Dicranopygium wedelii Harling

MONRO, A. K., SANTAMARÍA-AGUILAR, D., GONZÁLEZ, F., CHACÓN, O., SOLANO, D., RODRÍGUEZ, A., ZAMORA, N., FEDELE, E. & CORREA, M., 2017, A first checklist to the vascular plants of La Amistad International Park (PILA), Costa Rica-Panama, Phytotaxa 322 (1), pp. 448-450 : 448-450

Dicranopygium wedelii Harling

Epipetric herbs. Distribution in PILA 0–1500 m, low Sphaeradenia pachystigma Harling elevation mixed forest, mixed forest in transition to low Facultatively epiphytic herbs. Distribution in PILA 800– elevation forest, mixed forest in transition to cloud forest. 2300 m, mixed forest in transition to low elevation forest, Global distribution: Nicaragua to Panama. Records: G. mixed forest in transition to cloud forest, cloud forest, Herrera 3039 (MO). low elevation oak forest. Global distribution: Costa Rica, Panama. Records: J. H. Kirkbride Jr. 140 (MO).

Evodianthus funifer (Poit.) Lindm. Chidra, Chirrabaca, Tucuso (CR) . Sphaeradenia praetermissa R.Erikss. Hemi-epiphytic herbs. Distribution in PILA 0–1500 Facultatively epiphytic herbs. Distribution in PILA 1000–

m, low elevation mixed forest, mixed forest in transition 2500 m, mixed forest in transition to low elevation forest, to low elevation forest, mixed forest in transition to cloud mixed forest in transition to cloud forest, cloud forest, forest. Global distribution: Nicaragua to Peru, Venezuela, low elevation oak forest, pasture. Global distribution: Guayanas, Trinidad, Tobago, Brazil. Records: A. Chacón Endemic to Costa Rica. Records: A. Rodríguez 11546 275 (CR, INB, MO), A. Fernández 1000 (INB), A.K. (BM, INB, MO, PMA), G. Davidse 29090 (CR, MO), G. Monro 6263 (PMA), A. Rodríguez 11346 (BM, INB, Herrera 2664 (INB, MO). PMA), A. Rodríguez 11620 (BM, INB, PMA), D. Santamaría 6790 (BM, INB, PMA), D. Santamaría 7687 ε-PILA Sphaeradenia rostellata R.Erikss. (PMA) , D. Santamaría 7853 (PMA). Facultatively epiphytic herbs. Distribution in PILA 1500–2300 m, mixed forest in transition to cloud forest,

Sphaeradenia chiriquensis Harling cloud forest, low elevation oak forest. Global distribution: Facultatively epiphytic herbs. Distribution in PILA Endemic to Costa Rica. Records: A. Fernández 741 1500–3300 m, mixed forest in transition to cloud forest, (INB), A. Fernández 970 (INB), D. Solano 3900 (BM, cloud forest, low elevation oak forest, páramo. Global INB, PMA), J. Bittner 1788 (INB, MO). distribution: Costa Rica, Panama. Records: G. Davidse 25206 (MO), G. Davidse 28639 (CR, MO), G. Davidse CYPERACEAE 28920 (MO), G. McPherson 15903 (MO). Carex bonplandii Kunth

Sphaeradenia garciae Harling Terrestrial herbs. Distribution in PILA 2400–3600 m, low Facultatively epiphytic herbs. Distribution in PILA elevation oak forest, páramo. Global distribution: USA, 600–1700 m, low elevation mixed forest, mixed forest Guatemala, Costa Rica to Colombia. Records: E. Alfaro in transition to low elevation forest, mixed forest in 3283 (INB), G. Davidse 23896 (CR, MO), G. Davidse transition to cloud forest. Global distribution: Costa Rica 25915 (CR, MO), G. Davidse 26000 (CR, MO), G. to Ecuador. Records: D. Santamaría 6481 (BM, INB, Davidse 29304 (CR, MO), G. Davidse 29330 (CR, MO), PMA), G. Herrera 5377 (INB). G. Davidse 29371 (CR, MO), G. Davidse 29385 (CR, MO).

