Palmorchis trilobulata L.O.Williams

MONRO, A. K., SANTAMARÍA-AGUILAR, D., GONZÁLEZ, F., CHACÓN, O., SOLANO, D., RODRÍGUEZ, A., ZAMORA, N., FEDELE, E. & CORREA, M., 2017, A first checklist to the vascular plants of La Amistad International Park (PILA), Costa Rica-Panama, Phytotaxa 322 (1), pp. 448-450 : 448-450

Palmorchis trilobulata L.O.Williams Pleurothallis longipedicellata Ames & C.Schweinf.

Terrestrial herbs. Distribution in PILA 0–2000 m, low Epiphytic herbs. Distribution in PILA 600–2500 m, low elevation mixed forest, mixed forest in transition to low elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest, elevation forest, mixed forest in transition to cloud forest, cloud forest. Global distribution: Nicaragua to Panama cloud forest, low elevation oak forest. Global distribution: (possibly South America). Records: D. Solano 4894 ( BM, Costa Rica, Panama, Ecuador. Records: E. Alfaro 3461 INB, MO, PMA), L.D. Gómez 23093 ( MO) . ( INB) .

Platystele compacta (Ames) Ames Pleurothallis nitida Luer Epiphytic herbs. Distribution in PILA 200–2500 m, low Epiphytic herbs. Distribution in PILA 1700– 1700 m, elevation mixed forest, mixed forest in transition to low cloud forest. Global distribution: Costa Rica, Panama. elevation forest, mixed forest in transition to cloud forest, Records: Luer 1337 ( SEL). cloud forest, low elevation oak forest. Global distribution: Mexico to Panama. Records: D. Solano 4778 ( BM, INB, Pleurothallis peculiaris Luer MO, PMA ). Epiphytic herbs. Distribution in PILA 700– 700 m, low elevation mixed forest. Global distribution: Costa Rica ,

Platystele oxyglossa (Schltr.) Garay Panama. Records: F. Solís 25 (CR, INB, MO). Epiphytic herbs. Distribution in PILA 1700–2600 m, cloud forest, low elevation oak forest, high elevation oak Pleurothallis phyllocardia Rchb.f. Epiphytic herbs. Distribution in PILA 700–2500 m,

low elevation mixed forest, mixed forest in transition to 1600 m, mixed forest in transition to cloud forest. Global low elevation forest, mixed forest in transition to cloud distribution: Costa Rica, Panama. Records: M. Moraga forest, cloud forest, low elevation oak forest, pasture. 435 ( INB, MO) .

Global distribution: Costa Rica to Panama. Records: A.

Fernández 914 (CR, MO), A. Rodríguez 10810 (BM, Ponthieva formosa Schltr.

INB, MO, PMA), B. Hammel 1425 (MO), D. Solano 3916 Epiphytic herbs. Distribution in PILA 1600–2500 m, (BM, INB, MO, PMA), D. Solano 5224 (BM, INB, MO, mixed forest in transition to cloud forest, cloud forest, PMA), E. Alfaro 4687 (INB), G. Davidse 29171 (MO), low elevation oak forest. Global distribution: Mexico to G. Herrera 5252 (CR, INB, MO), G. McPherson 15883 Panama. Records: E. Fletes 179 (INB).

(MO), K. Utley 5677 (MO), L. Angulo 337 (INB).

Prescottia stachyodes (Sw.) Lindl.

Pleurothallis rhodoglossa Schltr. Terrestrial herbs. Distribution in PILA 0–2200 m, low Epiphytic herbs. Distribution in PILA 500–1900 m, low elevation mixed forest, mixed forest in transition to low elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest, elevation forest, mixed forest in transition to cloud forest. cloud forest, low elevation oak forest. Global distribution: Global distribution: Costa Rica to Ecuador. Records: G. Mexico to Paraguay, Greater Antilles, Lesser Antilles. Herrera 5253 (CR, INB, MO). Records: G. Herrera 6093 (INB).

