Aspidogyne utriculata (Dressler) Szlach.

MONRO, A. K., SANTAMARÍA-AGUILAR, D., GONZÁLEZ, F., CHACÓN, O., SOLANO, D., RODRÍGUEZ, A., ZAMORA, N., FEDELE, E. & CORREA, M., 2017, A first checklist to the vascular plants of La Amistad International Park (PILA), Costa Rica-Panama, Phytotaxa 322 (1), pp. 448-450 : 448-450

Aspidogyne utriculata (Dressler) Szlach.

Terrestrial herbs. Distribution in PILA 700– 700 m, low Brachionidium satyreum Luer elevation mixed forest. Global distribution: Mexico to Terrestrial herbs. Distribution in PILA 1400–1500 Panama. Records: G. Herrera 3245 (INB, MO). m, mixed forest in transition to cloud forest. Global distribution: Endemic to Panama. Records: R. Dressler

Barbosella geminata Luer 5809 (MO). Epiphytic herbs.Distribution in PILA 1800–2600 m, cloud forest, low elevation oak forest, high elevation oak forest. Calanthe calanthoides (A.Rich. & Galeotti) Hamer & Garay Global distribution: Endemic to Costa Rica. Records: G. Cebollín, Legua de vaca (CR). Herrera 6107 (CR, INB). Terrestrial herbs. Distribution in PILA 800–3200 m, mixed forest in transition to low elevation forest, mixed

Bletia campanulata Lex forest in transition to cloud forest, cloud forest, low Terrestrial herbs. Distribution in PILA 1100–1700 m, elevation oak forest, páramo. Global distribution: Mexico mixed forest in transition to low elevation forest, mixed to Colombia, Greater Antilles, Lesser Antilles. Records: forest in transition to cloud forest. Global distribution: B. Gamboa 561 (INB, MO), D. Solano 5215 (BM, INB, Mexico to Argentina. Records: D. Santamaría 4415 MO, PMA), E. Alfaro 3202 (INB), G. Davidse 25819 (CR, (INB). MO), G. Davidse 28699 (MO), G. Davidse 23970 (MO), G. Herrera 3413 (MO), F. Quesada 1608 (INB, MO), F.

ε-PILA Brachionidium calypso Luer Quesada 1625 (INB), L. Acosta 2408 (INB), L.D. Gómez Epiphytic herbs. Distribution in PILA 2200– 2200 m, 21861 (MO), L. González 1187 (INB), M.A. Thomas 1585 low elevation oak forest. Global distribution: Endemic to (MO), T. Croat 26987 (MO). Panama. Records: R. Dressler 5838 (MO). • ε-PILA Campylocentrum amistadense Bogarín

Brachionidium cruziae L.O.Williams Epiphytic herbs. Distribution in PILA – 1600 m, mixed Epiphytic herbs. Distribution in PILA 2300–2600 m, low forest in transition to cloud forest. Global distribution: elevation oak forest, high elevation oak forest. Global Costa Rica, Panama? Records : E. Fletes & A. Mora 204 distribution: Endemic to Costa Rica. Records: G. Davidse (INB). 28727 (MO), G. Davidse 28807 ( CR, MO) . Cischweinfia dasyandra (Rchb.f.) Dressler & N.H.Williams

?ε-PILA Brachionidium dentatum Luer & Dressler Epiphytic herbs. Distribution in PILA 600–1700 m, low Terrestrial herbs. Distribution in PILA 2800– 2800 m, low elevation mixed forest, mixed forest in transition to low elevation oak forest, high elevation oak forest. Global elevation forest, mixed forest in transition to cloud forest. distribution: Endemic to Panama. Records: R. Dressler Global distribution: Costa Rica to Ecuador. Records: E. s.n. (MO). Navarro 206 (INB, MO). This species is only known from the type, a cultivated plant collected from the vicinity of PILA. Coccineorchis bracteosa (Ames & C.Schweinf.) Garay Facultatively epiphytic herbs. Distribution in PILA 1200–