Sphaeradenia laucheana (Mast.) Harling Facultatively epiphytic herbs. Distribution in PILA Carex donnellsmithii L.H.Bailey 1600 –2800 m, mixed forest in transition to cloud forest, Terrestrial herbs. Distribution in PILA 1300–3500 m, cloud forest, low elevation oak forest, high elevation oak mixed forest in transition to cloud forest, cloud forest, low forest. Global distribution: Costa Rica, Colombia to Peru. elevation oak forest, páramo. Global distribution: Mexico Records: A. Soto 287 (INB), B. Gamboa 1240 (INB). to Panama. Records: A.K. Monro 5250 (BM, INB, MO), A. S. Weston 10184 (MO), B. Gamboa 1731 (INB), E.

Sphaeradenia magniglobula R.Erikss. Alfaro 3292 (INB), G. Davidse 25114 (CR, MO), G. Terrestrial herbs. Distribution in PILA 1500–2500 m, Davidse 25266 (CR, MO), G. Davidse 25381 (CR, MO), mixed forest in transition to cloud forest, cloud forest, G. Davidse 25508 (CR, MO), G. Davidse 25815 (CR, low elevation oak forest. Global distribution: Costa Rica, MO), G. Davidse 25842 (CR, MO), G. Davidse 25954 Panama. Records: G. Davidse 29108 (MO), G. Herrera (CR, MO) , G. Davidse 26036 (CR, MO), G. Davidse 5438 (CR, INB, MO), J. Bittner 1865 (INB). 29322 (CR, MO), G. Davidse 29351 (MO), G. Davidse 23918 (MO), L. Angulo 211 (INB, MO), L.D. Gómez

Sphaeradenia occidentalis R.Erikss. 22449 (MO), M. Grayum 10384 (INB, MO), S. Knapp Facultatively epiphytic herbs. Distribution in PILA 10032 (BM, INB). 1500–1800 m, mixed forest in transition to cloud forest. Global distribution: Endemic to Costa Rica. Records: A. Carex jamesonii Boott Rodríguez 10643 (BM, INB, MO, PMA). Terrestrial herbs. Distribution in PILA 1500–3400 m, mixed forest in transition to cloud forest, cloud forest, Cyperus brevifolius (Rottb.) Hassk. low elevation oak forest, páramo. Global distribution: Terrestrial herbs. Distribution in PILA 0–2000 m, low Mexico to Venezuela. Records: A. Chacón 561 (INB, elevation mixed forest, mixed forest in transition to low MO), A. Fernández 907 (CR, INB), E. Alfaro 3284 (INB), elevation forest, mixed forest in transition to cloud forest, G. Davidse 25180 (MO), G. Davidse 28571 (CR, MO), cloud forest. Global distribution: Pantropical. Records: G. Davidse 28593 (MO), G. Davidse 23931 (MO), G. A.K. Monro 6087 (PMA). Herrera 5890 (CR, INB), F. Quesada 1561 (INB). Cyperus hermaphroditus (Jacq.) Standl.

Carex lemanniana Boott Zacate jacintillo (CR). Terrestrial herbs. Distribution in PILA 1600–3500 m, Terrestrial herbs. Distribution in PILA 0–3400 m, low mixed forest in transition to cloud forest, cloud forest, low elevation mixed forest, mixed forest in transition to low elevation oak forest, páramo. Global distribution: Costa elevation forest, mixed forest in transition to cloud forest, Rica to Peru. Records: G. Davidse 25334 (CR, MO), cloud forest, low elevation oak forest, páramo. Global G. Davidse 25399 (MO), G. Davidse 29322A (MO), G. distribution: Mexico to Argentina, Greater Antilles, Lesser Davidse 29336 (CR, MO), G. Davidse 29401 (CR, MO), Antilles. Records: C. Hamilton 3651 (MO), C. Hamilton M.A. Thomas 1551 (MO). 704 (MO), C. Hamilton 721 (MO), S. Bridgewater 4217 (BM, INB, MO, PMA).

Carex polystachya Sw. ex Wahlenb. Terrestrial herbs. Distribution in PILA 300–2400 m, low Cyperus niger Ruiz & Pav. elevation mixed forest, mixed forest in transition to low Terrestrial herbs. Distribution in PILA 1000–2500 elevation forest, mixed forest in transition to cloud forest, m, mixed forest in transition to low elevation forest, cloud forest, low elevation oak forest. Global distribution: mixed forest in transition to cloud forest, cloud forest, USA to Bolivia, Greater Antilles. Records: A. Rodríguez low elevation oak forest. Global distribution: USA to 9849 (INB), B. Gamboa 637 (INB), D. Solano 2889 (INB), Argentina, Africa. Records: C. Hamilton 733 (MO). D. Solano 3195 (INB), G. Davidse 28448 (CR, MO). Cyperus reflexus Vahl.

