Pison bicellulare Pulawski, 2018

Pulawski, Wojciech J., 2018, A Revision of the Wasp Genus Pison Jurine, 1808 of Australia and New Zealand, New Guinea, and the Pacific Islands (Hymenoptera: Crabronidae), Proceedings of the California Academy of Sciences 65, pp. 1-584 : 108

Pison bicellulare Pulawski , species nova

Figures 192 -196.

NAME DERIVATION.– From the Latin prefix bis -, meaning two, and cellularis (neuter: cellulare), cellular; with reference to the presence of only two submarginal cells in this species.

RECOGNITION.– Pison bicellulare is all black (including the mid- and hindtibial spurs), with two submarginal cells ( Fig. 193 ), the posterior margin of the second submarginal cell equal to 1.7-1.8 × its height, the tegula finely punctate throughout, the eye asetose, and the propodeal dorsum without carinae bordering the enclosure. In the female (the male is unknown), the integument is depressed between the postspiracular carina and the episternal sulcus ( Fig. 194 ), a feature shared with P. aberrans and P. incurvatum . Unlike these species, however, the scutum of P. bicellulare lacks short longitudinal ridges adjacent to its posterior margin.

DESCRIPTION.– Frons slightly swollen above antennal socket, dull, minutely punctate, punctures less than one diameter apart. Distance between antennal socket and orbit slightly smaller than socket width. Gena narrow in dorsal view ( Fig. 193 ). Labrum emarginate mesally. Anteromedian pronotal pit transversely elongate, about twice as long as midocellar diameter. Scutum slightly foveate along flange, without longitudinal ridges adjacent to posterior margin; scutal punctures minute, less than one diameter apart. Scutellum inconspicuously foveate along anterior margin. Tegula enlarged, minutely punctate throughout, nearly entirely covering humeral plate. Mesopleural punctures fine, averaging about one diameter apart; interspaces inconspicuously microsculptured. Postspiracular carina present, slightly longer than midocellar diameter; integument depressed between postspiracular carina and episternal sulcus ( Fig. 194 ). Metapleural sulcus not costulate between dorsal and ventral metapleural pits. Propodeum with irregular longitudinal carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle; dorsum obliquely ridged (ridges evanescent laterally); side punctate, with or without ridges anteriorly; posterior surface transversely ridged, punctate between ridges. Forewing with two submarginal cells ( Fig. 195 ); posterior margin of second submarginal cell equal to 1.7-1.8 × its height. Hindcoxal dorsum with outer margin obtusely carinate. Punctures of tergum I fine, less than one diameter apart. Sterna finely punctate throughout, punctures of sternum II 1-2 diameters apart mesally.

Setae silvery, appressed on frons, postocellar area, scutum, and tergum I; inconspicuous on frons, not concealing integument on clypeus; straight; on lower gena straight, almost appressed, shorter than midocellar diameter. Apical depressions of terga II and III with silvery, setal fasciae.

Body black, mandible dark reddish in apical half; mid- and hindtibial spurs black.

♀.– Upper interocular distance equal to 0.96-0.98 × lower interocular distance; ocellocular distance equal to 0.9-1.2 × hindocellar diameter, distance between hindocelli equal to 1.2 × hindocellar diameter; eye height equal to 1.00-1.06 × distance between eye notches. Free margin of clypeal lamella with minute, obtuse median point ( Fig. 192 ). Dorsal length of flagellomere I

Figures

FIGURES 192–195. Pison bicellulare Pulawski, sp. nov., female. (192) Clypeus; (193) Head in dorsal view; (194) Thorax in lateral view (arrow shows depressed area of mesopleuron); (195) Distal part of forewing. FIGURE 196. Collecting localities of Pison bicellulare Pulawski, sp. nov.