Quateiella quatei Cook

Amorim, Dalton de Souza & Brown, Brian V., 2020, Urban Scatopsidae (Diptera) of Los Angeles, California, United States, Insect Systematics and Diversity 4 (1), No. 1, pp. 1-41 : 25-35

Quateiella quatei Cook

( Figs. 9, 10 , 23 , 24, and 67–74)

Quateiella quatei (Cook) : 1956a: 9 ( Rhexoza ). Type locality: United States, California, Davis. Type ♂, (UCD).

Ref.– Cook, 1956a: figs. 4C (male terminalia), 4E (female terminalia), 4G (male sternite 7 and tergite 7), 4H (wing); Cook, 1975: figs 1 (thorax pleura), 2 (male segment 7, lateral view), 5 (male terminalia); Cook, 1967 (Nearctic catalog).

Diagnosis. Male tergite 9 laterally compressed, with a conspicuous beak-like ventral process. Female tergite 7 posterior border with gentle lateral emargination; tergite 8 medially divided.

Material Examined. 1 ♀, United States, California, Los Angeles Co., Los Angeles, University Park , 34.034°N 118.281°W, BioSCAN site 3, 3–10 December 2014, Malaise trap, 63 m, Coll. Peggy Hentschke, BioSCAN sample 16969 ( LACM) ; 1 ♀, same data, but BioSCAN sample 16251 ( LACM) ; 1 ♂, 1 ♀, same data, but BioSCAN sample 16072 ( LACM) ; 1 ♂, 2 ♀♀, same data, but 30 April–7 May 2014, BioSCAN sample 16068 ( LACM) ; 1 ♂, same data, but 29 October–5 November 2014, BioSCAN sample 16774 ( LACM) ; 1 ♂, same data, but BioSCAN sample 16603 ( LACM) ; 1 ♂, United States, California, Los Angeles Co., Los Angeles, Los Feliz , 34.118°N 118.284°W, BioSCAN site 4, 29 March–4 April 2014, Malaise trap, 220 m, Coll. Mark and Jane Pisano, BioSCAN sample 15950 ( LACM) ; 5 ♂♂, 4 ♀♀, United States, California, Los Angeles Co., Los Angeles, Silverlake , 34.093°N 118.274°W, BioSCAN site 5, 28 December 2013 – 4 January 2014, Malaise trap, 166.1 m, Coll. Walter Renwick, BioSCAN sample 15419 ( LACM) ; 1 ♀, same data, but BioSCAN sample 16268 ( LACM) ; 1 ♀, same data, but 3–10 May 2014, BioSCAN sample 16109 ( LACM) ; 2 ♂♂, 1 ♀, same data, but 2–9 November 2013, BioSCAN sample 15318 ( LACM) (1♂, 1♀ on slide) ; 5 ♂♂, 3 ♀♀, same data, but 1–8 November 2014, BioSCAN sample 16796 ( LACM) ; 1 ♂, 4 ♀♀, same data, but 28 September–5 October 2013, BioSCAN sample 15226 ( LACM) ; 4 ♀♀, United States, California, Los Angeles Co., Los Angeles, Los Feliz , 34.116°N 118.279°W, BioSCAN site 6, 29 March–14 April 2014, Malaise trap, 199 m, Coll. Jeff, Aaron and Jacob Koch, BioSCAN sample 15939 ( LACM) ; 2 ♂♂, 1 ♀, same data, but 1–8 February 2014, BioSCAN sample 15572 ( LACM) ; 1 ♀, same data, but 1–5 January 2014, BioSCAN sample 15568 ( LACM) ; 3 ♀♀, same data, but BioSCAN sample 16107 ( LACM) ; 1 ♀, same data, but 1–9 March 2014, BioSCAN sample 15746 ( LACM) ; 3 ♀♀, same data, but 4–10 May 2014, BioSCAN sample 16104 ( LACM) ; 1 ♀, same data, but 2–9 November 2013, BioSCAN sample 15306 ( LACM) ; 2 ♀♀, same data, but BioSCAN sample 16644 ( LACM) ; 1 ♀, same data, but 15–20 September 2013, BioSCAN sample 15207 ( LACM) ; 2 ♀♀, same data, but BioSCAN sample 16463 ( LACM) ; 2 ♂♂, United States, California, Los Angeles Co., Los Angeles, Silverlake , 34.102°N 118.257°W, BioSCAN site 7, 29 March–5 