Carex porrecta Reznicek & Camelbeke Terrestrial herbs. Distribution in PILA – 2400 m, low Terrestrial herbs. Distribution in PILA 1800–3100 m, elevation oak forest. Global distribution: USA to Mexico, cloud forest, low elevation oak forest, high elevation oak Costa Rica, Venezuela to Argentina. Records: A. Chacón forest, páramo. Global distribution: Costa Rica to Ecuador, 556 (INB), G. Herrera 3702 (INB, MO). Venezuela. Records: G. Davidse 29166 (CR, MO). Cyperus tabina Steud. ex Boeck.

Carex purdiei Boott Terrestrial herbs. Distribution in PILA 1300–2500 m, Terrestrial herbs. Distribution in PILA 2500–3500 m, mixed forest in transition to cloud forest, cloud forest, low elevation oak forest, páramo. Global distribution: low elevation oak forest. Global distribution: Costa Guatemala, Costa Rica to Ecuador. Records: G. Davidse Rica, Panama, Ecuador, Bolivia. Records: A. Fernández 25350 (CR, MO), G. Davidse 29339 (MO), G. Davidse 751 (CR, INB), A. Rodríguez 10617 (BM, INB, MO, 29374 (CR, MO). PMA), A. Rodríguez 10916 (BM, INB, MO, PMA), D. Santamaría 5734 (BM, INB, MO, PMA), D. Solano 3944

Carex pygmaea Boeck. (BM, INB, MO, PMA) , E. Navarro 2 (INB), M. Grayum Terrestrial herbs. Distribution in PILA 2300–3500 m, low 10297 (INB, MO). elevation oak forest, páramo. Global distribution: Costa Rica to Peru. Records: G. Davidse 23932 (CR, MO), G. Cyperus tenerrimus J.Presl & C.Presl Davidse 25312 (CR, MO), G. Davidse 25420 (MO), G. Botón de amor (CR). Davidse 25988 (CR, MO), G. Davidse 28566 (CR, MO), Terrestrial herbs. Distribution in PILA 0–0 m, low G. Davidse 29274 (CR, MO), G. Herrera 3654 (MO), G. elevation mixed forest. Global distribution: Mexico to Herrera 3698 (MO), L.D. Gómez 22451 (MO). Venezuela, Bolivia. Records: E. Alfaro 4663 (INB).

Cladium mariscus (L.) Pohl Eleocharis acicularis (L.) Roem. & Schult. Terrestrial herbs. Distribution in PILA – 1100 m, mixed Terrestrial herbs. Distribution in PILA 3200–3500 m, forest in transition to low elevation forest. Global páramo. Global distribution: USA to Ecuador, Bolivia, distribution: USA to Honduras to Panama, Peru, Greater Antilles, Lesser Antilles, Europe, Asia. Records: Venezuela, Bolivia, Argentina, Greater Antilles. Records: G. Davidse 25330 (CR, MO). A. Rodríguez 11352 (BM, INB, MO, PMA).

Eleocharis elegans (Kunth) Roem. & Schult.

Tristra (CR).

Aquatic or terrestrial herbs. Distribution in PILA Ecuador, Bolivia, Venezuela, Brazil, Uruguay, Argentina. 0–3000 m, low elevation mixed forest, mixed forest in Records : G. Davidse 26143 ( MO) , G. Davidse 29257 transition to low elevation forest, mixed forest in transition ( MO) . to cloud forest, cloud forest, low elevation oak forest, high elevation oak forest. Global distribution: Mexico to Rhynchospora hirsuta (Vahl) Vahl Argentina, Greater Antilles. Records : E. Alfaro 766 ( INB) , Aquatic or terrestrial herbs. Distribution in PILA 400– R. Quirós 93 ( INB) , S. Bridgewater 4218 ( BM, INB, MO, 400 m, low elevation mixed forest. Global distribution: PMA). Mexico to the Guyanas , Brazil, Greater Antilles, Lesser Antilles. Records: B. Gamboa 1732 ( INB) .

