Protoita istvani Ulmer & Krogmann, 2023

UImer, Jonah M., Jansta, Petr, Azar, Dany & Krogmann, Lars, 2023, At the dawn of megadiversity - Protoitidae, a new family of Chalcidoidea (Hymenoptera) from Lower Cretaceous Lebanese amber, Journal of Hymenoptera Research 96, pp. 879-924 : 879

publication ID

https://dx.doi.org/10.3897/jhr.96.105494

publication LSID

lsid:zoobank.org:pub:DB845B36-62BA-4DA1-8370-5B36E5916BB0

persistent identifier

https://treatment.plazi.org/id/7550D3A4-0E73-444C-9692-E6AFA6915815

taxon LSID

lsid:zoobank.org:act:7550D3A4-0E73-444C-9692-E6AFA6915815

treatment provided by

Journal of Hymenoptera Research by Pensoft

scientific name

Protoita istvani Ulmer & Krogmann
status

sp. nov.

Protoita istvani Ulmer & Krogmann sp. nov.

Fig. 6 View Figure 6

Diagnosis.

The female of P. istvani differs from all others in the genus by the following combination of characters: Head flattened antero posteriorly, disc-like (Fig. 6D, G View Figure 6 ). Fl1-3 elongate, about 4 × as long as wide (Fig. 6B View Figure 6 ). Clava 2-segmented (Fig. 6B View Figure 6 ).

Description.

Female. Body length 918. Coloration auburn with dark antennae. Eyes white. Wings hyaline, uniformly setose. Metasoma damaged, tergal segments detached from body (Fig. 6A View Figure 6 ). Head broad, wider than mesosoma in dorsal view, anteroposteriorly narrow, head longer than wide in lateral view (Fig. 6D, G View Figure 6 ). Frons 2 × as long as face (Fig. 6F, G View Figure 6 ). Eyes large, 0.8 × height of head, about 1.6 × as high as long. Malar sulcus present; malar space short, only about 0.15 × as long as eye height (Fig. 6D View Figure 6 ). POL slightly shorter than LOL. Toruli equal in distance from each other as to inner eye margin. Maxillary palps 3-segmented (Fig. 6G View Figure 6 :mxp) Antenna length 676. Scape 2 × as long as pedicel, gradually broadened from base (Fig. 6B View Figure 6 ). Pedicel same length or slightly shorter than fu1, about 1.5 × as wide as fu1 width. Flagellomeres gradually widening; fu1-3 about 4 × as long as wide, fu6 3 × as long as wide, fu7-10 elongated, but less than 2 × as long as wide. Clava 2 segmented. MPS present on all segments (Fig. 6B View Figure 6 ). Mesosoma length 299. Mesosoma in lateral view heavily sloped; nearly 135° relative to mesosoma (Fig. 6C View Figure 6 ). Mesonotum roughly 0.5 × as long as mesosoma. Notauli present as simple depressions. Mesopleuron dorsoventrally elongate, 2.4 × as long as wide (Fig. 6C View Figure 6 :pl2). Axilla not advanced, not extending beyond the anterior margin of mesoscutellum (Fig. 6A View Figure 6 :axl). Mesoscutellum ⅓x as long as mesonotum. Metanotum band-like, 0.3 × as long as mesoscutellum (Fig. 6C View Figure 6 :no3). Propodeum is only slightly longer than mesoscutellum (Fig. 6C View Figure 6 :prp). Wings. Fore wing length roughly 1.1 × as long as body, 3.2 × as long as wide. Marginal setae of fore wing short, length of longest seta 26. Cubital vein ephemeral, roughly 1.8 × as long as basal vein. Basal vein pigmented. Submarginal vein long and narrow, length 393. Marginal vein narrow, about 0.2 × as long as submarginal vein. Postmarginal vein short and tapering, slightly shorter than marginal vein (63:77) (Fig. 6E View Figure 6 ). Stigmal vein at 80° angle to wing margin. Uncus present. Hind wing obscured by fore wing. Legs. Tarsomere 4 on all tarsi very short relative to other tarsal segments (Fig. 6A View Figure 6 ). Basitarsal comb of fore leg present. Hindtibia elongate, equal in length to tarsi (Fig. 6A View Figure 6 ). Forecoxa greatly impressed into prosternum (possibly an artifact). Hind coxa elongated, about 3.1 × as long as broad (Fig. 6C View Figure 6 :cx3). Metasoma 1.3 × as long as mesosoma. Metasoma broadly associated with mesosoma; remnants of Mt2 visible (Fig. 6C View Figure 6 ). Most of tergal and sternal segments damaged. Only 3 tergites countable, all with transverse setal row posteromedially. Two elongate sensillae visible off of terminal sternite. Metasomal cerci digitiform (visible only on left side; Fig. 6F View Figure 6 :cer). Ovipositor length 140, not extending beyond distal point of ovipositor sheath; ovipositor sheaths roughly equal in length to ovipositor, uniform width along entire length, about 4.2 × as long as wide (Fig. 5F View Figure 5 :3v).

Male. Unknown

Material examined.

Holotype: female, Roum - Aazour - Homsiyyeh , Caza Jezzine , Mouhafazet South Lebanon; lower Barremian. In amber mounted in Canada Balsam. Deposited at Natural History Museum of the Lebanese University, accession/specimen number: HAR26.

Type condition.

Specimen with bubbles surrounding MPS on several flagellomeres, mesosoma detached from the amber, and metasomal tergites damaged because they are detached.

Etymology.

The specific epithet is a patronym in honor of István Mikó, the first author’s mentor and close friend who instilled in him his passion for entomology and morphology.