Guarea bullata Radlk.

MONRO, A. K., SANTAMARÍA-AGUILAR, D., GONZÁLEZ, F., CHACÓN, O., SOLANO, D., RODRÍGUEZ, A., ZAMORA, N., FEDELE, E. & CORREA, M., 2017, A first checklist to the vascular plants of La Amistad International Park (PILA), Costa Rica-Panama, Phytotaxa 322 (1), pp. 448-450 : 448-450

Guarea bullata Radlk. Guarea macrocalyx Al.Rodr.

Terrestrial shrubs or trees. Distribution in PILA 0–1200 Terrestrial shrubs. Distribution in PILA 0–700 m, low m, low elevation mixed forest, mixed forest in transition elevation mixed forest. Global distribution: Endemic to to low elevation forest. Global distribution: Mexico, Costa Rica. Records: D. Santamaría 6001 ( BM, INB, Guatemala, Nicaragua, Costa Rica, Panamá. Records: A. MO, PMA), D. Solano 4276 ( BM, INB, MO, PMA) , D. Rodríguez, Jul. 22, 2007 ( Field obs.). Solano 4309 ( BM, INB, MO, PMA) .

Guarea chiricana Standl. ψ Guarea rhopalocarpa Radlk. Terrestrial shrubs. Distribution in PILA 0–1300 m, low Cacao de ardilla, Cocora, Pocora, Ocora (CR). elevation mixed forest, mixed forest in transition to low Terrestrial shrubs or trees. Distribution in PILA elevation forest, mixed forest in transition to cloud forest. 0–1500 m, low elevation mixed forest, mixed forest Global distribution: Nicaragua, Costa Rica. Records: A. in transition to low elevation forest, mixed forest in Rodríguez 11336 (BM, INB, MO, PMA), D. Santamaría transition to cloud forest. Global distribution: Nicaragua 6722 (BM, INB, MO, PMA). to Colombia. Records: G. Herrera 3278 (INB, MO). Uses: timber. Regional conservation status: Vulnerable

Guarea glabra Vahl (VU) , B1ab(i,iii). Terrestrial shrubs or trees. Distribution in PILA 0–1800 m, low elevation mixed forest, mixed forest in transition Guarea tonduzii C.DC. to low elevation forest, mixed forest in transition to Terrestrial trees. Distribution in PILA 300–1600 m, low cloud forest, cloud forest. Global distribution: Mexico, elevation mixed forest, mixed forest in transition to low Honduras to Panama. Records: Acosta 2320 (INB), elevation forest. Global distribution: Mexico to Honduras, Azofeifa 148 (INB), González 453 (INB), González 480 Costa Rica, Panama. Records: E. Alfaro 5557 (BM, (INB), Herrera 5927 (CR, INB, MO), Mora 302 (INB), INB, MO), Rodríguez 11455 (BM, INB, MO, PMA), D. D. Solano 4876 (BM, INB, MO, PMA). Santamaría 6532 (BM, INB, MO, PMA), D. Solano 4695 (BM, INB, MO, PMA), N. Zamora, Aug. 4, 2008 (Field

Guarea grandifolia DC. obs.). Caobilla , Uña de gato, Ocora , Pocora , Pochote de bajura, Cedro macho, Cortez negro, Cedro cóbano, Cola de pavo ψ Ruagea glabra Triana & Planch. (CR) , Chuchupate , Cedro macho, Tres bocas (P). Caoba, Cedro cóbano (CR). Terrestrial shrubs or trees. Distribution in PILA Terrestrial shrubs or trees. Distribution in PILA 0–1200 m, low elevation mixed forest, mixed forest in 300–2200 m, low elevation mixed forest, mixed forest transition to low elevation forest. Global distribution: in transition to low elevation forest, mixed forest in Nicaragua to Bolivia,, Venezuela, Trinidad, Guayanas , transition to cloud forest, cloud forest, low elevation oak Brazil, Paraguay. Argentina, Greater Antilles. Records: forest, pasture. Global distribution: Costa Rica to Bolivia. A. Rodríguez, Sept 29, 2007 (Field obs.), D. Santamaría, Records: A. De Sedas 681 ( PMA) , A.K. Monro 4865 Oct. 24, 2007 ( Field obs.), D. Solano, Oct. 24, 2007 (Field ( BM, INB, MO), A.K. Monro 5015 ( BM, INB, MO), A. obs.), D. Solano, Oct. 27, 2007 (Field obs.), D. Solano, Rodríguez 11593 ( BM, INB, MO, PMA) , D. Santamaría Oct. 28, 2007 (Field obs.). Uses: timber. 5722 ( BM, INB, MO, PMA) , D. Santamaría 7470 ( PMA) , D. Santamaría 7487 ( PMA) , G. Herrera 5505 (CR, INB,