Brachionidium dressleri Luer 2500 m, mixed forest in transition to low elevation forest, Epiphytic herbs. Distribution in PILA 1200–2500 m, mixed forest in transition to cloud forest, cloud forest, low mixed forest in transition to low elevation forest, mixed elevation oak forest. Global distribution: Nicaragua to forest in transition to cloud forest, cloud forest, low Panama, Ecuador. Records: B. Gamboa 563 (INB, MO), elevation oak forest. Global distribution: Costa Rica, D. Solano 3945 (BM, INB, MO, PMA), E. Alfaro 3465 Panama. Records: K. Sytsma 4908 (MO). (INB), E. Alfaro 3822 (INB), G. Davidse 28869 (CR, MO), G. Davidse 28901 (CR, MO), G. Davidse 29036

ε-PILA Brachionidium lucanoideum Luer (CR, MO) , L. González 1178 (INB). Epiphytic herbs. Distribution in PILA 2200– 2200 m, low elevation oak forest. Global distribution: Endemic to Coccineorchis cernua (Lindl.) Garay Panama. Records: S. Knapp 4829 (BM, PMA, MO). Facultatively epiphytic herbs. Distribution in PILA 1000– 2600 m, mixed forest in transition to low elevation forest,

Brachionidium polypodium Luer mixed forest in transition to cloud forest, cloud forest, Terrestrial herbs. Distribution in PILA 1200–1500 m, low elevation oak forest, high elevation oak forest. Global mixed forest in transition to low elevation forest, mixed distribution: Mexico to Peru. Records: G. Herrera 3419 forest in transition to cloud forest, pasture. Global (MO). distribution: Costa Rica, Panama. Records: A. Rodríguez 11537 (INB). Coccineorchis standleyi (Ames & C.Schweinf.) Garay Facultatively epiphytic herbs. Distribution in PILA

400–2800 m, low elevation mixed forest, mixed forest in elevation mixed forest, mixed forest in transition to low transition to low elevation forest, mixed forest in transition elevation forest, mixed forest in transition to cloud forest, to cloud forest, cloud forest, low elevation oak forest, cloud forest, low elevation oak forest. Global distribution: high elevation oak forest. Global distribution: Nicaragua Guatemala to Honduras, Costa Rica, Panama. Records: to Colombia. Records: D. Santamaría 5767 (BM, INB, G. Davidse 28407 (CR, MO), L. Acosta 2410 (INB). MO, PMA). Dichaea eligulata Folsom

Cochleanthes aromatica (Rchb.f.) R.E.Schult. & Garay Epiphytic herbs. Distribution in PILA 600– 600 m, low Epiphytic herbs. Distribution in PILA 900–1500 m, mixed elevation mixed forest. Global distribution: Costa Rica, forest in transition to low elevation forest, mixed forest in Panama. Records: G. Herrera 6099 (CR, INB, MO). transition to cloud forest. Global distribution: Costa Rica, Panama. Records: D. Solano 5569 (PMA). Dichaea oxyglossa Schltr. Epiphytic herbs. Distribution in PILA 1000–1700 m,

Cranichis lankesteri Ames mixed forest in transition to low elevation forest, mixed Terrestrial herbs. Distribution in PILA 1200–2300 m, forest in transition to cloud forest. Global distribution: mixed forest in transition to low elevation forest, mixed Costa Rica, Panama. Records: M. Chinchilla 196 (INB). forest in transition to cloud forest, cloud forest, low elevation oak forest. Global distribution: Costa Rica to Dichaea trulla Rchb.f. Panama. Records: S. Knapp 1500 (MO). Epiphytic herbs. Distribution in PILA 200–500 m, low elevation mixed forest. Global distribution: Guatemala to

