Rinhatiana, Irwin & Winterton, 2024

Irwin, Michael E. & Winterton, Shaun L., 2024, New genera of stiletto flies endemic to Madagascar (Therevidae: Therevinae), European Journal of Taxonomy 932 (1), pp. 1-33 : 30-31

publication ID	https://doi.org/ 10.5852/ejt.2024.932.2507
publication LSID	lsid:zoobank.org:pub:EA652637-A88D-41A7-A586-8D57770B9C0E
DOI	https://doi.org/10.5281/zenodo.11035248
persistent identifier	https://treatment.plazi.org/id/255759B6-BAFF-4386-84B7-8A4B38A44CD3
taxon LSID	lsid:zoobank.org:act:255759B6-BAFF-4386-84B7-8A4B38A44CD3
treatment provided by	Plazi
scientific name	Rinhatiana
status	gen. nov.

Treatment

Key to genera of Malagasy Therevidae View in CoL View at ENA and species of Rinhatiana gen. nov. and Tianarinha gen. nov.

1. Femora with regular filiform setae of varying lengths, lacking adpressed scale-like setae; costal vein ending between veins R 4 and CuA+CuP, never circumambient around wing; female terminalia with, at most, a single set of acanthophorite spines ..............................................................2 ( Therevinae View in CoL )

– Femora with regular filiform setae of varying lengths, admixed with adpressed scale-like setae, especially on mid and hind legs; costal vein circumambient around wing; female terminalia with two distinct sets of acanthophorite spines on each side (A1 and A2) ...................................................... 3

2. Dorsal surface of wing vein R 1 with setulae; palpus with apical pit; female sternite 8 without strong macrosetae; genitalia with three spermathecae and no spermathecal sac ........................................... ........................................................................................ Rueppellia Wiedemann, 1830 View in CoL ( Phycusinae )

– Wing vein R 1 without setulae; palpus without an apical pit; female sternite 8 with strong macrosetae; genitalia with two spermathecae and spermathecal sac is present ...................................................... ................................................................................. Microgephyra Lyneborg, 1972 View in CoL ( Xestomyzinae View in CoL )

3. Wing vein R 1 without setulae ............................................................................................................ 4

– Dorsal surface of wing vein R 1 with setulae ..................................................................................... 5

4. Sternum with setae present medially in furrow; mid coxae with setae on posterior surface .............. ................................................................................................................... Irwiniella Lyneborg, 1976

– Sternum with setae present laterally only, rarely in medial furrow; mid coxae without setae on posterior surface .................................................................................................. Stenopomyia Lyneborg, 1976

5. Prosternum with setae medially; one pair of scutellar macrosetae; frons with dark setae, pubescence grayish-brown with a dark band from base of antenna to eye margin ( Fig. 14 View Fig ); gena with dark setae ventrally; abdomen predominantly brown .................................................. Tianarinha gen. nov....6

– Prosternum without setae; two pairs of scutellar macrosetae; frons usually lacking setae, pubescence silver or metallic tan around antennal bases, matte black above this and frequently grayish-silver dorsally to ocellar tubercle (e.g., Fig. 9 View Fig ); gena with pale setae; abdomen predominantly bright yellow, sometimes brown laterally along tergites ................................................... Rinhatiana gen. nov....7

6. Extensive dark setae on the lower part of occiput, gena ( Figs 10 View Fig ; 14A–B View Fig ) and coxae; flagellum style relatively elongate; few dark setae on frons just above antennae; setae on posterior surface of hind coxae ( Fig. 11B View Fig ) ..................................................................................... T. goodmani gen. et sp. nov.

– Setae on the lower head and coxae fewer in number and shorter ( Fig. 12A View Fig , 13A View Fig ); flagellum style relatively short; frons only with setae dorsally, no setae just above antennae; setae absent on posterior surface of hind coxae ( Fig. 13A View Fig ) ................................................................... T. micet gen. et sp. nov.

7. Two pairs of supra alar macrosetae; dorsocentral macrosetae present ( Figs 5–6 View Fig View Fig ); Male frons width at narrowest point slightly wider than anterior ocellus ( Fig. 9E View Fig ); short, dark setae present dorsally on frons .................................................................................. R. distincta ( Lyneborg, 1976) comb. nov.

– One pair of supra alar macrosetae; dorsocentral macrosetae absent; male frons width at narrowest point narrower than anterior ocellus, eyes almost contiguous ( Fig. 9A, C, G View Fig ); frons lacking setae .. ........................................................................................................................................................... 8

8. Frons with metallic-tan pubescence around base of antennae ( Fig. 9C, D View Fig ); abdomen dark brown (yellow dorsally in female), cream-white posteriorly on each segment, silver pubescence absent ( Figs 3–4 View Fig View Fig ) ................................................................................................ R. cracentis gen. et sp. nov. – Frons with silver pubescence around base of antennae and matte black area medially ( Fig. 9A, G View Fig ); abdomen predominantly yellow, sometimes brown laterally along tergites, triangular patches of silver pubescence laterally on at least tergites 2 and 3 ............................................................................... 9

9. Male frons with matte-black pubescence relatively narrow, brown pubescence dorsally ( Fig. 9A View Fig ); female frons with dorsomedial stripe of matte-black pubescence relatively narrow, suffused with brown pubescence laterally ( Fig. 9B View Fig ); male terminalia black ( Figs 1–2 View Fig View Fig ) ...... R. arctifestuca gen. et sp. nov.

– Male frons with matte-black pubescence broader, lacking brown pubescence dorsally ( Fig. 9G View Fig ); female with dorsomedial stripe of matte-black pubescence broader, mostly lacking brown pubescence in upper portion ( Fig. 9H View Fig ); male terminalia yellow ( Figs 7–8 View Fig View Fig ) ............ R. latifestuca gen. et sp. nov.