April 2014, Malaise trap, 158.9 m, Coll. Joe Hogg, BioSCAN sample 15834 ( LACM) ; 1 ♀, same data, but 28 December 2013 – 4 January 2014, BioSCAN sample 15533 ( LACM) ; 1 ♀, same data, but 1–8 July 2016, BioSCAN sample 18324 ( LACM) ; 1 ♀, same data, but BioSCAN sample 16189 ( LACM) ; 1 ♂, 1 ♀, same data, but BioSCAN sample 18248 ( LACM) ; 1 ♀, same data, but 2–9 November 2013, BioSCAN sample 15281 ( LACM) ; 1 ♀, United States, California, Los Angeles Co., Los Angeles, Elysian Valley , 34.103°N 118.243°W, BioSCAN site 9, 14–21 September 2013, Malaise trap, 106.7 m, Coll. John Rodriguez, BioSCAN sample 15198 ( LACM) ; 1 ♀, United States, California, Los Angeles Co., Los Angeles, Korea Town , 34.072°N 118.291°W, BioSCAN site 11, 5–12 April 2014, Malaise trap, 85.6 m, Coll. Peter Ralph, BioSCAN sample 16056 ( LACM) ; 1 ♀, same data, but 2–9 March 2014, BioSCAN sample 15796 ( LACM) ; 1 ♂, same data, but 4–13 May 2014, BioSCAN sample 16060 ( LACM) ; 2 ♀♀, United States, California, Los Angeles Co., Los Angeles, Mid-City , 34.047°N 118.334°W, BioSCAN site 12, Malaise trap, 52.6 m, Coll. Sharon Oxborough, BioSCAN sample 16323 ( LACM) ; 1 ♀, same data, but 3–10 May 2014, BioSCAN sample 15948 ( LACM) ; 1 ♂, 1 ♀, same data, but 2–9 November 2013, BioSCAN sample 15293 ( LACM) ; 2 ♂♂, same data, but 14–21 September 2013, BioSCAN sample 15202 ( LACM) ; 3 ♂♂, United States, California, Los Angeles Co., Glendale , 34.149°N 118.218°W, BioSCAN site 13, Malaise trap, 245.4 m, Coll. Karen Hoffman, BioSCAN sample 16207 ( LACM) ; 1 ♂, same data, but 2–9 November 2013, BioSCAN sample 15352 ( LACM) ; 1 ♀, same data, but 29 November–6 December 2014, BioSCAN sample 16893 ( LACM) ; 1 ♀, same data, but 28 September–5 October 2013, BioSCAN sample 15245 ( LACM) ; 3 ♀♀, United States, California, Los Angeles Co., Los Angeles, Elysian Park , 34.078°N 118.234°W, BioSCAN site 14, 29 March–5 April 2014, Malaise trap, 149 m, Coll. Eric and Victoria Harding, BioSCAN sample 15828 ( LACM) ; 1 ♂, 1 ♀, same data, but 28 December 2013 – 4 January 2014, BioSCAN sample 15476 ( LACM) ; 2 ♀♀, same data, but BioSCAN sample 16347 ( LACM) ; 1 ♂, 5 ♀♀, same data, but BioSCAN sample 16166 ( LACM) ; 5 ♂♂, 3 ♀♀, same data, but 1–8 March 2014, BioSCAN sample 15824 ( LACM) ; 1 ♂, 3 ♀♀, same data, but 3–10 May 2014, BioSCAN sample 15992 (on slide) ; 15 ♂♂, 4 ♀♀, same data, but 2–9 November 2013, BioSCAN sample 15345 ( LACM) ; 3 ♂♂, 4 ♀♀, same data, but 1–8 November 2014, BioSCAN sample 16702 ( LACM) ; 10 ♂♂, 17 ♀♀, same data, but 28 September–5 October 2013, BioSCAN sample 15238 ( LACM) ; 1 ♂, United States, California, Los Angeles Co., Los Angeles, Gardena , 33.876°N 118.288°W, BioSCAN site 15, 29 March–5 April 2014, Malaise trap, 9 m, Coll. Betty Defibaugh, BioSCAN sample 15903 ( LACM) ; 10 ♀♀, United States, California, Los Angeles Co., Los Angeles, Hollywood , 34.095°N 118.334°W, BioSCAN site 16, 2–9 April 2014, Malaise trap, 100 m, Coll. Tony Hein, BioSCAN sample 15980 ( LACM) ; 3 ♀♀, same data, but BioSCAN sample 16345 ( LACM) ; 1 ♀, same data, but 29 January–5 February 2014, BioSCAN sample 