Goodyera ovatilabia Schltr.

MONRO, A. K., SANTAMARÍA-AGUILAR, D., GONZÁLEZ, F., CHACÓN, O., SOLANO, D., RODRÍGUEZ, A., ZAMORA, N., FEDELE, E. & CORREA, M., 2017, A first checklist to the vascular plants of La Amistad International Park (PILA), Costa Rica-Panama, Phytotaxa 322 (1), pp. 448-450 : 448-450

Goodyera ovatilabia Schltr. Lepanthes erinacea Rchb.f.

Terrestrial herbs. Distribution in PILA 2500–3000 m, low Epiphytic herbs. Distribution in PILA 1000–2000 m, elevation oak forest, high elevation oak forest. Global mixed forest in transition to low elevation forest, mixed distribution: Costa Rica, Panama. Records: G. Davidse forest in transition to cloud forest. Global distribution: 23963 (CR, MO), G. Davidse 25733 (MO). Endemic to Costa Rica. Records: L.D. Gómez 23156 (CR, MO).

Govenia liliacea (La Llave & L ex.) Lindl.

Terrestrial herbs. Distribution in PILA 1400–1400 Lepanthes excedens Ames & Correll

m, mixed forest in transition to cloud forest. Global Epiphyticherbs.Distribution in PILA1300– 1300m,mixed distribution: Mexico to Costa Rica. Records: D. Solano forest in transition to cloud forest. Global distribution: 3190 (INB). Endemic to Panama. Records: Luer 9314 (SEL).

Habenaria distans Griseb. Lepanthes fimbriata Ames. Terrestrial herbs. Distribution in PILA 200–2400 m, low Epiphytic herbs.Distribution in PILA 1300–1600m,mixed elevation mixed forest, mixed forest in transition to low forest in transition to cloud forest. Global distribution: elevation forest, mixed forest in transition to cloud forest, Costa Rica, Panama. Records: Luer 9279 (SEL). cloud forest, low elevation oak forest. Global distribution: USA to Argentina, Greater Antilles, Lesser Antilles. Lepanthes helleri A.D. Hawkes Records : F. Quesada 1981 (INB), M.A. Thomas 1669 Epiphyticherbs.Distribution in PILA 1500–1700m,mixed (MO). forest in transition to cloud forest. Global distribution: Nicaragua to Panama. Records: Luer 1331 (SEL).

Habenaria heptadactyla Rchb.f. Terrestrial herbs. Distribution in PILA – 700 m, low Lepanthes horichii Luer elevation mixed forest. Global distribution: Costa Rica Epiphytic herbs. Distribution in PILA 700–800 m, low to Brazil, Lesser Antilles. Records: A. Rodríguez 9767 elevation mixed forest, mixed forest in transition to low (INB). elevation forest. Global distribution: Endemic to Costa Rica. Records: D. Solano 3821 (BM, INB, MO, PMA).

Lepanthes chameleon Ames Epiphytic herbs. Distribution in PILA 1800–2400 m, Lepanthes horrida Rchb. f. cloud forest, low elevation oak forest. Global distribution: Epiphytic herbs. Distribution in PILA 1500–2600 m, Endemic to Costa Rica. Records: G. Davidse 28647 (CR, mixed forest in transition to cloud forest, cloud forest, MO). low elevation oak forest, high elevation oak forest. Global distribution: Costa Rica, Panama. Records: E.

Lepanthes davidsei Luer Alfaro 3689 (INB), E. Alfaro 3690 (INB), E. Alfaro 4592 Epiphytic herbs. Distribution in PILA 2400–2600 m, low (INB), E. Alfaro 4644 (INB), E. Alfaro 4645 (INB), G. elevation oak forest, high elevation oak forest. Global Herrera 6130 (CR, INB, MO), G. Herrera 6221 (INB), distribution: Endemic to Costa Rica. Records: A.K. M.A. Thomas 1643 (MO). Monro 5598 (BM, INB, MO, PMA), G. Davidse 28648 (CR, MO), G. Davidse 28801 (CR, MO). Lepanthes lindleyana Oerst. & Rchb.f. Epiphytic herbs.Distribution in PILA 1400–2000m,mixed

