CLAVAGELLIDAE d’Orbigny, 1844

Coan, Eugene V., Pacaud, Jean-Michel & Kabat, Alan R., 2024, Gérard-Paul Deshayes (1796 - 1875) and his taxa, Geodiversitas 46 (16), pp. 553-930 : 797

Family CLAVAGELLIDAE d’Orbigny, 1844

Buccodus – Deshayes, 1839-TraitElem-1(2): 17, ex Guettard ms. Incorrect subsequent spelling of Bunodus Guettard, 1770 ; in the synonymy of Clavagella .

bacillaris , Clavagella – Deshayes, 1830a-EncyMeth: 239-240; 1833b: 231; 1835b: 84; 1835-HistNatAnim2: 24; 1839-HistNatAnim3: 511; 1839-TraitElem-1(2): 23, 24-25, 1, pl. 1, figs 4-10; 1840-RegAnim: pl. 119, fig. 3, 3a-e; 1843d: 5, pl. 3, figs 8, 8a-f; 1845- Algér: 16-17. — Type locality: Sicily (Italiy). — Type age: Pliocene. — Current status: Stirpulina bacillum [ Brocchi, 1814 ( Teredo )] bacillaris (Deshayes, 1830) ( Stallwood 1995: 87, 88; Coan & Kabat 2017: 108). Clavagella bacillaris (Deshayes, 1830) ( Zenetos et al. 2005: 5, as “1832”).

brongniarti , Clavagella – Deshayes, 1824a: 250, 257, pl. 15, fig. 1; 1824-DescrCoqFoss: 11, 1, pl. 1, figs 1-5; 1830a-EncyMeth: 240; 1828-DictClass4: 195; 1843d: 4-5, pl. 2, fig. 1;1857-DescrCoqFoss: 93 (with additional synonyms).— Type locality:Valmondois. — Type age: Eocene (Bartonian). — Current status: Clavagella (Clavagella) brongniarti Deshayes, 1824 ( Glibert & Van de Poel 1966a: 11; Le Renard & Pacaud 1995: 67).

caillati , Clavagella –Deshayes, 1856-DescrCoqFoss: 1, pl. 1, figs 1-4; 1857-DescrCoqFoss: 88. — Type locality: Thiverval-Grignon. — Type age: Eocene (Lutetian). Clavagella (Stirpulina) caillati Deshayes, 1856 ( Glibert & Van de Poel 1966a: 12; Le Renard & Pacaud 1995: 67). Stirpulina caillati (Deshayes, 1856) ( Stallwood 1995: 87, 88, as “1866”). — Current status: Clavagella (Clavagella) caillati Deshayes, 1856 ( Pacaud 2008: 85).

coronata , Clavagella – Deshayes, 1824-DescrCoqFoss: 8-9, 3, pl. 5, figs 15-16; 1830a-EncyMeth: 239; 1835-HistNatAnim2: 23-24; Deshayes in Guérin-Méneville 1835: pl. 33, fig. 6, 6a, 6b; 1839-HistNatAnim3: 511; 1839-TraitElem-1(2): 23-24, 2, pl. 1, fig. 11; 1843d: 4, pl. 2, figs 5-9; Deshayes in Guérin-Méneville 1844: 51-52; 1856-DescrCoqFoss: 1, pl. 1, figs 5-15; 1857: 89-90. — Type localities: Lizy-sur-Ourq & Pauliac near Bordeaux. — Type age: Eocene (Bartonian). — Type species (SD Stoliczka, 1871) of Stirpulina Stoliczka, 1870 . Stirpulina was conserved and senior synonym Tubolana Bivona Bernardi, 1832 , was supressed by ICZN Opinion 2325 (2013). (The type species of the suppressed Tubolana was T. digitata Bivona Bernardi, 1832 , now regarded as a synonym of Clavagella bacillaris Deshayes, 1830 .) Clavagella (Stirpulina) coronata Deshayes, 1824 ( Glibert & Van de Poel 1966a: 13; Glibert 1980: 47; Le Renard & Pacaud 1995: 67). — Current status: Stirpulina coronata (Deshayes, 1824) ( Stallwood 1995: 88; Nevesskaja et al. 2013: 284, fig. 103-9).

