Hyptis obtusiflora C.Presl ex Benth.

MONRO, A. K., SANTAMARÍA-AGUILAR, D., GONZÁLEZ, F., CHACÓN, O., SOLANO, D., RODRÍGUEZ, A., ZAMORA, N., FEDELE, E. & CORREA, M., 2017, A first checklist to the vascular plants of La Amistad International Park (PILA), Costa Rica-Panama, Phytotaxa 322 (1), pp. 448-450 : 448-450

Hyptis obtusiflora C.Presl ex Benth. Salvia pteroura Briq.

Terrestrial herbs or shrubs. Distribution in PILA 0–1700 Terrestrial herbs or shrubs. Distribution in PILA 900–2000 m, low elevation mixed forest, mixed forest in transition m, mixed forest in transition to low elevation forest, mixed to low elevation forest, mixed forest in transition to cloud forest in transition to cloud forest. Global distribution: El forest , cloud forest. Global distribution: Belize, Nicaragua Salvador, Costa Rica, Panama. Records: A. Rodríguez to Bolivia. Records: A. Rodríguez 11372 ( BM, INB, MO, 9841 ( INB), D. Solano 2892 ( INB).

PMA), E. Alfaro 3697 (INB), L. Acosta 2393 (INB).

Scutellaria costaricana H.Wendl.

Hyptis sinuata Pohl ex Benth Pavoncillo rojo (CR).

Terrestrial herbs. Distribution in PILA 0–2000 m, low Terrestrial herbs or shrubs. Distribution in PILA elevation mixed forest, mixed forest in transition to low 0–1700 m, low elevation mixed forest, mixed forest elevation forest, mixed forest in transition to cloud forest, in transition to low elevation forest, mixed forest in cloud forest. Global distribution: Mexico to Colombia, transition to cloud forest, cloud forest. Global distribution: Brazil, Bolivia, Greater Antilles, Lesser Antilles. Records: Costa Rica, Panama. Records: A. Rodríguez 11418 (BM, A. Rodriguez 2811 (INB, MO), E. Alfaro 3696 (INB), M. INB, PMA), D. Santamaría 5909 (BM, INB, PMA), G. Alfaro 143 (INB). Gallardo 104 (INB).

Hyptis vilis Kunth & C.D.Bouché Scutellaria glabra Leonard Terrestrial herbs. Distribution in PILA 100–1600 m, Platanillo (CR). low elevation mixed forest, mixed forest in transition Terrestrial herbs or shrubs. Distribution in PILA to low elevation forest. Global distribution: Nicaragua 600–2200 m, low elevation mixed forest, mixed forest to Panama, Venezuela. Records: A. Picado 4 ( CR, INB, in transition to low elevation forest, mixed forest in MO), E. Navarro 189 ( INB) . transition to cloud forest, cloud forest, low elevation oak forest. Global distribution: Costa Rica, Panama. Records:

Lepechinia schiedeana (Schltdl.) Vatke D. Solano 4288 (BM, INB, PMA). Terrestrial herbs. Distribution in PILA 1100–3500 m, mixed forest in transition to low elevation forest, mixed Scutellaria isocheila Donn.Sm. forest in transition to cloud forest, cloud forest, low Terrestrial herbs. Distribution in PILA 1400– 1400 m, mixed elevation oak forest, páramo. Global distribution: Mexico, forest in transition to cloud forest.Global distribution: Costa Guatemala, Costa Rica to Colombia. Records: G. Davidse Rica, Panama. Records: F. González 773 ( INB, PMA) . 26012 (MO). ε-PILA Scutellaria talamancana A.Pool

Salvia alvajaca Oerst. Terrestrial herbs. Distribution in PILA 1100–1200 m, Alabaca cimarrona, Chirristate blanco (CR). mixed forest in transition to low elevation forest. Global Terrestrial herbs. Distribution in PILA 800–3200 distribution: endemic to Costa Rica. Records: M. Grayum

m, mixed forest in transition to low elevation forest, 10909 (INB, MO). mixed forest in transition to cloud forest, cloud forest, low elevation oak forest, páramo. Global distribution: Scutellaria tenuipetiolata A.Pool Nicaragua to Ecuador. Records: A. Fernández 858 (INB, Terrestrial herbs. Distribution in PILA 0–700 m, low MO), A. Rodríguez 10890 (BM, INB, PMA), D. Solano elevation mixed forest. Global distribution: Costa Rica, 4069 (BM, INB, PMA), G. Herrera 6024 (CR, INB, Panama. Records: G. Herrera 3101 (MO). MO). Stachys costaricensis Briq.