Mapania cuatrecasasii T.Koyama Terrestrial herbs. Distribution in PILA 600–1200 m, Rhynchospora kunthii Nees ex Kunth low elevation mixed forest, mixed forest in transition Terrestrial herbs. Distribution in PILA 2300–3000 m, low to low elevation forest. Global distribution: Costa Rica, elevation oak forest, high elevation oak forest. Global Colombia, Guyanas. Records: B. Hammel 17619 (MO), distribution: Mexico, Guatemala, Costa Rica to Colombia. G. Herrera 3282 ( INB, MO) , L.D. Gómez 23860 ( MO) . Records: E. Alfaro 4691 ( INB) , G. Davidse 28570 ( CR, MO) .

Oreobolus goeppingeri Suess. Terrestrial herbs. Distribution in PILA 2600–3500 m, low Rhynchospora locuples C.B.Clarke elevation oak forest, high elevation oak forest, páramo. Terrestrial herbs. Distribution in PILA 700–2800 m, low Global distribution: Costa Rica to Brazil. Records: G. elevation mixed forest, mixed forest in transition to low Davidse 23840 (CR, MO), G. Davidse 25310 (CR, MO), elevation forest, mixed forest in transition to cloud forest, G. Davidse 25514 (CR, MO), G. Davidse 25943 (CR, cloud forest, low elevation oak forest, high elevation oak MO), G. Davidse 26023 (MO), G. Davidse 29319 (MO). forest. Global distribution: Costa Rica to Bolivia. Records: K. Sytsma 4907 (MO), L.D. Gómez 22280 (MO).

Oreobolus venezuelensis Steyerm. Terrestrial herbs. Distribution in PILA 2300–3300 m, Rhynchospora macrochaeta Steud. ex Boeck. low elevation oak forest, páramo. Global distribution: Terrestrial herbs. Distribution in PILA 3100–3800 m, Costa Rica to Peru, Venezuela. Records: A. Chacón 564 páramo. Global distribution: Costa Rica to Bolivia. (INB, MO), E. Alfaro 3285 (INB), E. Alfaro 3377 (INB), Records: G. Davidse 26040 (CR, MO), G. Herrera 3658 E. Alfaro 3721 (INB), G. Davidse 25404 (CR, MO), G. (MO). Davidse 25867 (CR), G. Davidse 25877 (CR, MO), G. Davidse 28576 (CR, MO), G. Davidse 28992 (CR, MO), Rhynchospora oreoboloidea Gómez-Laur. G. Davidse 29272 (CR, MO), G. Davidse 29379 (MO), Terrestrial herbs. Distribution in PILA 3000–3400 m, G. Davidse 29404 (CR, MO), G. Herrera 3656 (MO), G. high elevation oak forest, páramo. Global distribution: Herrera 3697 (INB, MO), L.D. Gómez 22456 (MO). Costa Rica to Ecuador. Records: G. Davidse 23859 (CR, MO), G. Davidse 25309 (CR, MO), G. Davidse 25511

Rhynchospora cabecarae Gómez-Laur. (CR, MO) , G. Davidse 25987 (CR, MO), G. Davidse Terrestrial herbs. Distribution in PILA 2300– 2300 m, 29285 (CR, MO), G. Davidse 29378 (MO), L.D. Gómez low elevation oak forest. Global distribution: Endemic to 22459 (MO). Costa Rica. Records: M. Grayum 10354 (INB, MO). Rhynchospora polyphylla Vahl

Rhynchospora globosa (Kunth) Roem. & Schult. Terrestrial herbs. Distribution in PILA 0–2500 m, low Terrestrial herbs. Distribution in PILA 100–2000 m, low elevation mixed forest, mixed forest in transition to low elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest, elevation forest, mixed forest in transition to cloud forest. cloud forest, low elevation oak forest. Global distribution: Global distribution: Mexico to Argentina, Lesser Antilles , Mexico to Bolivia, Greater Antilles, Lesser Antilles, Greater Antilles. Records: A. Rodríguez 9833 (INB), D. Lesser Antilles. Records: A. Rodríguez 11780 (BM, INB, Santamaría 3495 (INB), D. Santamaría 3762 (INB), D. MO, PMA), A. Rodríguez 11927 (BM, INB, MO, PMA), Solano 2876 (INB), E. Alfaro 2210 (CR, INB), E. Alfaro D. Santamaría 7223 (BM, INB, MO, PMA). 4185 (INB), L. González 1218 (INB). Rhynchospora radicans (Schltdl. & Cham.) H.Pfeiff.