Guarea kegelii Turcz. MO ), J. Lezcano 338 (PMA), R. Aguilar 1130 (INB, MO), Terrestrial shrubs or trees. Distribution in PILA 100–1400 S. Bridgewater 4146 (BM, INB, MO, PMA), T. Croat m, low elevation mixed forest, mixed forest in transition 26972 (MO). to low elevation forest, mixed forest in transition to cloud forest. Global distribution: Mexico to Panama. Records: ψ Trichilia havanensis Jacq. A. Rodríguez 11187 (BM, INB, MO, PMA). Uruca, Caracolillo, Cedro cóbano, Ocora (CR). Terrestrial trees. Distribution in PILA 500–2600 m,

ψ Guarea kunthiana A.Juss. low elevation mixed forest, mixed forest in transition to Cocora, Ocora (CR). low elevation forest, mixed forest in transition to cloud Terrestrial trees. Distribution in PILA 0–2200 m, low forest, cloud forest, low elevation oak forest, high elevation elevation mixed forest, mixed forest in transition to low oak forest. Global distribution: Mexico to Venezuela, elevation forest, mixed forest in transition to cloud forest, Greater Antilles, Lesser Antilles. Records: A. Chacón 574 cloud forest, low elevation oak forest, pasture. Global (INB, MO). Uses: biological control, timber ( Guatemala), distribution: Nicaragua to Paraguay. Records: D. Solano medicinal, ornamental. Regional conservation status: 4158 (BM, INB, MO, PMA). Uses: timber. Regional Least Concern (LC). conservation status: Near Threatened (NT).

Trichilia martiana C.DC. low elevation mixed forest, mixed forest in transition Cedrillo, Cola de pava, Manteco, Guacatillo, Jocotillo, to low elevation forest, mixed forest in transition to Limoncillo (CR). cloud forest, cloud forest. Global distribution: USA to Terrestrial shrubs or trees. Distribution in PILA Argentina, Lesser Antilles , Africa, Madagascar Asia, 0–2000 m, low elevation mixed forest, mixed forest in Australia. Records: A. Soto 1248 (INB). Uses: medicinal. transition to low elevation forest, mixed forest in transition to cloud forest, cloud forest. Global distribution: Mexico Cissampelos tropaeolifolia DC. to Venezuela and Suriname, Brazil, Greater Antilles, Terrestrial twiners. Distribution in PILA 0–2100 m, low Lesser Antilles. Records: D. Solano 5047 (BM, INB, MO, elevation mixed forest, mixed forest in transition to low PMA), G. Davidse 28454 (CR, MO). Uses: medicinal. elevation forest, mixed forest in transition to cloud forest,

cloud forest, low elevation oak forest. Global distribution: Trichilia pallida Sw. Mexico to Paraguay. Records: J. González 814 (CR, INB, (CR), Terciopelo, Conejito colorado (P). MO). Terrestrial shrubs or trees. Distribution in PILA 0–1500 m, low elevation mixed forest, mixed forest in Hyperbaena smilacina Standl. transition to low elevation forest, mixed forest in transition Terrestrial lianas. Distribution in PILA 600–1500 m, low to cloud forest. Global distribution: Mexico to Paraguay, elevation mixed forest, mixed forest in transition to low Greater Antilles, Lesser Antilles. Records: A. Rodríguez elevation forest, mixed forest in transition to cloud forest. 11551 (BM, INB, MO, PMA), A. Rodríguez 11641 (BM, Global distribution: Nicaragua, Costa Rica, Colombia. INB, MO, PMA). Records: D. Santamaría 6185 (BM, INB, MO, PMA).

□ Trichilia pittieri C.DC.

Cedro cóbano (CR). METTENIUSSACEAE

Terrestrial shrubs or trees. Distribution in PILA

700–1200 m, low elevation mixed forest, mixed forest ψ Calatola costaricensis Standl.

in transition to low elevation forest. Global distribution: Azulillo, Cucaracho, Duraznillo, Erepe, Guaitil, Icacón, Costa Rica Panama, Ecuador,, Peru. Records: N. Zamora, Palo azul, Palo de papa, Tapa culo, Coquito, Azul, Aug. 8, 2008 (Field obs.). Azulillo, Erepe, Duraznillo ( CR), Jaguey (P) .