15611 ( LACM) ; 4 ♀♀, same data, but 1–18 January 2014, BioSCAN sample 15608 ( LACM) ; 6 ♀♀, same data, but BioSCAN sample 16341 ( LACM) ; 3 ♀♀, same data, but BioSCAN sample 16137 ( LACM) ; 2 ♀♀, same data, but 30 April–7 May 2014, BioSCAN sample 15984 ( LACM) (2♀♀ on slide) ; 2 ♀♀, same data, but BioSCAN sample 16686 ( LACM) ; 6 ♀♀, same data, but BioSCAN sample 16519 ( LACM) ; 1 ♂, United States, California, Los Angeles Co., Glendale , 34.159°N 118.247°W, BioSCAN site 17, Malaise trap, 190 m, Coll. Celeste Armstrong, BioSCAN sample 16196 ( LACM) ; 1 ♀, United States, California, Los Angeles Co., Los Angeles, Larchmont , 34.077°N 118.32°W, BioSCAN site 18, 29 March–5 April 2014, Malaise trap, 76 m, Coll. K.T. Wiegman, BioSCAN sample 15968 ( LACM) ; 1 ♂, same data, but 2–9 November 2013, BioSCAN sample 15322 ( LACM) ; 1 ♂, 1 ♀, same data, but 1–8 November 2014, BioSCAN sample 16843 ( LACM) ; 2 ♂♂, same data, but BioSCAN sample 16669 ( LACM) ; 1 ♂, same data, but BioSCAN sample 16501 ( LACM) ; 3 ♂♂, 1 ♀, United States, California, Los Angeles Co., Los Angeles, Carthay , 34.059°N 118.369°W, BioSCAN site 19, 1–8 April 2014, Malaise trap, 41.1 m, Coll. Teresa Dahl, BioSCAN sample 15812 ( LACM) ; 2 ♀♀, same data, but BioSCAN sample 16338 ( LACM) ; 3 ♀♀, same data, but 2–9 December 2014, BioSCAN sample 16876 ( LACM) ; 3 ♂♂, 12 ♀♀, same data, but 28 January–4 February 2014, BioSCAN sample 15593 ( LACM) ; 1 ♂, 1 ♀, same data, but BioSCAN sample 16334 ( LACM) ; 1 ♂, 1 ♀, same data, but BioSCAN sample 16147 ( LACM) ; 1 ♂, 3 ♀♀, same data, but BioSCAN sample 15977 ( LACM) ; 1 ♂, 4 ♀♀, same data, but 29 October–5 November 2013, BioSCAN sample 15325 ( LACM) ; 1 ♂, 5 ♀♀, same data, but 28 October–4 November 2014, BioSCAN sample 16871 ( LACM) ; 1 ♂, 1 ♀, same data, but BioSCAN sample 16868 ( LACM) ; 1 ♀, same data, but BioSCAN sample 16513 ( LACM) ; 1 ♂, 3 ♀♀, United States, California, Los Angeles Co., Burbank , 34.17°N 118.308°W, BioSCAN site 21, 1–8 March 2014, Malaise trap, 158 m, Coll. Chris Cianci, BioSCAN sample 15668 ( LACM) ; 1 ♂, same data, but 3–10 May 2014, BioSCAN sample 16025 ( LACM) ; 5 ♀♀, United States, California, Los Angeles Co., Los Angeles, Echo Park , 34.074°N 118.264°W, BioSCAN site 22, 2–8 April 2014, Malaise trap, 133.5 m, Coll. Erin Johnson, BioSCAN sample 15932 ( LACM) ; 1 ♀, same data, but BioSCAN sample 16302 ( LACM) ; 1 ♀, same data, but 2–9 December 2014, BioSCAN sample 16824 ( LACM) ; 3 ♀♀, same data, but 28 January–4 February 2014, BioSCAN sample 15577 ( LACM) ; 1 ♀, same data, but BioSCAN sample 16298 ( LACM) ; 1 ♂, 12 ♀♀, same data, but 29 April 2014 – 6 May 2014, BioSCAN sample 15936 ( LACM) ; 2 ♀♀, same data, but 28 October–4 November 2014, BioSCAN sample 16819 ( LACM) ; 2 ♀♀, same data, but BioSCAN sample 16650 ( LACM) ; 2 ♀♀, same data, but BioSCAN sample 16477 ( LACM) ; 6 ♀♀, United States, California, Los Angeles Co., Los Angeles, Atwater Village , 34.114°N 118.251°W, BioSCAN site 23, 2–10 April 2014, Malaise trap, 119 m, Coll. Sidney Higgins, BioSCAN sample 15858 ( LACM) ; 1 ♀, same data, but 2–8 January 2014, BioSCAN