Lepanthes disticha (A.Rich. & Galeotti) Garay & R.E.Schult. forest in transition to cloud forest. Global distribution: Epiphytic herbs. Distribution in PILA 400–1500 m, low Costa Rica, Panama. Records: R. Quirós 92 (INB). elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest. Lepanthes mariposa Luer Global distribution: Mexico to Panama. Records: G. Epiphytic herbs. Distribution in PILA 1100–1700 m, Herrera 5255 (CR, MO). mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest. Global distribution:

Lepanthes elata Rchb.f. Endemic to Panama. Records: Luer 1598 (SEL). Epiphytic herbs. Distribution in PILA 1500–2600 m, mixed forest in transition to cloud forest, cloud forest, Lepanthes psyche Luer low elevation oak forest, high elevation oak forest. Epiphytic herbs. Distribution in PILA 1000–1300 m, Global distribution: Costa Rica to Colombia. Records: A. mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest. Global distribution: Terrestrial herbs. Distribution in PILA 2400–2800 m, low Endemic to Panama. Records: Luer 1158 ( SEL) . elevation oak forest, high elevation oak forest. Global distribution: Endemic to Costa Rica. Records: G. Davidse ε-PILA Lepanthes trichidion Luer 29049 ( MO) . Epiphytic herbs. Distribution in PILA 2300– 2300 m, low elevation oak forest. Global distribution: Endemic to Masdevallia maduroi Luer Panama. Records: Luer 9307 ( SEL) . Epiphytic herbs. Distribution in PILA 1200– 1200 m, mixed forest in transition to low elevation forest. Global

Lepanthes valerioi Luer distribution: Endemic to Panama. Records : Luer 18932 Epiphytic herbs. Distribution in PILA 2200–2600 m, low (MO). elevation oak forest, high elevation oak forest. Global distribution: Endemic to Costa Rica. Records: M. Valerio Masdevallia nidifica Rchb.f. 179 (CR, MO). Epiphytic herbs. Distribution in PILA 500–2500 m, low elevation mixed forest, mixed forest in transition to low

Lepanthes wendlandii Rchb. f. elevation forest, mixed forest in transition to cloud forest, Epiphytic herbs. Distribution in PILA 1800–2900 m, cloud forest, low elevation oak forest. Global distribution: cloud forest, low elevation oak forest, high elevation Nicaragua to Peru. Records: G. Herrera 5173 (CR, INB, oak forest. Global distribution: Costa Rica, Panama. MO), G. Herrera 5257 (CR, MO). Records: B. Gamboa 1212 (INB, MO), E. Alfaro 3408 (INB), E. Alfaro 3652 (INB), E. Alfaro 3738 (INB), G. Masdevallia rafaeliana Luer Davidse 25131 (CR, MO), G. Davidse 28881 (CR, MO), Epiphytic herbs. Distribution in PILA 2600–3000 m, low G. Davidse 28899 (MO), G. Davidse 29241 (MO), G. elevation oak forest, high elevation oak forest. Global Herrera 3408 (MO), F. Quesada 1581 (INB), F. Quesada distribution: Costa Rica, Panama. Records: L.D. Gómez 1582 (INB), F. Quesada 1990 (INB, MO), L.D. Gómez 22225 (MO). 22231 (MO), S. Merz 350 (CR, MO). Masdevallia utriculata Luer

Liparis arnoglossophylla (Rchb.f.) Rchb.f. ex Hemsl. Epiphytic herbs. Distribution in PILA 1800–2200 m, Facultatively epiphytic herbs. Distribution in PILA cloud forest, low elevation oak forest. Global distribution: 2000–3000 m, cloud forest, low elevation oak forest, Endemic to Panama. Records: R. Dressler s.n. (MO). high elevation oak forest. Global distribution: Mexico to Panama. Records: F. Quesada 1620 (INB). Maxillaria adolphi (Schltr.) Ames & Correll Epiphytic herbs. Distribution in PILA 1900–3300 m,

Lockhartia amoena Rchb. f. cloud forest, low elevation oak forest, páramo. Global Trencilla (CR). distribution: Costa Rica, Panama. Records: A. Rodríguez Epiphytic herbs. Distribution in PILA 300–1800 m, 10850 (BM, INB, MO, PMA), B. Gamboa 1744 (INB, low elevation mixed forest, mixed forest in transition to MO), E. Alfaro 3265 (INB), G. Davidse 23870 (MO), low elevation forest, mixed forest in transition to cloud G. Davidse 25922 (MO), G. Davidse 29001 (CR, MO), forest. Global distribution: Guatemala to Peru. Records: G. Davidse 29255 (INB, MO), J. F. Morales 5806 (INB, L.D. Gómez 23117 (MO). MO), F. Quesada 1841 (INB).