Cranichis reticulata Rchb.f. Brazil. Records : D. Solano 5868 (PMA). Terrestrial herbs. Distribution in PILA 1500–2700 m, mixed forest in transition to cloud forest, cloud forest, Dracula erythrochaete (Rchb.f.) Luer low elevation oak forest, high elevation oak forest. Global Epiphytic herbs. Distribution in PILA 1500–2900 m, distribution: Endemic to Costa Rica. Records: G. Davidse mixed forest in transition to cloud forest, cloud forest, 28528 ( MO) . low elevation oak forest, high elevation oak forest. Global distribution: Costa Rica, Panama. Records: G. Davidse ε-PILA Cranichis talamancana Dressler 25135 ( MO) . Terrestrial herbs. Distribution in PILA 2400–2500 m, low elevation oak forest. Global distribution: Endemic to Dracula ripleyana Luer Costa Rica. Records: G. Davidse 28651 ( MO) . Epiphytic herbs. Distribution in PILA – 2200 m, low elevation oak forest. Global distribution: Endemic to

Crossoglossa blephariglottis (Schltr.) Dressler Costa Rica. Records: G. Herrera 6135 (CR, MO). Epiphytic herbs. Distribution in PILA 1100–2000 m, mixed forest in transition to low elevation forest, mixed Echinosepala aspasicensis (Rchb.f.) Pridgeon & M.W.Chase forest in transition to cloud forest. Global distribution: Epiphytic herbs. Distribution in PILA 1000–1300 m, Costa Rica, Panama. Records: A. Rodríguez 10744 ( BM, mixed forest in transition to low elevation forest, mixed INB, MO, PMA), B. Hammel 6108 ( MO) . forest in transition to cloud forest. Global distribution: Costa Rica to Peru, the Guyanas, Brazil. Records: F.

Cyrtochiloides panduriformis (Ames & C.Schweinf.) Quesada 1601 (INB). N.H.Williams & M.W.Chase Facultatively epiphytic herbs. Distribution in PILA 1200– Echinosepala uncinata (Fawc.) Pridgeon & M.W.Chase 2400 m, mixed forest in transition to low elevation forest, Epiphytic herbs. Distribution in PILA 0–1700 m, low mixed forest in transition to cloud forest, cloud forest, elevation mixed forest, mixed forest in transition to low low elevation oak forest. Global distribution: Costa Rica, elevation forest, mixed forest in transition to cloud forest, Panama. Records: G. Herrera 6109 (INB). cloud forest. Global distribution: Belize to Panama, Greater Antilles. Records: G. Herrera 5179 (CR, INB,

Dichaea ciliolata Rolfe MO ). Epiphytic herbs. Distribution in PILA 700–1600 m, low elevation mixed forest, mixed forest in transition to low Elleanthus aurantiacus (Lindl.) Rchb. f. elevation forest. Global distribution: Costa Rica, Panama. Facultatively epiphytic herbs. Distribution in PILA Records: A. Rodríguez 11341 (BM, INB, MO, PMA), A. 1100–2800 m, mixed forest in transition to low elevation Rodríguez 11345 (BM, INB, MO, PMA). forest, mixed forest in transition to cloud forest, cloud forest, low elevation oak forest, high elevation oak forest.

Dichaea cryptarrhena Rchb.f. ex Kraenzl. Global distribution: Nicaragua to Venezuela. Records: B. Epiphytic herbs. Distribution in PILA 300–2200 m, low Gamboa 1210 (INB, MO), D. Solano 3908 (BM, INB, Terrestrial herbs. Distribution in PILA 1200–3200 MO, PMA), M.A. Thomas 1651 (MO). m, mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest, cloud forest,

Elleanthus capitatus (Poepp. & Endl.) Rchb. f. low elevation oak forest, páramo. Global distribution: Epiphytic herbs. Distribution in PILA 2300– 2300 m, low Nicaragua, Costa Rica. Records: G. Davidse 25400 (MO), elevation oak forest. Global distribution: Mexico to Costa G. Davidse 25933 (INB, MO). Rica. Records: G. Davidse 28451 (CR, MO), F. Quesada 1682 (CR, INB, MO). Epidendrum adnatum Ames & C.Schweinf. Epiphytic herbs. Distribution in PILA 1100–1700 m,

