Stachys hakkariensis Akçiçek & Fırat, Phytotaxa

Akçiçek, Ekrem, 2020, Taxonomic revision of Stachys sect. Olisia (Lamiaceae: Lamioideae) in Turkey, Phytotaxa 449 (2), pp. 109-148 : 135-136

8. Stachys hakkariensis Akçiçek & Fırat, Phytotaxa 257: 168 (2016) ( Figs. 1D , 6, 8 )

Type:― TURKEY. C9 Hakkari: between Şine and Marinus (Çîyayêreş Mountain, Bemro region ), rocky slopes, eroded slopes, 37 o 29’53”N 43 o 27’47”E, 1200 m, 7 July 2015, M. Fırat 32600 (holo: GAZI!; iso: ISTE!, Hb. Akçiçek!, Hb. M.Fırat!)

Description:―Suffrutescent perennial herbs, without basal rosettes of leaves. Flowering stems 40‒80cm tall, ascending, slender, very much branched, glabrous with sparsely sessile glands. Cauline leaves linear to linear-lanceolate, 0.7‒7 × 0.1‒0.3(‒0.5) cm, entire at margin, acute at apex, glabrous with sparsely sessile glands. Floral leaves similar to cauline leaves, but smaller, linear to linear-lanceolate gradually shorter than verticillasters above, 2‒12(‒15) mm, entire at margin, acute at apex. Verticillasters (2‒)3‒5, remote, (1.2‒) 1.5‒6 cm distant, (1‒)2(‒3)-flowered. Bracteoles few, minute, setaceous, 1‒2 mm, margin antrorsely hairy. Pedicels 1.5‒2 mm long. Calyx ± regular, subcampanulate, 5.5‒7.5 mm, glabrous, mouth scarsely hairy (between calyx teeth); teeth ± equal, triangular-lanceolate, 1.5‒2 mm long, ca. 1/3 of the tube, slightly recurved, shortly mucronate-tipped (mucron ca. 0.2 mm long). Corolla white, 10‒12 mm long; lips streaked and spotted with pink inside, densely short adpressed hairy outside; tube subexserted, annulate; limb bilabiate, upper lip ± 5 mm long, the lower 3-lobed, middle lobe longer than 2 lateral lobes, 4‒5 mm long, rounded. Stamens 4, slightly exserted from corolla tube. Nutlets obovoid, faintly trigonous, slightly winged near base, 2.5‒3 × 1.7‒2 mm. blackish-brown.

Phenology: ―Flowering and fruiting from July to October.

Distribution and ecology: ―Endemic to Turkey (Eastern Anatolia). Rocky slopes, eroded slopes. 1150‒1200 m elevation.

Conservation status: ―EN [Criteria B2ab(i,iii)] ( IUCN 2017). This species is endemic to Hakkari Province in eastern Anatolia. It is only known from two locations at altitudes of 1150–1200 m, where the number of mature individuals is approximately 100. The habitat of this species is threatened by mineral extraction activities, soil erosion, and deforestation in the area.

Remarks: ― Stachys hakkariensis is closely related to S. sparsipilosa Bhattacharjee & Huber-Morath (1974: 287) , but it differs mainly by its many branched stems, herbaceous bracts, verticillasters 3‒5 with (1‒)2(‒3) flower, shorter pedicels, calyx ± regular with teeth 1.5‒2 mm long with short mucrone, corolla white and nutlets obovoid.

Figures

FIGURE 1. A. Stachys recta subsp. subcrenata (Ö. Güner 2412); B. S. atherocalyx (Akçiçek 5637); C. S. iberica subsp. iberica var. iberica (Akçiçek 5588); D. S. hakkariensis (M. Fırat 32600); E. S. sparsipilosa (Akçiçek 5598); F. S. aleurites (Akçiçek 5601); G. S. bombycina (Akçiçek 5550). Photograph D by M. Fırat.

FIGURE 8. Distribution map of Stachys angustifolia (▲), S. atherocalyx (●), S. hakkariensis (▼), S. recta subsp. subcrenata (▃) and S. sparsipilosa (⊕) in Turkey.