Terrestrial shrubs or trees. Distribution in PILA MENISPERMACEAE 0–2400 m, low elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition Abuta panamensis (Standl.) Krukoff & Barneby to cloud forest, cloud forest, low elevation oak forest,

pasture. Global distribution: Mexico, Belize, Honduras Terrestrial lianas. Distribution in PILA 0–1800 m, low

to Bolivia, Venezuela. Records: A.K. Monro 4461 (BM, elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest, MO), E. Navarro 688 (INB, MO), F. González 468 (INB),

J. González 803 (CR, MO). Uses: food, timber. Regional cloud forest. Global distribution: Mexico to Panama.

conservation status: Near Threatened

Records: A.K. Monro 6012 (PMA), L. Acosta 2309

(INB).

ψ Metteniusa tessmanniana (Sleumer) Sleumer Anomospermum reticulatum (Mart.) Eichler Terrestrial shrubs or trees. Distribution in PILA 100–1000 Manguito (CR). m, low elevation mixed forest, mixed forest in transition to low elevation forest. Global distribution: Costa Rica Terrestrial lianas. Distribution in PILA 200–800 m,

to Peru. Records: D. Santamaría 6840 (BM, INB, MO, low elevation mixed forest, mixed forest in transition to low elevation forest. Global distribution: Nicaragua to PMA). Regional conservation status: Vulnerable (VU),

B1ab(i,iii).

Bolivia. Records: G. Herrera 3119 (MO).

Cissampelos andromorpha DC. MORACEAE

Terrestrial twiners. Distribution in PILA 0–1000 m,

Castilla elastica Sessé

low elevation mixed forest, mixed forest in transition

Hule, Ule, Palo de hule, Tseni (Cabécar), Sini, Tsini to low elevation forest. Global distribution: Guatemala,

(Bribrí) (CR), Caucho, Mastate blanco, Hule (P).

Nicaragua to Bolivia, Venezuela, the Guyanas, Brazil.

Terrestrial tree. Distribution in PILA 0–100 m, low Records: G. Herrera 5293 (CR, INB, MO).

elevation mixed forest. Global distribution: Mexico to Cissampelos pareira L. Ecuador. Records: A. Rodríguez, Jul. 21, 2007 (Field obs.), A. Rodríguez, Jul. 22, 2007 (Field obs.) .

Bejuco azul, Venadera, Venadero (CR).

Terrestrial twiners. Distribution in PILA 0–2000 m, Clarisia biflora Ruiz & Pav. low elevation forest, mixed forest in transition to cloud Lechillo, Ojoche, Ojoche negro, Ojochillo, Quihra (CR). forest, cloud forest, low elevation oak forest. Global Terrestrial trees. Distribution in PILA 0–1300 m, distribution: Costa Rica to Peru, Venezuela. Records: M. low elevation mixed forest, mixed forest in transition to Moraga 589 (INB). Regional conservation status: Near low elevation forest, mixed forest in transition to cloud Threatened (NT). forest. Global distribution: Mexico to Brazil. Records: D. Santamaría 6180 (BM, INB, MO, PMA). Ficus citrifolia Mill. Higo, Higuerón (CR) .

Dorstenia choconiana S.Watson Shrubs or epiphytic trees. Distribution in PILA Terrestrial herbs. Distribution in PILA 0–1700 m, low 0–1700 m, low elevation mixed forest, mixed forest in elevation mixed forest, mixed forest in transition to low transition to low elevation forest, mixed forest in transition elevation forest, mixed forest in transition to cloud forest, to cloud forest, cloud forest. Global distribution: USA to cloud forest, dense thicket. Global distribution: Guatemala the Guyanas, Greater Antilles, Lesser Antilles. Records: to Panama. Records: A. Rodríguez 11781 (BM, INB, MO, J. González 784 (CR, INB, MO), J. González 821 (CR, PMA), D. Solano 5016 (BM, INB, MO, PMA), D. Solano INB, MO). 5038 (BM, INB, MO, PMA). Ficus crassiuscula Warb. ex Standl.