Lycaste leucantha (Klotzsch) Lindl. Maxillaria amabilis J.T.Atwood Epiphytic herbs. Distribution in PILA 1000–1700 m, Epiphytic herbs. Distribution in PILA 1200–1200 mixed forest in transition to low elevation forest, mixed m, mixed forest in transition to low elevation forest. forest in transition to cloud forest. Global distribution: Global distribution: Endemic to Costa Rica. Records: A. Costa Rica, Panama. Records: E. Alfaro 730 (INB, MO), Fernández 679 (INB). G. Davidse 28502 (CR, MO). Maxillaria biolleyi (Schltr.) L.O.Williams

Malaxis excavata (Lindl.) Kuntze Epiphytic herbs. Distribution in PILA 1400–3000 m, Terrestrial herbs. Distribution in PILA 1600–2500 m, mixed forest in transition to cloud forest, cloud forest, mixed forest in transition to cloud forest, cloud forest, low low elevation oak forest, high elevation oak forest. Global elevation oak forest. Global distribution: Guatemala to distribution: Costa Rica, Panama. Records: E. Alfaro 3393 Colombia. Records: G. Davidse 28541 (MO), G. Davidse (INB). 28662 (MO), G. Herrera 6091 (CR, INB, MO), M.A. Thomas 1649 (MO). Maxillaria bracteata (Schltr.) Ames & Correll Pajarillo (CR) .

Malaxis tonduzii (Schltr.) Ames Epiphytic herbs. Distribution in PILA 1100–2500 m, mixed forest in transition to low elevation forest, Epiphyticherbs.Distribution in PILA 1200–2500m,mixed mixed forest in transition to cloud forest, cloud forest, forest in transition to low elevation forest, mixed forest in low elevation oak forest. Global distribution: Costa Rica, transition to cloud forest, cloud forest, low elevation oak Panama. Records: M.A. Thomas 1672 (MO), M. Moraga forest. Global distribution: Costa Rica, Panama. Records: 503 (CR, INB). D. Solano 3853 (BM, INB, MO, PMA).

Maxillaria bradeorum (Schltr.) L.O.Williams Maxillaria horichii Senghas Facultatively epiphytic herbs. Distribution in PILA Epiphytic herbs. Distribution in PILA 900–1500 m, mixed 1000–2000 m, mixed forest in transition to low elevation forest in transition to low elevation forest, mixed forest in forest, mixed forest in transition to cloud forest. Global transition to cloud forest. Global distribution: Costa Rica, distribution: Costa Rica, Panama. Records: D. Solano Panama. Records: E. Navarro 207 (INB, MO). 4478 (CR). Maxillaria nutantiflorum L.O. Williams

Maxillaria brevilabia Ames & Correll Epiphytic herbs. Distribution in PILA 400–900 m, low Epiphytic herbs. Distribution in PILA 1600–2500 m, elevation mixed forest, mixed forest in transition to low mixed forest in transition to cloud forest, cloud forest, elevation forest. Global distribution: Costa Rica, Panama. low elevation oak forest. Global distribution: Costa Rica, Records: A. Fernández 791 (CR, INB), A. Rodríguez Panama. Records: J. F. Morales 5829 (INB, MO), M. 10659 (BM, INB, MO, PMA), D. Santamaría 5775 (BM, Valerio 169 (CR, INB, MO). INB, MO, PMA), D. Solano 4018 (BM, INB, MO, PMA), G. Herrera 5939 (INB, MO).

Maxillaria carinulata Rchb.f. Facultatively epiphytic herbs. Distribution in PILA 1100– Maxillaria microphyton Schltr. 2200 m, mixed forest in transition to low elevation forest, Epiphytic herbs. Distribution in PILA 1000–2000 m, mixed forest in transition to cloud forest, cloud forest, low mixed forest in transition to low elevation forest, mixed elevation oak forest. Global distribution: Costa Rica to forest in transition to cloud forest. Global distribution: Ecuador. Records: G. Herrera 5374 (INB, MO). Costa Rica, Panama. Records: G. Herrera 2724 (MO).