depressa , Saxicava – Deshayes, 1824a: 254-255, 258, pl. 15, fig. 10; 1824-DescrCoqFoss: 66, 5, pl. 9, figs 20-21; 1832-EncyMeth: 928; 1835-HistNatAnim2: 155; 1839-HistNatAnim3: 553; 1857-DescrCoqFoss: 93 (as synonym of Clavagella brongniarti Deshayes, 1824 ). — Type locality: Valmondois. — Type age: Eocene (Bartonian). — Current status: synonym of Clavagella brongniarti Deshayes, 1824 .

desmoulinsi , Clavagella – Deshayes, 1857-DescrCoqFoss: 90; Guérin-Méneville & Deshayes 1868: 66, pl. 32, fig. 6, 6a, 6b. The latter treatment were the same figures treated in Guérin-Méneville (1835, 1844), also as C. coronata (see above). Based on figures in Des Moulins (1829: 239-246, pl. 13, figs “1-4” [actually 1-5]) of what Des Moulins figured as Clavagella coronata Deshayes, 1824 . — Type localities: Lafite (Pauillac) & Saint-Estèphe. — Type age: Eocene (Bartonian). In subsequent literature, this has been mistakenly referred to as just C. “ moulinsi ” and/or given a second terminal “i”, as “ desmoulinsii ”. Clavagella desmoulinsi Deshayes, 1857 ( Dollfus & Dautzenberg 1902: 56). — Current status: Brechites desmoulinsi (Deshayes, 1857) (J.-F. Lesport, pers. comm., December 19, 2022).

lagenalis , Clavagella – Deshayes, 1856-DescrCoqFoss: 2, pl. 2, figs 4-6; 1857-DescrCoqFoss: 92-93, as C. lagenula . — Type localities: Ver-sur-Launette &Le Guépelle. — Type age: Eocene (Bartonian). — Current status: Clavagella (Clavagella) lagenalis Deshayes, 1856 ( Glibert & Van de Poel 1966a: 12; Glibert 1980: 47-48; Le Renard & Pacaud 1995: 67).

lamarckii , Clavagella – Deshayes, 1856-DescrCoqFoss: 2, pl. 2, figs 7-10 [fig. 10 not mentioned in text]; 1857-DescrCoqFoss: 93-94; 1865-DescrCoqFoss: 665 [this correction noted]. — Type locality: Les Bouleaux. — Type age: Eocene (Lutetian). — Current status: Clavagella (Clavagella) lamarckii Deshayes, 1856 ( Le Renard & Pacaud 1995: 67; Pacaud 2008: 85).

margaritacea , Saxicava – Deshayes, 1824a: 258, 254, pl. 15, fig. 9; 1824-DescrCoqFoss: 65, 5, pl. 9, figs 22-24; 1829-DictClass15: 24; 1832-EncyMeth: 928; 1835-HistNatAnim2: 155; 1839-HistNatAnim3: 553; 1857-DescrCoqFoss: 93 (as synonym of Clavagella brongniarti Deshayes, 1824 ). — Type locality: Valmondois. — Type age: Eocene (Bartonian). — Current status: synonym of Clavagella brongniarti Deshayes, 1824 .

primigenia , Clavagella – Deshayes, 1857-DescrCoqFoss: 94-95, 15, pl. 15, figs 1-2. — Type locality: Gannes. — Type age: Eocene (Lutetian). — Current status: Clavagella (Clavagella) primigenia Deshayes, 1857 ( Le Renard & Pacaud 1995: 67).

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lata , Clavagella – Deshayes, 1839-TraitElem-1(2): 25-26, 2, pl. 1, figs 12-14. Sherborn (1927: 3455) recorded this as being by Deshayes, separately from the Indo-Pacific Clavagella lata Broderip, 1834 , although Deshayes noted Broderip’s publication, perhaps because Deshayes was miscredited with the species on the plate explanation. — Current status: Bryopa lata (Broderip, 1834) [ Clavagella ].

philippiana , Clavagella – G. B. Sowerby II, 1872b: pl. 1, fig. 3, ex Deshayes ms. — Type locality: Mediterranean. — Type age: Recent. Petit (2009: 151). — Current status: synonym of Bryopa aperta (G. B. Sowerby I, 1823) [ Clavagella ].