Salvia carnea Kunth Terrestrial herbs. Distribution in PILA 1000–3500 m, Terrestrial herbs. Distribution in PILA 800–3400 m, mixed mixed forest in transition to low elevation forest, mixed forest in transition to low elevation forest, mixed forest in forest in transition to cloud forest, cloud forest, low transition to cloud forest, cloud forest, low elevation oak elevation oak forest, páramo. Global distribution: Mexico forest, páramo. Global distribution: Mexico, Guatemala, to Colombia. Records: M.A. Thomas 1660 (MO). Costa Rica to Ecuador, Venezuela. Records: A.K. Monro 4053 ( BM, INB) , A.K. Monro 4101 ( BM) , A.K. Monro Stachys pittieri Briq. 5244 ( BM, INB) , B.B. Klitgaard 800 ( BM, INB) , G. Terrestrial herbs. Distribution in PILA 1400–3400 m, mixed forest in transition to cloud forest, cloud forest, low elevation oak forest, páramo. Global distribution: Endemic LEPIDOBOTRYACEAE to Costa Rica. Records: M. Grayum 10364 ( CR) . Ruptiliocarpon caracolito Hammel & N.Zamora

ε-PILA Stachys riparia A.Pool Caracolito, Cedro caracolito, Lépido (CR) . Terrestrialherbs. DistributioninPILA 1400–1800m,mixed Terrestrial trees. Distribution in PILA 0–1100 m, forest in transition to cloud forest. Global distribution: low elevation mixed forest, mixed forest in transition Endemic to Costa Rica. Records : D. Santamaría 5839 to low elevation forest. Global distribution: Nicaragua (BM, INB, PMA), D. Solano 4107 (BM, INB, PMA). to Panama, Colombia to Peru. Records: D. Santamaría, Oct. 22, 2008 (Field obs.), D. Santamaría, Oct. 25, 2008

Stachys uniflora A.Pool (Field obs.). Uses: timber. Terrestrial herbs. Distribution in PILA 1600–2800 m, mixed forest in transition to cloud forest, cloud forest, LINDERNIACEAE low elevation oak forest, high elevation oak forest. Global distribution: Costa Rica, Panama. Records: A. Rodríguez Lindernia diffusa (L.) Wettst.

10771 (BM, INB, PMA), D. Santamaría 5648 (BM, INB, Terrestrial herbs. Distribution in PILA 0–1600 m, low

PMA), G. Davidse 26154 (CR, MO). elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest,

LENTIBULARIACEAE cloud forest. Global distribution: Mexico to Brazil, Lesser Antilles, Greater Antilles. Records: M. Alfaro 2 (INB).

Utricularia amethystina Salzm. ex A.St. -Hil. & Girard Aquatic or terrestrial herbs.Distribution in PILA 500–1400 LOASACEAE m, low elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to cloud Klaprothia mentzelioides Kunth forest. Global distribution: Mexico to Honduras, Costa Terrestrial herbs. Distribution in PILA 1200–2500 m, Rica to Bolivia, Venezuela, Trinidad, Guayanas, Brazil, mixed forest in transition to low elevation forest, mixed Greater Antilles. Records: D. Santamaría 3504 (INB), D. forest in transition to cloud forest, cloud forest, low Solano 2880 (INB). elevation oak forest. Global distribution: Mexico, Costa Rica to Bolivia. Records: A. Fernández 911 (CR, INB,