Maxillaria concavilabia Ames & Correll Maxillaria obscura Linden & Rchb.f. Epiphytic herbs. Distribution in PILA – m. Global Epiphytic herbs. Distribution in PILA 1500–3000 m, distribution: Endemic to Costa Rica. Records: M. Moraga mixed forest in transition to cloud forest, cloud forest, 500 (INB, MO). low elevation oak forest, high elevation oak forest. Global distribution: Costa Rica, Panama. Records: B. Gamboa

Maxillaria confusa Ames & C.Schweinf. 1223 (INB), E. Alfaro 3621 (INB), G. Herrera 5974 (CR, Epiphytic herbs. Distribution in PILA 0–1500 m, low INB, MO), F. Quesada 1954 (INB). elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest. Maxillaria paleata (Rchb.f.) Ames & Correll Global distribution: Belize to Panama. Records: F. Solís Epiphyticherbs.Distribution in PILA 1200–3300m,mixed 20 (CR, INB, MO). forest in transition to low elevation forest, mixed forest in transition to cloud forest, cloud forest, low elevation oak

Maxillaria falcata Ames & Correll forest, páramo. Global distribution: Nicaragua to Panama. Epiphytic herbs. Distribution in PILA 1500–2400 m, Records: A. Chacón 498 (MO), A. Rojas 3244 (INB, mixed forest in transition to cloud forest, cloud forest, MO), B. Gamboa 1205 (INB), B. Gamboa 1713 (INB, low elevation oak forest. Global distribution: Costa Rica, MO), B. Hammel 5795 (MO), G. Davidse 25123 (MO), Panama. Records: D. Santamaría 5733 (BM, INB, MO, G. Davidse 28610 (INB, MO), G. Davidse 28877 (CR, PMA), G. McPherson 15919 (MO), J. Bittner 1809 (INB, MO), G. Davidse 28919 (CR, MO), G. Davidse 28994 MO). (CR, MO), G. Davidse 29275 (INB, MO), G. Herrera 3628 (MO), G. Herrera 3708 (INB, MO), K. Sytsma 4925

Maxillaria fulgens (Rchb.f.) L.O.Williams (MO) , K. Sytsma 4939 (MO), M. Valerio 203 (INB), S. Epiphytic herbs. Distribution in PILA 500–2000 m, Knapp 2161 (MO). low elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to cloud Maxillaria parvilabia Ames & C.Schweinf. forest. Global distribution: Nicaragua to Peru, Venezuela. Epiphytic herbs. Distribution in PILA 1500–2300 m, Records: G. Davidse 28419 (CR, MO), J. F. Morales mixed forest in transition to cloud forest, cloud forest, 5828 (INB, MO). low elevation oak forest. Global distribution: Costa Rica, Panama. Records: D. Santamaría 5847 (BM, INB, MO,

Maxillaria gomeziana J.T.Atwood PMA ), J. Bittner 1775 (INB, MO), M. Grayum 10351 Terrestrial herbs. Distribution in PILA 1500–2100 m, (INB, MO). mixed forest in transition to cloud forest, cloud forest, low elevation oak forest. Global distribution: Costa Rica,

Maxillaria pseudoneglecta J.T.Atwood Panama. Records: J. Bittner 1910 (INB, MO). Epiphytic herbs. Distribution in PILA 1100– 1100 m, mixed forest in transition to low elevation forest. Global Myoxanthus colothrix (Luer) Luer distribution: Costa Rica, Panama. Records: E. Alfaro 729 Epiphytic herbs. Distribution in PILA 1700– 1700 m, (INB). cloud forest. Global distribution: Costa Rica, Panama, Ecuador. Records: E. Alfaro 3464 (INB).