Dorstenia contrajerva L. Higuerón (CR) . Contrayerba, Mano de sapo (CR). Terrestrial shrubs or trees. Distribution in PILA Terrestrial herbs. Distribution in PILA 0–1700 m, low 300–2000 m, low elevation mixed forest, mixed forest in elevation mixed forest, mixed forest in transition to low transition to low elevation forest, mixed forest in transition elevation forest, mixed forest in transition to cloud forest, to cloud forest. Global distribution: Guatemala to Peru, cloud forest. Global distribution: Mexico to Venezuela, Venezuela. Records: A. Fernández 1067 (CR, INB, MO), Greater Antilles, Lesser Antilles. Records: D. Solano C. von Hagen 2095 (MO), G. Herrera 6112 (CR, INB, 4375 (BM, INB, MO, PMA), D. Solano 4558 (BM, INB, MO). MO, PMA). Uses: medicinal. Ficus crassivenosa W.C.Burger

Ficus americana Aubl. Shrubs or epiphytic trees. Distribution in PILA 0–500 m, Higuito, Higuerón (CR). low elevation mixed forest. Global distribution: Nicaragua Shrubs or epiphytic trees. Distribution in PILA to Colombia. Records: A. Rodríguez 11028 (BM, INB, 0–3800 m, low elevation mixed forest, mixed forest in MO, PMA). transition to low elevation forest, mixed forest in transition to cloud forest, cloud forest, low elevation oak forest, Ficus macbridei Standl. páramo, dense thicket. Global distribution: Mexico to the Terrestrial trees. Distribution in PILA 500–2000 m, low Guyanas, Greater Antilles, Lesser Antilles. Records: A.K. elevation mixed forest, mixed forest in transition to low Monro 5074 (BM, INB), D. Santamaría 5953 (BM, INB, elevation forest, mixed forest in transition to cloud forest. MO, PMA). Global distribution: Nicaragua to Bolivia. Records: F. Quesada 1688 (CR, INB, MO), F. Quesada 1896 (INB).

Ficus cahuitensis C.C.Berg Hemi-epiphytic tree. Distribution in PILA 0–1200 m, low Ficus obtusifolia Kunth elevation mixed forest, mixed forest in transition to low Higuerón, Capulamate, Matapalo, Palo de agua (CR). elevation forest. Global distribution: Belize, Honduras, Hemi-epiphytic tree. Distribution in PILA 0–1100 Costa Rica, Panama. Records: L.D. Gómez 23832 (INB). m, low elevation mixed forest, mixed forest in transition to low elevation forest. Global distribution: Mexico to

ψ Ficus caldasiana Dugand Bolivia. Records: A. Soto 1251 (INB). Terrestrial shrubs. Distribution in PILA 500–1700 m, low elevation mixed forest, mixed forest in transition to low Ficus pertusa L.f. elevation forest, mixed forest in transition to cloud forest. Higuerón, Higuito (CR). Global distribution: Mexico to Ecuador, Venezuela. Shrubs or epiphytic trees. Distribution in PILA 0–2300 Records: N. Zamora, Aug.1, 2008 (Field obs.), N. Zamora, m, low elevation mixed forest, mixed forest in transition Jul. 29, 2008 (Field obs.). Regional conservation status: to low elevation forest, mixed forest in transition to cloud Vulnerable (VU), B1ab(i,iii). forest, cloud forest, low elevation oak forest. Global distribution: Mexico to the Guyanas, Greater Antilles.

ψ Ficus cervantesiana Standl. & L.O.Williams Records : D. Santamaría 7216 (BM, INB, MO, PMA), D. Hemi-epiphytic tree. Distribution in PILA 700–2100 m, Solano 5435 (PMA), J. Dwyer 6954 (MO). low elevation mixed forest, mixed forest in transition to

Ficus schippii Standl. to low elevation forest, mixed forest in transition to Hemi-epiphytic shrubs. Distribution in PILA 0–600 m, cloud forest. Global distribution: Costa Rica to Ecuador. low elevation mixed forest. Global distribution: Belize, Records: D. Solano 4727 (BM, INB, MO, PMA), F. Nicaragua to Peru, Venezuela. Records: G. Herrera 3113 González 452 (INB), J. González 825 (CR, INB, MO).

(MO).

ψ Naucleopsis naga Pittier

ψ Ficus tonduzii Standl. Amargo, Naga, Quina (CR) .

