Liothorax innexus ( Say, 1835 )

Dellacasa, Marco, Gordon, Robert D. & Dellacasa, Giovanni, 2007, Systematic review to the genus Liothorax (Coleoptera: Scarabaeoidea: Scarabaeidae: Aphodiinae), Zootaxa 1407, pp. 23-42 : 28-29

Liothorax innexus ( Say, 1835)

( Figs. 13–17 )

Aphodius innexus: Say 1835: 177 , Harold 1871: 270, Bates 1887: 85, Blackwelder 1944: 212, Dellacasa et al. 1998: 155 Aphodius flavocinctus: Harold 1860: 614 , Harold 1871: 270 (as a synonym of A. innexus ) Aphodius circumcinctus: Germar in Harold 1871: 270 (nomen nudum)

Aphodius (Nialus) innexus: Schmidt 1913: 168 ; Dellacasa 1988: 144

Liothorax innexus: Dellacasa et al., 2002: 166 (new combination)

Type locality: Mexico (for Liothorax innexus ).

Type depository: Type series lost (for Liothorax innexus ) (see Mawdsley 1993).

Redescription: Length 5.0–6.0 mm; body subelongate, parallel, convex, shiny glabrous. Color piceous; front angles of pronotum, humeral angle, sides and apex of elytra dirty yellow. Head anteriomedially gibbose, anteriorly with transverse weak carina, nearly smooth at center, with coarse punctures only toward clypeal margin. Clypeus sinuate medially, rounded laterally, finely bordered, border weakly upturned, glabrous; genae broadly rounded, with elongate setae, not protruding beyond eyes; frontal suture weakly gibbous. Pronotum transverse, with dual sparse punctures, nearly smooth on disc; lateral margins thinly bordered, bor- der with elongate setae; base weakly bisinuate, not bordered. Scutellum with curved sides, smooth. Elytra subparallel-sided, distinctly striate; striae strongly crenulate; interstriae flat, with very fine and sparse punctures, sometimes punctures almost imperceptible. Hind tibiae with superior apical spur shorter than first tarsomere; first tarsomere nearly as long as following three combined. Male head with anterior carina more distinct than in female; pronotum almost parallel-sided. Female with pronotum weakly narrowed forward.

Distribution: Mexico (Aguascalientes, Distrito Federal, Guanajuato, Guerrero, Hidalgo, México, Michoacán, Oaxaca, Puebla).

Bionomy: This is a coprophagous species, which is found in summer and autumn. Other bionomical details are unknown.

Systematic remarks: This species has been recorded from California, U.S.A. ( Harold 1860: 614) but this record was based on misidentified specimens.

Figures

FIGURES 13 – 17. Liothorax innexus (Say, 1835) (Lago Zumpango, env. San Juan Zitlaltepec, E. do México, Mexico): 13. habitus of male; 14 – 15. aedeagus, dorsal and lateral view; 16. epipharynx; 17. habitus.