Maxillaria quadrata Ames & Correll Facultatively epiphytic herbs. Distribution in PILA Nidema boothii (Lindl.) Schltr. 1700 –2800 m, cloud forest, low elevation oak forest, Epiphytic herbs. Distribution in PILA 0–1400 m, low high elevation oak forest. Global distribution: Costa Rica, elevation mixed forest, mixed forest in transition to low Panama. Records: G. Davidse 25771 (MO). elevation forest, mixed forest in transition to cloud forest. Global distribution: Mexico to Panama. Records: L.D.

ε-PILA Maxillaria synsepalum J.T.Atwood Gómez 23286 (CR, MO). Epiphytic herbs. Distribution in PILA 2200– 2200 m, low elevation oak forest. Global distribution: Endemic to Oncidium bryolophotum Rchb.f. Panama. Records: Van der Werff 6435 (MO, SEL). Facultatively epiphytic herbs. Distribution in PILA 1100–2000 m, mixed forest in transition to low elevation

Maxillaria tigrina C.Schweinf. forest, mixed forest in transition to cloud forest. Global Epiphytic herbs. Distribution in PILA 2000–2500 m, distribution: Costa Rica to Peru. Records: D. Solano 3891 cloud forest, low elevation oak forest. Global distribution: (BM, INB, MO, PMA). Costa Rica, Panama. Records: G. Davidse 28634 (CR, MO), F. Quesada 1596 (INB). Oncidium cheirophorum Rchb.f. Facultatively epiphytic herbs. Distribution in PILA ε-PILA Maxillaria tricarinatum J.T.Atwood 1100 –2000 m, mixed forest in transition to low elevation Epiphytic herbs. Distribution in PILA 1500–1500 forest, mixed forest in transition to cloud forest. Global m, mixed forest in transition to cloud forest. Global distribution: Mexico to Colombia. Records: J. Dwyer 440 distribution: Endemic to Panama. Records: R. Dressler (MO). 5810 (SEL). Oncidium lineoligerum Rchb.f. & Warsz.

Maxillaria tubercularis J.T.Atwood Facultatively epiphytic herbs. Distribution in PILA Epiphytic herbs. Distribution in PILA 1300–3000 m, – m. Global distribution: Elevation unkown. Costa Rica, mixed forest in transition to cloud forest, cloud forest, Panama. Records: L.D. Gómez 23257 (CR, MO). low elevation oak forest, high elevation oak forest. Global distribution: Costa Rica, Panama. Records: A. Chacón Oncidium maduroi Dressler 490 (INB, MO), A. Soto 333 (INB), B. Gamboa 565 (INB, Facultatively epiphytic herbs. Distribution in PILA MO), G. Davidse 23979 (CR, MO). 1200–1400 m, mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest. Global

Maxillaria umbratilis L.O. Williams distribution: Costa Rica, Panama. Records: R. Dressler Epiphytic herbs. Distribution in PILA 900–3200 m, mixed 6220 (MO). forest in transition to low elevation forest, mixed forest in transition to cloud forest, cloud forest, low elevation oak Oncidium nebulosum Lindl. forest, páramo. Global distribution: Costa Rica, Panama. Facultatively epiphytic herbs. Distribution in PILA 1000– Records: E. Alfaro 3367 ( INB) , G. Herrera 6231 ( CR, 2100 m, mixed forest in transition to low elevation forest, INB, MO). mixed forest in transition to cloud forest, cloud forest, low elevation oak forest. Global distribution: Costa Rica to

Maxillaria vaginalis Rchb.f. Peru. Records: J. Dwyer 514 (MO). Facultatively epiphytic herbs. Distribution in PILA 1800– 2500 m, cloud forest, low elevation oak forest. Global Oncidium poikilostalix (Kraenzl.) M.W.Chase & distribution: Costa Rica, Panama. Records: A. Fernández N.H.Williams 1854 (INB), E. Navarro 475 (INB, MO), G. Davidse Epiphytic herbs. Distribution in PILA 1200–1500 m, 28539 (INB, MO), G. Davidse 28926 (MO). mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest. Global distribution:

Microchilus nigrescens (Schltr.) Ormerod Mexico to Ecuador. Records: Davidse 28460 (CR, MO), forest. Global distribution: Mexico to Brazil. Records: E. Davidse 28459 (CR, MO), Herrera 5102 (CR, INB, Alfaro 4643 (INB), G. Davidse 28802 (MO), G. Herrera MO ). 6106 (CR, INB, MO).