Rhynchospora hieronymi Boeck. Terrestrial herbs. Distribution in PILA 0–1500 m, low Terrestrial herbs. Distribution in PILA 1500–3200 m, elevation mixed forest, mixed forest in transition to low mixed forest in transition to cloud forest, cloud forest, low elevation forest, mixed forest in transition to cloud forest. elevation oak forest, páramo, sabana natural grassland. Global distribution: Mexico to Bolivia, Greater Antilles, Global distribution: Guatemala, Costa Rica, Panama, Lesser Antilles. Records: G. Herrera 5314 (MO). cloud forest, low elevation oak forest, páramo. Global

Rhynchospora rugosa (Vahl) Gale distribution: Mexico to Panama, Guyanas. Records: Terrestrial herbs. Distribution in PILA 400–2800 m, low E. Alfaro 748 (INB), G. Davidse 28450 (CR, MO), G. elevation mixed forest, mixed forest in transition to low Herrera 5962 (CR, INB), J. H. Kirkbride Jr. 160 (MO), elevation forest, mixed forest in transition to cloud forest, L. Acosta 2382 (INB), M. Alfaro 41 (INB), T. Croat 26970 cloud forest, low elevation oak forest, high elevation oak (MO). forest. Global distribution: Mexico to Argentina, Greater Antilles, Lesser Antilles. Records: D. Santamaría 3759 Rhynchospora vulcani Boeck. (INB) , D. Santamaría 3772 (INB), D. Solano 2870 (INB), Terrestrial herbs. Distribution in PILA 700–3500 m, low E. Alfaro 2207 (INB), E. Alfaro 3731 (INB), E. Alfaro elevation mixed forest, mixed forest in transition to low 4199 (INB), L. González 1197 (INB), M.A. Thomas 1548 elevation forest, mixed forest in transition to cloud forest, (MO). cloud forest, low elevation oak forest, páramo. Global distribution: Mexico to Bolivia. Records: G. Davidse

Rhynchospora ruiziana Boeck. 105427 (CR), G. Davidse 25348 (CR, MO), G. Davidse Terrestrial herbs. Distribution in PILA 0–3500 m, low 25371 (CR, MO), G. Davidse 25919 (MO), G. Davidse elevation mixed forest, mixed forest in transition to low 28889 (CR, MO), G. Davidse 29098 (CR, MO), G. elevation forest, mixed forest in transition to cloud forest, Davidse 29287 (CR, MO), G. Davidse 29329 (CR, MO), cloud forest, low elevation oak forest, páramo. Global G. Davidse 29347 (CR, MO), G. Davidse 23839 (CR, distribution: Mexico, Costa Ricato Bolivia, Venezuela, MO). Guyanas. Records: A. Chacón 588 (INB, MO), A. Rodríguez 10737 (BM, INB, MO, PMA), D. Solano Scleria distans Poir. 3995 (BM, INB, MO, PMA), E. Alfaro 762 (INB), G. Terrestrial herbs. Distribution in PILA 1000–2000 m, Davidse 23938 (CR, MO), G. Davidse 25246 (CR, MO), mixed forest in transition to low elevation forest, mixed G. Davidse 25972 (MO), G. Davidse 28602 (MO), G. forest in transition to cloud forest. Global distribution: Davidse 28650 (CR, MO), G. Davidse 28713 (CR, MO), USA to the Guyanas, Chile Brazil, the Guyanas, Argentina, G. Davidse 28988 (CR, MO), G. Herrera 6242 (CR, Greater Antilles, Lesser Antilles, Africa, Madagascar. INB), M. Grayum 10337 (CR, INB, MO). Records: A. Rodríguez 9764 (INB), D. Santamaría 3496 (INB), D. Solano 2926 (INB).

Rhynchospora schiedeana Kunth Navajuela (CR) . Scleria secans (L.) Urb. Terrestrial herbs. Distribution in PILA 1300–3400 m, Navajuela (CR). mixed forest in transition to cloud forest, cloud forest, low Terrestrial herbs. Distribution in PILA 0–1300 m, low elevation oak forest, páramo. Global distribution: Mexico elevation mixed forest, mixed forest in transition to low to Bolivia. Records: B.B. Klitgaard 838 (BM, INB, MO), elevation forest, mixed forest in transition to cloud forest. G. Davidse 23959 (CR, MO), G. Davidse 25844 (CR, Global distribution: Mexico to the Guyanas, Greater MO), G. Davidse 26201 (MO), G. Davidse 28529 (CR, Antilles, Lesser Antilles. Records: B. Hammel 17620 MO), G. Davidse 24002 (MO). (MO).