Pleurothallis rowleei Ames Prosthechea brassavolae (Rchb.f.) W.E.Higgins Epiphytic herbs. Distribution in PILA 1100–2500 m, Epiphytic herbs. Distribution in PILA 1500–2600 m, mixed forest in transition to low elevation forest, mixed mixed forest in transition to cloud forest, cloud forest, forest in transition to cloud forest, cloud forest, low low elevation oak forest, high elevation oak forest. Global elevation oak forest. Global distribution: Costa Rica to distribution: Mexico to Panama. Records: A. Fernández Ecuador. Records: G. Davidse 29077 (CR, MO), G. 1851 (INB), B. Gamboa 567 (INB, MO), D. Solano 3897 Davidse 29210 (CR, MO). (BM, INB, MO, PMA), M.A. Thomas 1667 (MO), T. Croat 26936 (MO).

Pleurothallis ruscifolia (Jacq.) R.Br. Epiphytic herbs. Distribution in PILA 400–1800 m, low Prosthechea ochracea (Lindl.) W.E.Higgins elevation mixed forest, mixed forest in transition to low Epiphytic herbs. Distribution in PILA 1300–1900 elevation forest, mixed forest in transition to cloud forest. m, mixed forest in transition to cloud forest. Global Global distribution: Mexico to Brazil, Greater Antilles, distribution: Mexico to Costa Rica. Records: A. Rodríguez Lesser Antilles. Records: D. Solano 4191 (BM, INB, MO, 11383 (INB). PMA). Prosthechea pseudopygmaea (Finet) W.E.Higgins

Pleurothallis tonduzii Schltr. Epiphytic herbs. Distribution in PILA 1100–2200 m, Epiphytic herbs. Distribution in PILA 700–1700 m, low mixed forest in transition to low elevation forest, mixed elevation mixed forest, mixed forest in transition to low forest in transition to cloud forest, cloud forest, low elevation forest, mixed forest in transition to cloud forest. elevation oak forest. Global distribution: Mexico to Costa Global distribution: Costa Rica, Panama. Records: B. Rica. Records: C. Hamilton 3909 (MO), G. Davidse Gamboa 642 (INB), G. Davidse 28437 (CR, MO). 29160 (CR, MO).

Pleurothallis volcanica Luer Prosthechea vespa (Vell.) W.E.Higgins Epiphytic herbs. Distribution in PILA 1300–1600 Epiphytic herbs. Distribution in PILA 700–1700 m, low m, mixed forest in transition to cloud forest. Global elevation mixed forest, mixed forest in transition to low distribution: Costa Rica, Panama. Records: G. Davidse elevation forest, mixed forest in transition to cloud forest. 28380 (MO), F. Quesada 1725 (INB, MO). Global distribution: Nicaragua to Brazil, Greater Antilles, Lesser Antilles. Records: A. Rodríguez 11403 (BM, INB,

Polystachya masayensis Rchb.f. MO, PMA ), D. Santamaría, Oct. 26, 2007 ( Field obs.), G. Epiphytic herbs. Distribution in PILA 1000–1900 m, Herrera 2686 (INB), J. Dwyer 8774 ( MO) . mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest. Global distribution: Pseudocentrum hoffmannii (Rchb.f.) Rchb.f. Mexico to Panama, Venezuela. Records: L.D. Gómez Terrestrial herbs. Distribution in PILA 2200–3000 m, low 23265 ( MO) . elevation oak forest, high elevation oak forest. Global distribution: Costa Rica, Panama. Records: B. Hammel

Ponthieva brenesii Schltr. 5777 (MO), G. Davidse 23990 (CR, MO), G. Herrera Facultatively epiphytic herbs. Distribution in PILA 1400– 3672 (INB, MO). 3685 (INB), E. Alfaro 4661 (INB), G. Davidse 28805 (CR, MO), G. Herrera 6101 (CR, INB, MO).

Psilochilus sp. A . Terrestrial herbs. Distribution in PILA 3400–3500 m, Scaphyglottis acostae (Schltr.) C.Schweinf. páramo. Global distribution: Endemic to Costa Rica. Epiphytic herbs. Distribution in PILA 1000–2700 m, Records: A. Rodríguez 11536 (INB). mixed forest in transition to low elevation forest, mixed The above material was determined as being of a forest in transition to cloud forest, cloud forest, low currently undescribed species by R.L. Dressler in the elevation oak forest, high elevation oak forest. Global Manual de Plantas de Costa Rica (MO, Hammel et al. distribution: Costa Rica, Panama. Records: D. Solano 2003). 5142 (BM, INB, MO, PMA), D. Solano 5216 (BM, INB, MO, PMA), W. R. Maxon 5399 (CR).