Elleanthus glaucophyllus Schltr. mixed forest in transition to low elevation forest, mixed Facultatively epiphytic herbs. Distribution in PILA forest in transition to cloud forest. Global distribution: 500–2400 m, low elevation mixed forest, mixed forest Costa Rica, Panamá. Records: G. Herrera 2669 (INB, in transition to low elevation forest, mixed forest in MO), G. Herrera 5489 (CR, MO). transition to cloud forest, cloud forest, low elevation oak forest. Global distribution: Costa Rica, Panama. Records: Epidendrum alfaroi Ames & C.Schweinf. A. Rodríguez 10666 (BM, INB, MO, PMA), D. Solano Epiphytic herbs. Distribution in PILA 400–2200 m, low 3905 (BM, INB, MO, PMA), E. Alfaro 2266 (INB), G. elevation mixed forest, mixed forest in transition to low Herrera 3262 (INB, MO), L. Angulo 330 (INB). elevation forest, mixed forest in transition to cloud forest, cloud forest, low elevation oak forest. Global distribution:

Elleanthus graminifolius (Barb. Rodr.) Lojtnant Costa Rica to Colombia. Records: F. Solís 23 (INB, Epiphytic herbs. Distribution in PILA 0–900 m, low MO). elevation mixed forest, mixed forest in transition to low elevation forest. Global distribution: Mexico to Brazil, Epidendrum anoglossoides Ames & C.Schweinf. Greater Antilles, Lesser Antilles. Records: G. Gallardo Epiphytic herbs. Distribution in PILA 1600–3200 m, 302 (CR, INB, MO). mixed forest in transition to cloud forest, cloud forest, low elevation oak forest, páramo. Global distribution: Costa

Elleanthus hymenophorus (Rchb.f.) Rchb.f. Rica , Panama. Records: E. Alfaro 3622 (INB), G. Davidse Facultatively epiphytic herbs. Distribution in PILA 25929 (INB, MO), G. Davidse 26070 (MO), G. Davidse 200–3000 m, low elevation mixed forest, mixed forest in 28833 (CR, MO), G. Herrera 6233 (CR, INB, MO). transition to low elevation forest, mixed forest in transition to cloud forest, cloud forest, low elevation oak forest, Epidendrum anoglossum Schltr. high elevation oak forest. Global distribution: Mexico to Epiphytic herbs. Distribution in PILA 1300–3100 m, Ecuador. Records: D. Solano 3193 (INB), D. Solano 5211 mixed forest in transition to cloud forest, cloud forest, low (BM, INB, MO, PMA), E. Fletes 199 (INB), L. Angulo elevation oak forest, high elevation oak forest, páramo. 319 (INB). Global distribution: Costa Rica, Panama. Records: E. Alfaro 3274 (INB), G. Davidse 25134 (CR, MO), F.

Elleanthus tonduzii Schltr. Quesada 1998 (INB), M. Valerio 172 (CR, MO). Facultatively epiphytic herbs. Distribution in PILA 1000– 3100 m, mixed forest in transition to low elevation forest, Epidendrum atrorugosum Hágsater mixed forest in transition to cloud forest, cloud forest, low Epiphytic herbs. Distribution in PILA 2300–2400 m, elevation oak forest, high elevation oak forest, páramo. low elevation oak forest. Global distribution: Endemic Global distribution: Costa Rica, Panama. Records: A. to Costa Rica. Records: G. Davidse 28737 (MO), M. Rodríguez 10842 (BM, INB, MO, PMA), D. Santamaría Grayum 10334 (INB, MO). 5763 (BM, INB, MO, PMA), E. Alfaro 3263 (INB), G. Davidse 25139 (MO), G. Davidse 28580 (MO), G. Epidendrum barbae Rchb.f. Davidse 28785 (MO), G. Herrera 5454 (CR, INB, MO), Epiphytic herbs. Distribution in PILA 1300–2600 m, F. Quesada 1828 (INB), L. González 3620 (INB). mixed forest in transition to cloud forest, cloud forest, low elevation oak forest, high elevation oak forest. Global

