Bocchus schmalhauseni, Perkovsky & Olmi & Vasilenko & Capradossi & Guglielmino, 2020

Perkovsky, Evgeny E., Olmi, Massimo, Vasilenko, Dmitry V., Capradossi, Leonardo & Guglielmino, Adalgisa, 2020, First Bocchus Ashmead (Hymenoptera: Dryinidae) from Upper Eocene Rovno amber: B. schmalhauseni sp. nov., Zootaxa 4819 (3), pp. 544-556 : 545

Bocchus schmalhauseni sp. nov.

( Figs 1–3 , 5 )

Type material. Holotype: ♀, SIZK L-119: UKRAINE: specimen in Upper Eocene Rovno amber, collected in Vladimirets district near Voronki village , Rovno region, Ukraine ( SIZK). Horizon : Priabonian (35–38 Ma).

Diagnosis. Macropterous female of Bocchus ( Fig. 1 ) with OOL more than three times as long as OPL; forewing uniformly darkened ( Fig. 1 ); petiole distinctly visible ( Fig. 1 ); enlarged claw ( Figs 2 , 3 , 5 ) with one row of small teeth and one medial lamella on inner margin; protarsomere 5 ( Figs 2 , 3 , 5 ) with inner band, one preapical lamella and no bristles.

Description of female holotype ( Figs 1–3 , 5 ). Fully winged ( Fig. 1 ); length 3.3 mm.

Black. Antenna not visible. Head dull, apparently completely granulate, not reticulate rugose; frontal line not visible; occipital carina complete; ocelli only partly visible; OOL = 2; OPL = 7; TL = 6. Mesosoma longer than head (47:28), shorter than metasoma (47:53). Pronotum crossed by strong transverse impression, with sculpture not distinct; pronotal tubercle reaching tegula. Dorsal side of mesosoma not visible. Epicnemium concealed. Metapectal-propodeal disc apparently reticulate rugose; propodeal declivity dull and rugose. Forewing uniformly darkened ( Fig. 1 ); distal part of 2r-rs&Rs vein longer than proximal part (18:7). Petiole very long ( Fig. 1 ), much shorter than rest of metasoma (6:53). Proleg ratio: 17 (coxa): 9 (trochanter): 22 (femur): 22 (tibia): 12 (protarsomere 1): 2 (protarsomere 2): 2 (protarsomere 3): 13 (protarsomere 4): 17 (protarsomere 5). Enlarged claw ( Figs 2 , 3 , 5 ) slightly shorter than protarsomere 5 (15:17), with one medial lamella and one row of small teeth (number not distinct) situated in the distal half. Protarsomere 5 ( Figs 2 , 3 , 5 ) with inner band, one preapical lamella and no bristles. Rudimentary claw present. Arolium length about 21% of fifth protarsomere length (3.6:17). Mesoleg ratio: 12 (coxa): 5 (trochanter): 20 (femur): 19 (tibia): 26 (tarsus). Metaleg ratio: 20 (coxa): 4 (trochanter): 26 (femur): 26 (tibia): 31 (tarsus). Tibial spurs not visible.

Male. Unknown.

Remarks. The new species is attributed to the genus Bocchus mainly because it has the protarsomere 5 with an inner band and has one preapical lamella. This special shape of protarsomere 5 can be found only in Bocchus ( Olmi et al. 2019) .

Hosts. Unknown.

Etymology. This species is named after Ivan Ivanovich Schmalhausen, the famous evolutionary biologist.

Figures

FIGURES 1–2. Bocchus schmalhauseni sp. nov.: female holotype: 1, body in lateral view; 2, foreleg.

FIGURES 3–4. 3. Bocchus schmalhauseni sp. nov.: female holotype: chela. 4. Bocchus cenomanianus Olmi, Rasnitsyn & Guglielmino, female holotype (from Olmi et al. 2010). Scale bar: fig. 3 = 90 µm, fig. 4 = 1.5 mm.

FIGURES 5–6. Chelae. 5, Bocchus schmalhauseni sp. nov.: female holotype. 6, Bocchus primaevus Martins & Melo (CGC 8339). a = arolium; b = long bristle; e = enlarged claw; i = inner band; rc = rudimentary claw; t = protarsomere 5. Scale bar: fig. 5 = 70 µm, fig. 6 = 60 µm.