Quediellus debilis (Horn, 1878), comb. res.

Brunke, Adam J., 2022, A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages, ZooKeys 1134, pp. 129-170 : 129

Quediellus debilis (Horn, 1878), comb. res.

Figs 5C, D , 8K-Q , 9H, I , 11E (map), 12

Quedius debilis Horn, 1878: 156, 165.

Quediellus debilis ; Casey 1915 (redescription).

Quediellus helenae Casey, 1915: 403 syn. nov.

Quediellus humilis Casey, 1915: 403 syn. nov.

Quediellus nanulus Casey, 1915: 402 syn. nov.

Quedius (Raphirus) debilis ; Smetana 1971 (redescription); Brunke et al. 2021 (phylogeny, as Q. nanulus ).

Type locality.

California (unknown locality), United States.

Type material.

Lectotype (female, MCZ, examined digitally): ‘Cala’ [white typed label] / Q. debilis H. [handwritten label] / Type 7270 [red label].

The female lectotype of this species is a classic example of the ‘debilis’ morphotype (see below): a larger specimen with elytra at sides much longer than pronotum at midline, elytra at apex wider than pronotum.

Figures

Figure 5. A-C dorsal head A, B showing extent of head punctation (white arrow) and color of first antennomere base (black arrow) D-F elytral punctation and microsculpture A Paraquedius puncticeps (Horn) B P. marginicollis sp. nov. C, D Quediellus debilis (Horn) E Quedionuchus longipennis (Mannerheim) F Quedius densiventris (Casey). Abbreviations: af = anterior frontal puncture; b = basal puncture; pf = posterior frontal puncture; sa = supra-antennal puncture. Scale bars: 0.5 mm.

Figure 8. A aedeagus in ventral view B, C, D, G, H, K, L median lobe of aedeagus B, G, K, L in ventral view, C, D, H, O-Q in lateral view E, F, I, J, M, N underside of paramere showing peg setae A-F Paraquedius marginicollis sp. nov. E atypical specimen from Clallam County, Washington, United States G-J Paraquedius puncticeps (Horn) I lectotype, from Vancouver, British Columbia, Canada J non-type specimen from Vancouver Island K-Q Quediellus debilis (Horn), specimens from east (K, M) and west (L, M) sides of the continental divide. Filled circles = typical ' nanulus' morphotype; open circles = typical ' debilis' morphotype. Scale bars: 0.1 mm.

Figure 9. A-I female tergite X A Iratiquedius amabilis (Smetana) B I. mutator (Smetana) C I. prostans (Horn) D I. seriatus (Horn), evenly convex E I. uncifer sp. nov., discal impression F Paraquedius puncticeps (Horn) G P. marginicollis sp. nov. H-I Quediellus debilis (Horn). Pigmented area on tergite in E not shown for clarity but similar to I. seriatus. Scale bars: 0.1 mm.

Figure 11. Distributions of A Iratiquedius amabilis (Smetana) (circles), I. mutator (Smetana) (squares) B I. prostans (Horn) (black - specimens examined, white - literature records) C I. seriatus (Horn) (circles), I. uncifer sp. nov. (squares) D Paraquedius puncticeps (Horn) (circles), P. marginicollis sp. nov. (squares) E Quediellus debilis (Horn) (circles) (black - specimens examined, white - literature records).