Utricularia jamesoniana Oliv. MO ), A. Rodríguez 10961 (BM, INB, MO, PMA), B. Epiphytic herbs. Distribution in PILA 700–2200 m, low Hammel 5809 (MO), D. Santamaría 5726 (BM, INB, elevation mixed forest, mixed forest in transition to low MO, PMA), D. Santamaría 5782 (BM, INB, MO, PMA), elevation forest, mixed forest in transition to cloud forest, D. Solano 3993 (BM, INB, MO, PMA), G. Herrera 6009 cloud forest, low elevation oak forest. Global distribution: (CR, INB, MO), J. Bittner 1835 (CR, INB, MO), M. Guatemala, Nicaragua to Bolivia, Venezuela, Guayanas, N Grayum 10361 (CR, INB, MO). Brazil, Antilles. Records: A. Chacón 193 (CR, INB, MO), A. Chacón 279 (CR, MO), D. Santamaría 4463 (INB), G. ψ Nasa speciosa (Donn.Sm.) Weigend Herrera 3288 (INB, MO), G. Herrera 5348 (INB, MO), Terrestrial herbs to shrubs. Distribution in PILA 1900– G. Herrera 5348,1 (INB), S. Knapp 4836 (MO). 3200 m, cloud forest, low elevation oak forest, páramo. Global distribution: Costa Rica, Panama. Records: B.

Utricularia praetermissa P.Taylor Hammel 1453 (MO), F. Almeda 3508 (MO), G. Davidse Facultatively epiphytic herbs. Distribution in PILA 800– 25780 (CR, MO), G. Herrera 3669 (INB), G. Herrera 2000 m, mixed forest in transition to low elevation forest, 6249 (CR, INB, MO). Global conservation status: mixed forest in transition to cloud forest, sabana natural Endangered (EN), B1ab(i,iii). grassland. Global distribution: Nicaragua to Panama. Records: A. Soto 325 (INB), F. Solís 13 (CR, INB, MO). Nasa triphylla (Juss.) Weigend Terrestrial herbs to shrubs.Distribution in PILA1600–2600

Utricularia unifolia Ruiz & Pav. m, mixed forest in transition to cloud forest, cloud forest, Facultatively epiphytic herbs. Distribution in PILA 1000– low elevation oak forest, high elevation oak forest. Global 2500 m, mixed forest in transition to low elevation forest, distribution: Mexico, Guatemala, Nicaragua to Panama, mixed forest in transition to cloud forest, cloud forest, low Colombia, Peru, Venezuela. Records: D. Santamaría elevation oak forest. Global distribution: Costa Rica to 5780 (BM, INB, MO, PMA), D. Santamaría 7247 (BM, Bolivia, Venezuela. Records: A. Chacón 569 (INB, MO), INB, MO, PMA), D. Solano 4112 (BM, INB, MO, PMA), G. Davidse 28586 (CR, MO), G. Herrera 3707 (INB, K. E. Blum 2670 (MO), L. Acosta 2301 (INB). MO). LOGANIACEAE Panamanthus panamensis (Rizzini) Kuijt Epiphytic shrubs. Distribution in PILA 1200–2500 m,

mixed forest in transition to low elevation forest, mixed

Spigelia scabra Cham. & Schltdl. forest in transition to cloud forest, cloud forest, low

Terrestrial herbs.Distribution in PILA 1500–1800m,mixed elevation oak forest. Global distribution: Costa Rica,

forest in transition to cloud forest. Global distribution: Panama. Records: F. Almeda 3536 (MO).

Mexico to Argentina. Records: D. Santamaría 7098 (BM,

INB, MO, PMA), G. Herrera 3090 (MO). Psittacanthus rhynchanthus (Benth.) Kuijt

Epiphytic shrubs. Distribution in PILA 0–1800 m, low

LORANTHACEAE elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest,

ψ Gaiadendron punctatum (Ruiz & Pav.) G.Don cloud forest. Global distribution: Mexico to Venezuela.

Matapalo (CR). Records: L. Acosta 2318 (INB).

Shrubs or facultatively epiphytic trees. Distribution in PILA 0–3800 m, low elevation mixed forest, mixed Psittacanthus scheryi Woodson

forest in transition to low elevation forest, mixed forest in Epiphytic shrubs. Distribution in PILA 1400–2500 m,

transition to cloud forest, cloud forest, low elevation oak mixed forest in transition to cloud forest, cloud forest,

forest, páramo. Global distribution: Mexico to Bolivia. low elevation oak forest. Global distribution: Costa Rica,

Records: A. Chacón 582 (INB, MO), A. Fernández 1053 Panama. Records: A.K. Monro 4483 (BM, INB), A.K.