sample 15540 ( LACM) ; 1 ♀, same data, but 26 February–5 March 2014, BioSCAN sample 15663 ( LACM) ; 1 ♀, same data, but 29 October–7 November 2014, BioSCAN sample 16707 ( LACM) ; 4 ♀♀, United States, California, Los Angeles Co., Los Angeles, Eagle Rock , 34.136°N 118.194°W, BioSCAN site 24, 29 March–5 April 2014, Malaise trap, 215 m, Coll. Jim Hogue, BioSCAN sample 15846 ( LACM) ; 1 ♂, same data, but BioSCAN sample 16406 ( LACM) ; 1 ♀, same data, but 28 December 2013 – 4 January 2014, BioSCAN sample 15485 ( LACM) ; 1 ♀, same data, but BioSCAN sample 16187 ( LACM) ; 1 ♀, same data, but 1–8 March 2014, BioSCAN sample 15842 ( LACM) ; 1 ♀, United States, California, Los Angeles Co., Los Angeles, Mid–Wilshire , 34.058°N 118.328°W, BioSCAN site 26, 28 December 2013 – 4 January 2014, Malaise trap, 58.7 m, Coll. Patty Lombard, BioSCAN sample 15465 ( LACM) ; 1 ♀, same data, but BioSCAN sample 16663 ( LACM) ; 1 ♀, United States, California, Los Angeles Co., Los Angeles, Glassell Park , 34.111°N 118.230°W, BioSCAN site 27, Malaise trap, 186.8 m, Coll. Glen Creason, BioSCAN sample 16067 ( LACM) ; 1 ♀, United States, California, Los Angeles Co., Los Angeles, Eagle Rock , 34.129°N 118.215°W, BioSCAN site 28, 28 December 2013 – 4 January 2014, Malaise trap, 160.7 m, Coll. Eric Keller, BioSCAN sample 15480 ( LACM) ; 1 ♀, same data, but BioSCAN sample 16245 ( LACM) ; 1 ♀, same data, but BioSCAN sample 16241 ( LACM) ; 1 ♂, same data, but 1–8 March 2014, BioSCAN sample 15714 ( LACM) ; 3 ♀♀, same data, but 3–10 May 2014, BioSCAN sample 16044 ( LACM) ; 5 ♀♀, same data, but 1–8 November 2014, BioSCAN sample 16755 ( LACM) ; 3 ♀♀, same data, but BioSCAN sample 16597 ( LACM) ; 2 ♂♂, 11 ♀♀, United States, California, Los Angeles Co., Los Angeles, Highland Park , 34.123°N 118.193°W, BioSCAN site 29, 29 March–5 April 2014, Malaise trap, 205.6 m, Coll. Natalie Brejcha, BioSCAN sample 15865 ( LACM) ; 1 ♂, 2 ♀♀, same data, but BioSCAN sample 16412 ( LACM) ; 4 ♂♂, 5 ♀♀, same data, but 21–28 December 2013, BioSCAN sample 15488 (4♂♂, 2♀♀ on slide) ( LACM) ; 1 ♂, 5 ♀♀, same data, but 28 December 2013 – 4 January 2014, BioSCAN sample 15489 ( LACM) ; 1 ♂, 1 ♀, same data, but BioSCAN sample 16233 ( LACM) ; 6 ♂♂, 3 ♀♀, same data, but BioSCAN sample 16229 ( LACM) ; 1 ♂, 5 ♀♀, same data, but 1–8 March 2014, BioSCAN sample 15707 ( LACM) ; 4 ♂♂, 5 ♀♀, same data, but 3–10 May 2014, BioSCAN sample 16032 (2♂♂, 4♀♀ on slide) ( LACM) ; 3 ♂♂, 5 ♀♀, same data, but 25 October–8 November 2014, BioSCAN sample 16786 ( LACM) ; 5 ♂♂, 3 ♀♀, United States, California, Los Angeles Co., Los Angeles, Highland Park , 34.125°N 118.189°W, BioSCAN site 30, 31 March–8 April 2014, Malaise trap, 227.8 m, Coll. Candace Franco, BioSCAN sample 15887 ( LACM) ; 1 ♂, same data, but BioSCAN sample 16423 ( LACM) ; 1 ♂, same data, but 28 December 2013 – 4 January 2014, BioSCAN sample 15527 ( LACM) ; 15 ♂♂, 4 ♀♀, same data, but BioSCAN sample 16239 ( LACM) ; 22 ♂♂, 2 ♀♀, same data, but BioSCAN sample 16235 ( LACM) ; 13 ♂♂, 5 ♀♀, same data, but BioSCAN sample 15767 ( LACM) ; 15 ♂♂, 1 ♀, same data, but 3–12 May 2014, BioSCAN