Batsu, Higuerón, Chilamate (CR). Terrestrial trees. Distribution in PILA 0–1700 m, low Terrestrial trees. Distribution in PILA 0–1000 m, elevation mixed forest, mixed forest in transition to low low elevation mixed forest, mixed forest in transition to elevation forest, mixed forest in transition to cloud forest, low elevation forest,, dense thicket. Global distribution: cloud forest, dense thicket. Global distribution: Honduras Guatemala to Bolivia, Venezuela. Records: D. Solano to Ecuador. Records: G. Herrera 2674 (INB, MO). Uses: 4563 (BM, INB, MO, PMA), E. Navarro 212 (MO). timber. Regional conservation status: Near Threatened Regional conservation status: Near Threatened (NT). (NT).

ψ Ficus tuerckheimii Standl. Naucleopsis ulei (Warb.) Ducke

Higuerón (CR). Quina, Amargo (CR). Hemi-epiphytic tree. Distribution in PILA 1200–1800 Terrestrial trees. Distribution in PILA 0–400 m, low

m, mixed forest in transition to low elevation forest, mixed elevation mixed forest. Global distribution: Costa Rica to forest in transition to cloud forest. Global distribution: Bolivia. Records: A. Chacón 8 (INB, MO).

Mexico to Panama. Records: J. González 829 (CR, INB,

MO), M. Chinchilla 182 (INB). Regional conservation Perebea hispidula Standl.

status: Near Threatened ( NT). Terrestrial shrubs or trees. Distribution in PILA 0–700 m, low elevation mixed forest. Global distribution: Nicaragua

ψ Ficus velutina Humb. & Bonpl. ex Willd. to Peru. Records: A. Rodríguez 11743 (BM, INB, MO,

Amate, Higuerón, Matapalo, Barrú, Barú (CR). PMA), A. Rodríguez 11759 (BM, INB, MO, PMA). Hemi-epiphytic tree. Distribution in PILA 700–1900

m, low elevation mixed forest, mixed forest in transition Poulsenia armata (Miq.) Standl.

to low elevation forest, mixed forest in transition to cloud Calugo, Chilamate espina, Mastate, Calabacillo de playa,

forest. Global distribution: Mexico to Ecuador, Venezuela. Chilamate espinoso, Chilamate (CR), Cocua, Cucua,

Records: L. Acosta 2283 (INB). Regional conservation Mastate (P).

status: Near Threatened (NT). Terrestrial trees. Distribution in PILA 0–1100 m, low elevation mixed forest, mixed forest in transition to low

ψ Helicostylis tovarensis (Klotzsch & H.Karst.) C.C.Berg elevation forest. Global distribution: Mexico, Guatemala,

Terrestrial shrubs or trees. Distribution in PILA 500–1500 Costa Rica to Bolivia. Records: D. Santamaría 6020 (BM,

m, low elevation mixed forest, mixed forest in transition INB, MO, PMA). Uses: timber.

to low elevation forest, mixed forest in transition to cloud forest, pasture. Global distribution: Costa Rica to ψ Pseudolmedia glabrata (Liebm.) C.C.Berg

Bolivia, Venezuela. Records: A. De Sedas 699 (PMA), Terrestrial trees. Distribution in PILA 200–1800 m, low

A.K. Monro 5013 (BM, INB), D. Santamaría 5939 (BM, elevation mixed forest, mixed forest in transition to low

INB, PMA), D. Santamaría 6679 (BM, INB, PMA), G. elevation forest, mixed forest in transition to cloud forest.

Herrera 2697 (INB, MO). Regional conservation status: Global distribution: Guatemala to Costa Rica. Records:

Near Threatened (NT). L. Acosta 2351 (INB). Regional conservation status: Near Threatened (NT).

℮ Morus alba L.

Morera (CR). ψ Pseudolmedia spuria (Sw.) Griseb. Terrestrial shrubs. Distribution in PILA 1600–2500 m, Casanegro, Guapinolillo, Guiubra, Ojochillo, Ojoche mixed forest in transition to cloud forest, cloud forest, low colorado (CR).

elevation oak forest. Global distribution: USA, Bolivia, Terrestrial trees. Distribution in PILA 0–1900 m, low

Guyanas, Madagascar, Phillipines. Records: M. Moraga elevation mixed forest, mixed forest in transition to low

417 (CR, INB). Uses: food, medicinal. elevation forest, mixed forest in transition to cloud forest, cloud forest. Global distribution: Guatemala to Costa

Naucleopsis capirensis C.C.Berg Rica. Records : N. Zamora, Aug. 4, 2008 (Field obs.), N.

Terrestrial shrubs or trees. Distribution in PILA 700–1700 Zamora, Jul. 30, 2008 (Field obs.). Regional conservation

m, low elevation mixed forest, mixed forest in transition status: Near Threatened (NT).