(INB), A. Fernández 684 (CR, INB, MO), A. Fernández Monro 4936 (BM, INB, MO), M. Chinchilla 192 (CR,

922 (CR, INB), A.K. Monro 4097 (BM, INB, MO), A.K. INB).

Monro 4307 (BM, INB, MO), A.K. Monro 5176 (BM,

INB, MO), A.K. Monro 5187 (BM, INB, MO), A.K. Psittacanthus schiedeanus (Schltdl. & Cham.) G.Don

Monro 5234 (BM, INB, MO), A. Soto 322 (INB), B.B. Matapalo (CR).

Klitgaard 799 (BM, INB, MO), B. Gamboa 1237 (INB), Epiphytic shrubs. Distribution in PILA 1200–3400 m,

B. Gamboa 1723 (INB, MO), B. Gamboa 585 (CR, INB, mixed forest in transition to low elevation forest, mixed

MO), D. Solano 5223 (BM, INB, MO, PMA), E. Alfaro forest in transition to cloud forest, cloud forest, low

3289 (INB), E. Alfaro 4696 (INB), G. Davidse 23866 (CR, elevation oak forest, páramo. Global distribution: Mexico

MO), G. Davidse 25379 (CR, MO), G. Davidse 25854 to Panama. Records: A.K. Monro 4496 (BM, INB), D.

(CR, MO), G. Davidse 25899 (MO), G. Davidse 28756 Santamaría 5899 (BM, INB, MO, PMA), G. Herrera

(CR, MO), G. Herrera 6110 (CR, INB, MO), J. Bittner 6039 (CR, INB, MO), R. J. Schmalzel 1348 (MO).

1810 (INB, MO), F. Quesada 1835 (INB), F. Quesada

1947 (INB), L. Angulo 240 (CR, INB), L. González 2487 Struthanthus burgeri Kuijt

(INB), L. González 3635 (INB), M. Grayum 10346 (CR, Epiphytic shrubs. Distribution in PILA 800– 800 m,

INB, MO), M. Moraga 395 (INB), M. Moraga 429 (CR, mixed forest in transition to low elevation forest. Global

INB), R. Aguilar 1109 (INB, MO), R. Aguilar 1119 (INB), distribution: Endemic to Costa Rica. Records: G. Herrera

S. Knapp 9974 (BM, INB, MO). Regional conservation 3326 (INB, MO).

status: Near Threatened (NT). Struthanthus cansjerifolius (Oliv.) Eichler

Oryctanthus occidentalis (L.) Eichler Epiphytic shrubs.Distribution in PILA 800–2200 m, mixed

Epiphytic shrubs. Distribution in PILA 0–1900 m, low forest in transition to low elevation forest, mixed forest

elevation mixed forest, mixed forest in transition to low in transition to cloud forest, cloud forest, low elevation

elevation forest, mixed forest in transition to cloud forest, oak forest. Global distribution: El Salvador, Costa Rica

cloud forest. Global distribution: Nicaragua to Bolivia, to Panama. Records: A.K. Monro 5611 (BM, INB, MO,

Greater Antilles. Records: D. Solano 5490 (PMA), G. PMA), A. Rodríguez 11919 (BM, INB, MO, PMA), D.

Herrera 5209 (CR, INB, MO). Solano 4057 (BM, INB, MO, PMA), G. Herrera 6096 (CR, INB), N. Zamora 4097 (BM, INB, MO, PMA).

Oryctanthus spicatus (Jacq.) Eichler

Epiphytic shrubs. Distribution in PILA 200–2000 m, Struthanthus hartwegii (Benth.) Standl.

low elevation mixed forest, mixed forest in transition to Epiphytic shrubs. Distribution in PILA 1300–2300 m,

low elevation forest, mixed forest in transition to cloud mixed forest in transition to cloud forest, cloud forest,

forest. Global distribution: Belize, Guatemala, Costa low elevation oak forest. Global distribution: Mexico to

Rica, Venezuela to Bolivia. Records: A.K. Monro 5991 Honduras, Costa Rica, Panama. Records: R. Villalobos

(PMA), D. Santamaría 7473 (PMA), D. Santamaría 7597 218 (CR, INB).