sample 16038 ( LACM) ; 16 ♂♂, 1 ♀, same data, but 1–8 November 2014, BioSCAN sample 16762 ( LACM) ; 2 ♂♂, same data, but 29 November–6 December 2014, BioSCAN sample 16955 ( LACM) ; 2 ♂♂, 4 ♀♀, same data, but BioSCAN sample 16595 ( LACM) ; 2 ♂♂, same data, but BioSCAN sample 16591 ( LACM) ; 3 ♀♀, United States, California, Los Angeles Co., Los Angeles , 34.103°N – 118.216°W, BioSCAN site 31, 29 March–5 April 2014, Malaise trap, 74 m, Coll. Julian Donahue, BioSCAN sample 15840 ( LACM) ; 1 ♂, 2 ♀♀, same data, but BioSCAN sample 16202 ( LACM) ; 3 ♂♂, 6 ♀♀, same data, but 3–10 May 2014, BioSCAN sample 16009 (1 ♂, on slide) ( LACM) ; 2 ♀♀, same data, but 1–8 November 2014, BioSCAN sample 16721 ( LACM) ; 1 ♂, United States, California, Los Angeles Co., Santa Monica. 34.021°N 118.489°W, BioSCAN site 32, 1–8 August 2016, Malaise trap, 53.7 m, Coll. Brian Palatucci, BioSCAN sample 18404 ( LACM) ; 1 ♀, United States, California, Los Angeles Co., Los Angeles, Pasadena , 34.163°N 118.118°W, BioSCAN site 33, 1–8 August 2016, Malaise trap, 280 m, Coll. Leslie Harris, BioSCAN sample 18235 ( LACM) ; 2 ♀♀, same data, but 1–8 June 2016, BioSCAN sample 18325 ( LACM) ; 1 ♀, United States, California, Los Angeles Co., Los Angeles, Rosemead , 34.05°N 118.108°W, BioSCAN site 34, 1–8 April 2016, Malaise trap, 135 m, Coll. Deniece Lascano, BioSCAN sample 18230 ( LACM) ; 16 ♂♂, 16 ♀♀, United States, California, Los Angeles Co., Monrovia. 34.158°N 117.999°W, BioSCAN site 35, 1–8 April 2016, Malaise trap, 214 m, Coll. Brian Brown, BioSCAN sample 18197 ( LACM) ; 5 ♂♂, 4 ♀♀, same data, but 4–11 August 2016, BioSCAN sample 18370 ( LACM) ; 1 ♀, same data, but BioSCAN sample 18136 ( LACM) ; 6 ♂♂, 4 ♀♀, same data, but 1–15 June 2016, BioSCAN sample 18277 ( LACM) ; 1 ♂, 4 ♀♀, same data, but 1–8 March 2016, BioSCAN sample 18155 ( LACM) ; 3 ♂♂, 9 ♀♀, same data, but 1–8 May 2016, BioSCAN sample 18212 ( LACM) ; 3 ♂♂, 1 ♀, same data, but 1–8 October 2016, BioSCAN sample 18422 ( LACM) ; 1 ♂, 1 ♀, same data, but 1–8 September 2016, BioSCAN sample 18375 ( LACM) ; 2 ♀♀, United States, California, Los Angeles Co., San Dimas , 34.086°N 117.822°W, BioSCAN site 36, 1–8 April 2016, Malaise trap, 298 m, Coll. Carla Zorko, BioSCAN sample 18245 ( LACM) ; 1 ♂, same data, but 1–8 August 2016, BioSCAN sample 18401 ( LACM) ; 1 ♂, same data, but 1–8 July 2016, BioSCAN sample 18342 ( LACM) ; 1 ♂, 6 ♀♀, same data, but BioSCAN sample 18246 ( LACM) ; 10 ♀♀, United States, California, Los Angeles Co., Claremont , 34.09°N 117.711°W, BioSCAN site 37, 1–8 April 2016, Malaise trap, 341 m, Coll. Kathryn Turner, BioSCAN sample 18249 ( LACM) ; 1 ♀, same data, but 1–8 February 2016, BioSCAN sample 18163 ( LACM) ; 11 ♀♀, same data, but 1–8 June 2016, BioSCAN sample 18337 ( LACM) ; 5 ♂♂, 2 ♀♀, same data, but 1–8 March 2016, BioSCAN sample 18164 ( LACM) ; 3 ♀♀, same data, but BioSCAN sample 18250 ( LACM) ; 2 ♀♀, United States, California, San Bernardino Co., Rancho Cucamonga , 34.139°N 117.521°W, BioSCAN site 38, 1–8 August 2016, Malaise trap, 451 m, Coll. Lucila Tagliani, BioSCAN sample 18407 ( LACM) ; 2 ♀♀, same data, but 1–8 July 2016, BioSCAN sample 18299 ( LACM) ; 1 ♀, same data, but 1–8 June 2016, BioSCAN sample 18298 ( LACM) ; 1 ♀, same data, but BioSCAN sample 18226 ( LACM) ; 1 ♀, same data, but BioSCAN sample 18409 ( LACM) ; 1 ♂, United States, California, San Bernardino Co., Grand Terrace , 34.024°N 117.315°W, BioSCAN site 39, 1–8 February 2016, Malaise trap, 322 m, Coll. Dave Goodward, BioSCAN sample 18182 ( LACM) ; 1 ♀, United States, California, Los Angeles Co., El Segundo. 33.925°N 118.425°W, BioSCAN site 40, 2–16 March 2016, Malaise trap, 35 m, Coll. Jack Fulmer, BioSCAN sample 18153 ( LACM) ; 1 ♀, United States, California, Los Angeles Co., Gardena. 33.885°N 118.287°W, BioSCAN site 41, Malaise trap, 1–8 August 2016, Malaise trap, 14 m, Coll. Ron Matsumoto, BioSCAN sample 18386 ( LACM) ; 1 ♂, same data, but 1–8 December 2015, BioSCAN sample 18103 ( LACM) ; 1 ♀, same data, but 1–8 January 2016, BioSCAN sample 18104 ( LACM) ; 2 ♂♂, same data, but 1–8 February 2016, BioSCAN sample 18148 ( LACM) ; 2 ♂♂, 4 ♀♀, same data, but 1–8 April 2016, BioSCAN sample 18237 ( LACM) ; 1 ♀, same data, but 1–8 May 2016, BioSCAN sample 18238 ( LACM) ; 1 ♀, United States, California, Los Angeles Co., La Mirada , 33.914°N 118.005°W, BioSCAN site 43, 1–8 December 2015, Malaise trap, 63 m, Coll. Carmen Gendusa, BioSCAN sample 18131 ( LACM) ; 1 ♂, 2 ♀♀, same data, but 1–8 June 2016, BioSCAN sample 18321 ( LACM) ; 1 ♀, same data, but 1–8 March 2016, BioSCAN sample 18147 ( LACM) ; 3 ♀♀, same data, but 1–8 April 2016, BioSCAN sample 18265 ( LACM) ; 2 ♂♂, 4 ♀♀, same data, but 1–8 May 2016, BioSCAN sample 18266 ( LACM) ; 2 ♀♀, same data, but 1–8 October 2016, BioSCAN sample 18385 ( LACM) ; 3 ♂♂, 3 ♀♀, same data, but 1–8 September 2016, BioSCAN sample 18384 ( LACM) ; 1 ♂, United States, California, Los Angeles Co., Walnut. 33.998°N 117.842°W, BioSCAN site 44, Malaise trap, 189 m, Coll. Emilia Park, BioSCAN sample 18252 ( LACM) ; 1 ♀, United States, California, San Bernardino Co., Chino Hills , 34.003°N 117.734°W, BioSCAN site 45, 1–8 August 2016, Malaise trap, 225 m, Coll. Joe Williams, BioSCAN sample 18448 ( LACM) ; 1 ♀, same data, but 1–8 June 2016, BioSCAN sample 18339 ( LACM) ; 2 ♀♀, United States, California, Riverside Co., Norco. 33.912°N 117.57°W, BioSCAN site 46, 1–8 April 2016, Malaise trap, 180 m, Coll. Sara Taylor, BioSCAN sample 18227 ( LACM) ; 2 ♀♀, United States, California, Riverside Co., Norco. 33.912°N 117.57°W, BioSCAN site 46, 1–8 March 2016, Malaise trap, 180 m, Coll. Sara Taylor, BioSCAN sample 18177 ( LACM) ; 1 ♀, same data, but BioSCAN sample 18228 ( LACM) ; 8 ♂♂, 4 ♀♀, United States, California, Riverside Co., Riverside , 33.984°N 117.313°W, BioSCAN site 47, 1–8 April 2016, Malaise trap, 375 m, Coll. Justin Betz, BioSCAN sample 18223 ( LACM) ; 11 ♂♂, 7 ♀♀, same data, but 1–8 June 2016, BioSCAN sample 18279 ( LACM) ; 3 ♂♂, 1 ♀, same data, but 1–8 March 2016, BioSCAN sample 18179 ( LACM) ; 1 ♂, United States, same data, but 1–8 October 2016, BioSCAN sample 18412 ( LACM) (1 ♂, 3 ♀♀ BioSCAN sample 15992 at MZUSP, all other specimens at LACM) .

