit & Harasewych (2005: 92). — Current status: synonym of Trigonostoma umbilicare ( Brocchi, 1814) ( Brunetti et al. 2011: 112) .

separata , Cancellaria – Deshayes, 1864-DescrCoqFoss: 97-98, 72, pl. 72, figs 20-22. — Type localities: Thiverval-Grignon, Parnes, Chaussy, Damery & Saint-Félix. — Type age: Eocene (Lutetian). — Type material: UCBL-EM 30568-30575, syntypes. Listed by Petit & Harasewych (2005: 42). — Current status: Unitas separata (Deshayes, 1864) ( Le Renard & Pacaud 1995: 121; Pacaud 2008: 71).

speciosa , Cancellaria – Deshayes, 1864-DescrCoqFoss: 100-101, 73, pl. 73, figs 1-3. — Type locality: Vregny. — Type age: Eocene (Ypresian). — Type material: UCBL-EM 30580, holotype. Listed by Petit & Harasewych (2005: 96). — Current status: Unitas speciosa (Deshayes, 1864) ( Le Renard & Pacaud 1995: 121) .

spectabilis , Cancellaria – Deshayes, 1864-DescrCoqFoss: 102, 72, pl. 72, figs 23-25. — Type locality: Mouchy-le-Châtel. — Type age: Eocene (Lutetian). — Type material: UCBL-EM 30581, holotype. Listed by Petit & Harasewych (2005: 96). — Current status: Unitas spectabilis (Deshayes, 1864) ( Le Renard & Pacaud 1995: 121; Pacaud 2008: 71).

spengleriana , Cancellaria – Deshayes, 1830a-EncyMeth: 185-186; 1843-HistNatAnim2: 415; 1843-HistNatAnim3: 648. — Type locality: none given; Lisbonne collection. — Type age: Recent. — Type material: MNHN-IM-2000-2174, holotype. — Type species (OD) of Sydaphera Iredale, 1929 , through the synonymy of Sydaphera renovata Iredale, 1929 . — Current status: the Japanese Sydaphera spengleriana (Deshayes, 1830) ( Qi 2004: 109, pl. 66, fig. B; Petit & Harasewych 2005: 18, 96; Okutani 2017: 1053, pl. 343, fig. 2).

striatulata , Cancellaria – Deshayes, 1835-DescrCoqFoss: 503, 42 [as C. “ striatula ”], pl. 79, figs 29-30;1837-DescrCoqFoss: 814, correction noted; 1843-HistNatAnim2: 428-429; 1843-HistNatAnim3: 653; 1864-DescrCoqFoss: 106. — Type locality: Mouchy-le-Châtel. — Type age: Eocene (Lutetian). Bonellita striatulata (Deshayes, 1835) ( Le Renard & Pacaud 1995: 121) . Listed by Petit & Harasewych (2005: 98). — Current status: Admetula striatulata (Deshayes, 1835) ( Pacaud 2008: 71) .

turriculatum, Triton – Deshayes, 1835b: 187-188, pl. 19, figs 58-60, non Deshayes, 1835-DescrCoqFoss [p. 608] (see below). — Type locality: Peloponnese Peninsula, Greece. — Type age: Recent. — Current status: synonym of Cumia intertexta (Helbling, 1779) [ Murex (Fusus) ] (MolluscaBase).

turriculatum, Triton – Deshayes, 1835-DescrCoqFoss: 608, 42, pl. 80, figs 7-12 [figs 10-12 not cited in text; 1837]: 813 [correction made], non Deshayes, 1835b (see above); 1865-DescrCoqFoss: 312. — Type localities: Thiverval-Grignon, Mantes & Mouchy-le-Châtel. — Type age: Eocene (Lutetian). Plesiotriton deshayesianus Beu & P. A. Maxwell, 1987 , replacement name. The latter is the type species (OD) of Colubratriton Pacaud, Ledon & Loubry, 2015 . Exact dates of both Deshayes’ 1835 works unknown, so either could have been replaced. — Current status: Colubratriton deshayesianus (Beu & P. A. Maxwell, 1987) ( Beu & Maxwell 1987: 25-26, pl. 23, figs a-e, h; Pacaud et al. 2015: 85-86, fig. 4, pl. 11, figs 1-5).

uniangulata , Cancellaria – Deshayes, 1830a-EncyMeth: 181-182; 1843-HistNatAnim2: 423;1843-HistNatAnim3: 651.— Type locality: Asti. — Type age: Pliocene (Piacenzian-Zanclean). — Current status: Tribia uniangulata (Deshayes, 1830) ( Petit & Harasewych 2005: 110; Brunetti et al. 2011: 126; Koskeridou et al. 2017: 270, fig. 2, 272).

●

asperula , Cancellaria “Deshayes” – Deshayes in Guérin-Méneville, 1835: pl. 17, fig. 3; Cancellaria asperella “Deshayes ”. Deshayes, in: 1844: 33, error C. asperella Lamarck, 1822 , correctly credited in Guérin-Méneville & Deshayes (1868).

elegans , Triton – Leymerie, 1841: 320, nomen nudum; 1842: 14, 31, pl. 17, fig. 13a-b, ex Deshayes ms, non Triton (Pusio) elegans J. E. Gray , in Griffith & Pidgeon, 1833. — Type locality: Courtaoult. — Type age: Cretaceous (Albian). Secondary homonym of Fusus elegans Lamarck, 1803 ; Fusus elegans J. de C. Sowerby, 1824 ; Fusus elegans T. Brown, 1827 ; and other species, when d’Orbigny placed Deshayes’ and other species into Fusus . Fusus subelegans d’Orbigny, 1850b (d’Orbigny 1850b: 133, no. 180, citing Deshayes’ species under his own treatment in the Paléontologie Française, and as “ non T. Brown, 1827”), replacement name. Listed in Snyder (2022: 77). Use of the replacement name Fusus subelegans d’Orbigny, 1850 , by many authors, makes the secondary homonym permanently invalid (ICZN Code Art. 59.3). — Current status: Paladmete subelegans (d’Orbigny, 1850) ( Kollmann 2005: 147) , which preoccupies Paladmete subelegans Stephenson, 1941 .