(PMA). Struthanthus leptostachyus (Kunth) G.Don forest. Global distribution: Costa Rica to Venezuela. Epiphytic shrubs. Distribution in PILA 0–2200 m, low Records: M. Correa 3625 (MO). elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest, MALPIGHIACEAE cloud forest, low elevation oak forest. Global distribution: Costa Rica to Peru. Records: A. Rodríguez 11887 (BM, □ Bunchosia costaricensis Rose INB, MO, PMA ), A. Soto 1256 (INB), D. Solano 3208 Terrestrial shrubs or trees. Distribution in PILA 700–1200 (INB), J. Corrales 30 (INB), F. Quesada 1883 (INB), R. m, low elevation mixed forest, mixed forest in transition Aguilar 1120 (INB). to low elevation forest. Global distribution: Costa Rica to Panama. Records: A. De Sedas 606 (INB, PMA), D.

Struthanthus quercicola (Schltdl. & Cham.) D.Don Santamaría 6450 (BM, INB, MO, PMA),. Epiphytic shrubs. Distribution in PILA 0–3400 m, low elevation mixed forest, mixed forest in transition to low Bunchosia macrophylla Rose elevation forest, mixed forest in transition to cloud forest, Muñeco (CR). cloud forest, low elevation oak forest, páramo. Global Terrestrial shrubs or trees. Distribution in PILA 0–900 distribution: Mexico to Panama. Records: A. Chacón 587 m, low elevation mixed forest, mixed forest in transition (INB, MO), A. Fernández 1855 (INB), A.K. Monro 4155 to low elevation forest. Global distribution: Nicaragua to (BM), A.K. Monro 5287 (BM, INB, MO), D. Santamaría Colombia. Records: D. Santamaría 7259 (BM, INB, MO, 4426 (INB), D. Santamaría 5755 (BM, INB, MO, PMA), PMA), D. Solano 5198 (BM, INB, MO, PMA). E. Alfaro 4639 (INB), E. Alfaro 4686 (INB), E. Alfaro 734 (INB), G. Davidse 25471 (MO), G. Davidse 25748 (CR, Bunchosia polystachia (Andrews) DC. MO ), G. Davidse 25753 (CR, MO), G. Davidse 29026 Terrestrial shrubs or trees. Distribution in PILA 800–800 (CR, MO), G. Davidse 29124 (CR, MO), J. Bittner 1812 m, mixed forest in transition to low elevation forest.Global (INB), F. Quesada 1617 (INB), F. Quesada 1945 (INB), distribution: Belize, El Salvador to Costa Rica, French M. Alfaro 37 (INB), M. Grayum 10330 (CR, INB, MO), Guyana, Greater Antilles, Lesser Antilles. Records: A. M. Moraga 402 (INB), S. Knapp 10007 (BM, INB), S. Fernández 833 (CR, INB). Knapp 9910 (BM, INB). Bunchosia ternata Dobson

LYTHRACEAE Terrestrial shrubs or trees. Distribution in PILA 1100– 2000 m, mixed forest in transition to low elevation

Cuphea appendiculata Benth. forest, mixed forest in transition to cloud forest. Global Terrestrial shrubs. Distribution in PILA 300–2700 m, low distribution: Costa Rica, Panama. Records: A.K. Monro elevation mixed forest, mixed forest in transition to low 5585 (BM, INB, MO, PMA), A. Rodríguez 10809 (BM, elevation forest, mixed forest in transition to cloud forest, INB, MO, PMA), D. Solano 3949 (BM, INB, MO, PMA), cloud forest, low elevation oak forest, high elevation oak G. Herrera 5878 (CR, INB). forest. Global distribution: Mexico to Panama. Records: A. Maroto 202 (INB, MO), A. Rodríguez 11362 (BM, Byrsonima arthropoda A.Juss. INB, MO, PMA ), C. Hamilton 3654 (MO), C. Hamilton Terrestrial shrubs or trees. Distribution in PILA 0–1000 719 (MO), C. Hamilton 745 (MO), L. Acosta 2394 (INB), m, low elevation mixed forest, mixed forest in transition S. Knapp 1516 (MO). to low elevation forest. Global distribution: Nicaragua to Brazil. Records: G. Herrera 3340 (INB, MO).