Elleanthus tricallosus Ames & C.Schweinf. distribution: Costa Rica, Panama. Records: E. Alfaro Epiphytic herbs. Distribution in PILA 900–2600 m, mixed 3280 (INB), E. Alfaro 3399 (INB), E. Alfaro 3963 (INB), forest in transition to low elevation forest, mixed forest G. Davidse 28544 (MO), G. Davidse 28716 (MO), G. in transition to cloud forest, cloud forest, low elevation Davidse 28870 (MO), F. Quesada 1589 (INB). oak forest, high elevation oak forest. Global distribution: Costa Rica, Panama. Records: E. Alfaro 3812 (INB). Epidendrum cardiochilum L.O.Williams Epiphytic herbs. Distribution in PILA 2400–2600 m, low

Elleanthus wercklei Schltr. elevation oak forest, high elevation oak forest. Global Rica, Panama. Records: G. Davidse 26196 (MO). distribution: Mexico to Costa Rica. Records: G. Davidse 28631 ( MO) , G. Davidse 28832 (MO). Epidendrum horichii Hágsater Epiphytic herbs. Distribution in PILA 1200–1800 m,

Epidendrum chirripoense Hágsater mixed forest in transition to low elevation forest, mixed Epiphytic herbs. Distribution in PILA 2600–3200 m, low forest in transition to cloud forest. Global distribution: elevation oak forest, high elevation oak forest, páramo. Endemic to Costa Rica. Records: A. Rodríguez 11365 Global distribution: Costa Rica, Panama. Records: G. (INB). Davidse 23912 (CR, MO). • ε-PILA Epidendrum jorgewarneri Karremans & Hágsater

Epidendrum confertum Ames & C.Schweinf. Epiphytic herbs. Distribution in PILA – 2500 m, low Epiphytic herbs. Distribution in PILA 200–400 m, low elevation oak forest, páramo. Global distribution:Endemic elevation mixed forest, pasture. Global distribution: Costa to Costa Rica. Records: A. Karremans et al. 5545 (CR, Rica, Panama. Records: A. Rodríguez 10673 (BM, INB, JBL). MO, PMA). Epidendrum kerichilum Hágsater

Epidendrum davidsei Hágsater Facultatively epiphytic herbs. Distribution in PILA 1700– Epiphytic herbs. Distribution in PILA 3000–3300 m, 2300 m, cloud forest, low elevation oak forest. Global high elevation oak forest, páramo. Global distribution: distribution: Endemic to Panama. Records: Hágsater Endemic to Costa Rica. Records: E. Alfaro 1323 (INB, 6519 (AMO, K, MO). MO). Epidendrum lacustre Lindl.

Epidendrum endresii Rchb.f. Epiphytic herbs. Distribution in PILA 1100–1900 m, Epiphytic herbs. Distribution in PILA 1400–2400 m, mixed forest in transition to low elevation forest, mixed mixed forest in transition to cloud forest, cloud forest, forest in transition to cloud forest. Global distribution: low elevation oak forest. Global distribution: Costa Rica, Nicaragua to Ecuador, Venezuela. Records: D. Solano Panama. Records: Chacón 580 (MO), Gamboa 1236 4763 (MEXU, MO, SEL). (INB, MO), McPherson 15917 (MO). Epidendrum lagenocolumna Hágsater

Epidendrum erythrostigma Hágsater Epiphytic herbs. Distribution in PILA 1400–2500 m, Epiphytic herbs. Distribution in PILA 2100–2400 m, mixed forest in transition to cloud forest, cloud forest, cloud forest, low elevation oak forest. Global distribution: low elevation oak forest. Global distribution: Honduras Costa Rica, Panama. Records: A. Fernández 680 (CR, to Panama. Records: D. Santamaría 7161 (CR, MO), D. INB, MO), G. Davidse 28784 (CR, MO), M. Grayum Santamaría 7246 (CR, MO), Zamora 4117 (CR, MO), N. 10345 (CR, MO). Zamora 4144 (CR, MO), N. Zamora 4150 (CR, MO).