Pterichis galeata Lindl. Terrestrial herbs. Distribution in PILA 2300–3300 m, low Scaphyglottis corallorrhiza (Ames) Ames, F.T.Hubb. & elevation oak forest, páramo. Global distribution: Costa C.Schweinf. Rica, Venezuela to Peru. Records: G. Davidse 28578 Epiphytic herbs. Distribution in PILA 1500–2800 m, (MO), G. Davidse 29005 (CR, MO), G. Davidse 29028 mixed forest in transition to cloud forest, cloud forest, (MO), G. Herrera 3664 (INB, MO). low elevation oak forest, high elevation oak forest. Global distribution: Costa Rica, Panama. Records: A.K. Monro

Pterichis habenarioides (F.Lehm. & Kraenzl.) Schltr. 5483 (BM, INB, MO, PMA), G. Davidse 28843 (INB, Terrestrial herbs. Distribution in PILA 2300–3500 m, low MO), G. Herrera 3713 (INB, MO). elevation oak forest, páramo. Global distribution: Costa Rica to Ecuador, Venezuela. Records: B. Gamboa 1718 Scaphyglottis sigmoidea (Ames & C.Schweinf.) (INB, MO) , E. Alfaro 3713 (INB), G. Davidse 23852 (CR, B.R.Adams MO), G. Davidse 28778 (MO), G. Davidse 29296 (CR, Epiphytic herbs. Distribution in PILA 1600–2200 m, MO), G. Davidse 29359 (CR, MO), G. Davidse 29414 mixed forest in transition to cloud forest, cloud forest, (MO), G. Herrera 3739 (INB, MO). low elevation oak forest. Global distribution: Costa Rica, Panama. Records: D. Solano 3930 (BM, INB, MO,

• ε-PILA Pterichis herrerae Kolan. PMA ). Terrestrial herbs. Distribution in PILA 3300– 3300 m, páramo. Global distribution: Endemic to Costa Rica. Sobralia amabilis (Rchb.f.) L.O.Williams Records : G. Herrera 3665 (INB, MO). Facultatively epiphytic herbs. Distribution in PILA 1100– 3100 m, mixed forest in transition to low elevation forest,

Rossioglossum krameri (Rchb.f.) M.W.Chase & mixed forest in transition to cloud forest, cloud forest, low N.H.Williams elevation oak forest, high elevation oak forest, páramo. Epiphytic herbs. Distribution in PILA 900–1100 m, Global distribution: El Salvador to Panama. Records: A. mixed forest in transition to low elevation forest. Global Chacón 552 (INB, MO), A. Fernández 955 (CR, INB, distribution: Nicaragua to Panama. Records: E. Alfaro MO), E. Alfaro 3969 (INB), E. Alfaro 4705 (INB), G. 3333 (INB). Herrera 6133 (CR, INB, MO), F. Quesada 1832 (INB), F. Quesada 1952 (INB).

Rossioglossum oerstedii (Rchb.f.) M.W.Chase & N.H.Williams Sobralia leucoxantha Rchb.f. Clavelillo (CR) . Facultatively epiphytic herbs. Distribution in PILA Epiphytic herbs. Distribution in PILA 1700–3100 600–2000 m, low elevation mixed forest, mixed forest

m, cloud forest, low elevation oak forest, high elevation in transition to low elevation forest, mixed forest in oak forest, páramo. Global distribution: Costa Rica, transition to cloud forest. Global distribution: Mexico to Panama. Records: E. Alfaro 3210 (INB), E. Navarro 479 Panama. Records: G. Herrera 2835 (MO). (INB, MO), G. Davidse 28612 (MO), M.A. Thomas 1646 (MO). Sobralia warszewiczii Rchb.f. Facultatively epiphytic herbs. Distribution in PILA