Distribution. United States, southern California.

Remarks. The type locality of Q. quatei is Davis, California. Additional specimens are known from Anaheim, Riverside, Rialto, and Olive, in the area around Los Angeles, as well as Coachella, more to the east in southern California ( Cook 1956a), and from San Bernardino ( Cook 1975). Cook (1956a) mentioned specimens collected with citronella bait traps and Cook (1975) refers to specimens reared from Echinocactus polycephalus . This cactus species is widespread in the Mohave and Sonoran desert in Arizona (Yuma and Mojave Co.), California (Inyo, San Bernardino Valley, Riverside, and Imperial Co.), and Nevada (Lincoln and Clark Co.), and in the northwestern Sonora desert in Mexico ( Chamberland 1997). Echinocactus polycephalus was not part of the original flora of the area of Los Angeles and is not particularly common in gardens in the urban area.

A total of 292 males and 435 females were collected in our traps. The species was collected in the largest number of BioSCAN sites— 42 of our 47 sampling sites, with 196 collecting events. The most abundant sites for Q. quatei were sites 14, 29, 30 (Phase I), and 35 (Phase II). Mid-spring is clearly the season with the highest abundance of Q. quatei , while a second, autumn peak is clear in Phase Inot detected in Phase II. It is possible that Q. quatei can also develop in other kinds of rotting cactuses. The BOLDSystems has no records for the genus Quateiella .

Figures

Figs. 7–10. Habitus. 7. Coboldia fuscipes (Meigen), male (black arrow indicates the diagnostic tergite 8 spatulate projection). 8. Coboldia fuscipes (Meigen), female. 9. Quateiella quatei (Cook), male. 10. Quateiella quatei (Cook), female.

Figs. 21–23. Abdomen. 21. Swammerdamella marginata Cook, male. 22. Swammerdamella marginata Cook, female. 23. Quateiella quatei (Cook), male. 24. Quateiella quatei (Cook), female.