Cuphea carthagenensis (Jacq.) J.F.Macbr. Santalucía, Nicaragüita (CR) . Byrsonima crassifolia (L.) Kunth Terrestrial herbs or shrubs. Distribution in PILA Nance, Nancite, Tsiki, Bek (Cabécar), Si+krá (Brunka), 0–3200 m, low elevation mixed forest, mixed forest in Skír-ko (Térraba) (CR), Nance (P). transition to low elevation forest, mixed forest in transition Terrestrial shrubs or trees. Distribution in PILA to cloud forest, cloud forest, low elevation oak forest, 0–1500 m, low elevation mixed forest, mixed forest páramo. Global distribution: USA to Argentina, Greater in transition to low elevation forest, mixed forest in Antilles, Lesser Antilles, Pacific Islands . Records: C. transition to cloud forest. Global distribution: Mexico Hamilton 697 (MO), L. Acosta 2315 (INB). to Brazil, Greater Antilles, Lesser Antilles. Records: D. Solano 2878 (INB). Uses: colouring, Edible, biological

Cuphea epilobiifolia Koehne control, timber, medicinal, Textil. Terrestrial herbs or shrubs. Distribution in PILA 0–1300 m, low elevation mixed forest, mixed forest in transition ψ Byrsonima herrerae W.R.Anderson to low elevation forest, mixed forest in transition to cloud Nance, Nancite, Tsiki (CR). Terrestrial trees. Distribution in PILA 500–1700 m, low elevation forest, mixed forest in transition to cloud low elevation mixed forest, mixed forest in transition forest, cloud forest, low elevation oak forest, sabana to low elevation forest, mixed forest in transition to natural grassland.Global distribution: Mexico to Paraguay, cloud forest. Global distribution: Costa Rica, Panama. Lesser Antilles. Records: L. González 3587 (INB). Records: D. Santamaría 6716 (BM, INB, MO, PMA), G. Herrera 2709 (MO), G. Herrera 3333 (INB, MO). Global ψ Heliocarpus appendiculatus Turcz. conservation status: Endangered (EN), B1ab(iii). Burío, Burillo, Buría, Burío patón, Buriogre, Tsa-ri (Cabécar), Stsa (Bribrí), Gsii+krá (Brunka), Rús-ura+gró

Heteropterys panamensis Cuatrec. & Croat (Térraba) (CR). Terrestrial lianas. Distribution in PILA 0–1300 m, low Terrestrial trees. Distribution in PILA 0–2000 m, low elevation mixed forest, mixed forest in transition to low elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest. elevation forest, mixed forest in transition to cloud forest, Global distribution: Mexico to Panama. Records : A. cloud forest. Global distribution: Mexico to Panama. Rodríguez 11294 (BM, INB, MO, PMA). Records: A. Rodríguez, Jul. 21, 2007 (Field obs.), A. Rodríguez, Jul. 25, 2007 (Field obs.), N. Zamora, Aug. 1,

Hiraea fagifolia (DC.) A.Juss. 2008 (Field obs.), N. Zamora, Jul. 29, 2008 (Field obs.). Terrestrial lianas. Distribution in PILA 0–1600 m, low Uses : handicrafts, cellulose. Regional conservation status: elevation mixed forest, mixed forest in transition to low Least Concern (LC). elevation forest, mixed forest in transition to cloud forest, cloud forest. Global distribution: Mexico to Argentina, □ Malvaviscus achanioides (Turcz.) Fryxell Lesser Antilles. Records : D. Santamaría 6244 (BM, INB, Terrestrial shrubs. Distribution in PILA 300–2200 m, low MO, PMA). elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest,

Hiraea smilacina Standl. cloud forest, low elevation oak forest. Global distribution: Terrestrial lianas. Distribution in PILA 0–1900 m, low Mexico to Honduras, Costa Rica, Panamá. Records: A. elevation mixed forest, mixed forest in transition to low Rodríguez 10979 (BM, INB, MO, PMA), D. Santamaría elevation forest, mixed forest in transition to cloud forest, 7265 (BM, INB, MO, PMA), D. Solano 4565 (BM, INB, cloud forest. Global distribution: Mexico to Honduras, MO, PMA), D. Solano 5257 (BM, INB, MO, PMA), Costa Rica to Ecuador. Records: E. Alfaro 2186 (CR, E. Alfaro 754 (INB), G. Davidse 28418 (CR, MO), N. INB). Zamora 4173 (BM, INB, MO, PMA).