Scaphosepalum microdactylum Rolfe 1000 –1800 m, mixed forest in transition to low elevation Epiphytic herbs. Distribution in PILA 0–2600 m, low forest, mixed forest in transition to cloud forest. Global elevation mixed forest, mixed forest in transition to low distribution: Nicaragua to Panama. Records: A. Rodríguez elevation forest, mixed forest in transition to cloud forest, 10309 ( INB). cloud forest, low elevation oak forest, high elevation oak forest. Global distribution: Mexico to Ecuador. Records: Solenocentrum costaricense Schltr. A. Fernández 985 ( INB), E. Alfaro 3676 ( INB), E. Alfaro Epiphytic herbs. Distribution in PILA 1500–2100 m ,

mixed forest in transition to cloud forest, cloud forest, Epiphytic herbs.Distribution in PILA 1100–2400 m, mixed low elevation oak forest. Global distribution: Costa Rica, forest in transition to low elevation forest, mixed forest in Panama. Records: G. Davidse 23757 (CR, MO). transition to cloud forest, cloud forest, low elevation oak forest. Global distribution: Mexico to Panama, Brazil. Specklinia endotrachys (Rchb.f.) Pridgeon & M.W.Chase Records : G. Herrera 5256 (CR, INB, MO).

Epiphytic herbs. Distribution in PILA 1000–2500 m,

mixed forest in transition to low elevation forest, mixed Stelis segoviensis (Rchb.f.) Pridgeon & M.W.Chase forest in transition to cloud forest, cloud forest, low Epiphytic herbs. Distribution in PILA 1000–2700 m, elevation oak forest. Global distribution: Mexico to mixed forest in transition to low elevation forest, mixed Venezuela. Records: E. Alfaro 3646 (INB), E. Alfaro 4597 forest in transition to cloud forest, cloud forest, low (INB), G. Davidse 28729 (MO), G. Davidse 28921 (INB, elevation oak forest, high elevation oak forest. Global MO), G. Herrera 3411 (INB, MO), G. McPherson 15941 distribution: Mexico to Ecuador. Records: T. Croat 26962 (MO), F. Quesada 1481 (INB), F. Quesada 1574 (INB). (MO).

Stanhopea ecornuta Lem. Stelis spathulata Poepp. & Endl. Flor de vaca, Torito sin cacho (CR). Epiphytic herbs. Distribution in PILA 1200–1400 m, Epiphytic herbs. Distribution in PILA 500– 500 m, low mixed forest in transition to low elevation forest, mixed elevation mixed forest. Global distribution: Guatemala to forest in transition to cloud forest. Global distribution: Panama. Records: A. Rodríguez 11242 (INB). Costa Rica to Peru, Venezuela. Records: L.D. Gómez 23239 (MO).

Stelis conochila (Luer) Pridgeon & M.W.Chase Epiphytic herbs. Distribution in PILA 1600–1600 Stelis superbiens Lindl. m, mixed forest in transition to cloud forest. Global Epiphytic herbs. Distribution in PILA 1100–2500 m, distribution: Endemic to Costa Rica. Records: F. Solís 24 mixed forest in transition to low elevation forest, mixed (INB, MO). forest in transition to cloud forest, cloud forest, low elevation oak forest. Global distribution: Mexico to Peru,

Stelis macrantha Pridgeon & M.W.Chase Venezuela. Records: G. Davidse 28695 (MO), G. Davidse Epiphytic herbs. Distribution in PILA 1900– 1900 m, 28953 ( MO) . cloud forest. Global distribution: Costa Rica, Panama. Records: G. Davidse 28820 ( MO) . Stelis tonduziana Schltr. Epiphytic herbs. Distribution in PILA 1500–1500

Stelis gelida (Lindl.) Pridgeon & M.W.Chase m, mixed forest in transition to cloud forest. Global Epiphytic herbs. Distribution in PILA 1100–1700 m, distribution: Endemic to Costa Rica. Records: G. Herrera mixed forest in transition to low elevation forest, mixed 2718 (INB, MO). forest in transition to cloud forest. Global distribution: USA to Bolivia, Greater Antilles, Lesser Antilles. Records: Telipogon costaricensis Schltr. L.D. Gómez 23274 (MO). Epiphytic herbs. Distribution in PILA 2500–3300 m, low elevation oak forest, páramo. Global distribution:Endemic ε-PILA Stelis montana L.O.Williams to Costa Rica. Records: G. Herrera 3621 (INB). Epiphytic herbs. Distribution in PILA 2000–2500 m, cloud forest, low elevation oak forest. Global distribution: Telipogon distantiflorus (Ames& C.Schweinf.) N.H.Williams Endemic to Panama. Records: Allen 1463 (MO). & Dressler Epiphytic herbs. Distribution in PILA – m. Global

Stelis morganii Dodson & Garay distribution: Elevation unknown. Endemic to Costa Rica. Epiphytic herbs.Distribution in PILA 2100– 2100 m, cloud Records: G. Herrera 5945 (CR, INB, MO). forest, low elevation oak forest. Global distribution: Costa Rica, Ecuador. Records: R. Aguilar 1141 (INB, MO). Telipogon gracilipes Schltr. Epiphytic herbs. Distribution in PILA 2300–3000 m, low

Stelis papillifera (Rolfe) Pridgeon & M.W.Chase elevation oak forest, high elevation oak forest. Global Epiphytic herbs. Distribution in PILA 1500–1600 distribution: Endemic to Costa Rica. Records: G. Davidse m, mixed forest in transition to cloud forest. Global 28786 (MO). distribution: Endemic to Costa Rica. Records: G. Davidse 28415 (MO). Telipogon leila-alexandrae Braas Epiphytic herbs. Distribution in PILA 2900–3200 m, low

Stelis parvula Lindl. elevation oak forest, high elevation oak forest, páramo. Global distribution: Endemic to Costa Rica. Records: G. Xylobium elongatum (Lindl. & Paxton) Hemsl. Herrera 6000 (INB, MO). Epiphytic herbs. Distribution in PILA 700–1700 m, low elevation mixed forest, mixed forest in transition to low

Telipogon storkii Ames & C.Schweinf. elevation forest, mixed forest in transition to cloud forest. Epiphytic herbs. Distribution in PILA 2900–3100 m, low Global distribution: Mexico to Peru. Records: G. Davidse elevation oak forest, high elevation oak forest, páramo. 28431 (CR, MO), F. Quesada 1659 (INB), F. Quesada Global distribution: Endemic to Costa Rica. Records: 1683 (INB), L. Acosta 2407 (INB), L.D. Gómez 23096 G. Davidse 26073 (MO), G. Davidse 28608 (MO), G. (MO). Davidse 28896 (MO). Trichopilia marginata Henfr. POACEAE Epiphytic herbs. Distribution in PILA 900–1300 m, mixed forest in transition to low elevation forest, mixed forest in Acroceras zizanioides (Kunth) Dandy transition to cloud forest. Global distribution: Nicaragua Terrestrial herbs. Distribution in PILA 0–1700 m, low to Colombia. Records: M. Moraga 421 (INB, MO). elevation mixed forest, mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest,

ε-PILA Trichopilia olmosii Dressler cloud forest. Global distribution: Mexico to Argentina, Epiphytic herbs. Distribution in PILA 1000–1200 m, Antilles, Asia, tropical Africa. Records: A. Rodríguez mixed forest in transition to low elevation forest. Global 11189 (BM, INB, MO, PMA). distribution: Endemic to Panama. Records: R. Dressler 6288 (MO). Agrostis bacillata Hack. Terrestrial herbs. Distribution in PILA 2200–3800 m,

Trichosalpinx arbuscula (Lindl.) Luer low elevation oak forest, páramo. Global distribution: Epiphytic herbs. Distribution in PILA 700–1200 m, Costa Rica, Panama. Records: A. Chacón 519 (MO), A. S. low elevation mixed forest, mixed forest in transition to Weston 10191 (MO), B. Gamboa 1728 (CR, INB, MO), low elevation forest. Global distribution: Costa Rica to E. Alfaro 3291 (INB), E. Alfaro 3301 (INB), E. Alfaro Bolivia. Records: A. Fernández 788 (INB). 3784 (INB), G. Davidse 23853 (CR, MO), G. Davidse 29277 (CR, MO), F. Quesada 1793 (INB).

Trichosalpinx cedralensis (Ames) Luer Epiphytic herbs. Distribution in PILA 1100–1500 m, Agrostis perennans (Walter) Tuck. mixed forest in transition to low elevation forest, mixed Terrestrial herbs. Distribution in PILA 2300–3400 m, low elevation oak forest, páramo. Global distribution: Mexico forest in transition to cloud forest. Global distribution: Guatemala to Bolivia, Venezuela. Records: G. Herrera to Panama, Bolivia. Records: A.K. Monro 4163 (BM), G. 5427 (CR, INB, MO), R. Aguilar 1164 (INB, MO). Davidse 28591 (MO).