Pison adnyamathanha Pulawski, 2018

Pulawski, Wojciech J., 2018, A Revision of the Wasp Genus Pison Jurine, 1808 of Australia and New Zealand, New Guinea, and the Pacific Islands (Hymenoptera: Crabronidae), Proceedings of the California Academy of Sciences 65, pp. 1-584 : 45-46

Pison adnyamathanha Pulawski , species nova

Figures 26 -29.

NAME DERIVATION.– Adnyamathanha, meaning rock people, is the name of the aboriginal tribe inhabiting the Flinders Ranges, an area where the holotype was collected.

RECOGNITION.– Pison adnyamathanha has three submarginal cells, the second recurrent vein contiguous with the second intersubmarginal vein or nearly so, the propodeum with an irregular longitudinal carina separating the side from the dorsum and the posterior surface and extending from the gastral socket area toward spiracle, and the setae appressed on tergum I, the gaster black (apical depressions of terga yellowish brown), and at least the hindfemur, tibiae, and tarsi ferruginous. It is further characterized by the setae of the lower gena straight, shorter than the midocellar diameter (but partly sinuous in the single female from Wilpena, South Australia), the frons setae silvery, and sterna densely punctate throughout. The female (the male is unknown) lacks specializations found in other species: the clypeus is evenly convex above the lamella, the lamella is prominently rounded, there are no psammophores on the gena and foreleg, and the tegula is not particularly elongate. It can be recognized by the following combination: ocellocular distance equal to 1.2-1.5 × hindocellar diameter, trimmal mandibular carina with rounded preapical tooth, scutal punctures less than one diameter apart, and tergum VI moderately broad ( Fig. 28 ); the clypeal lamella is protruding less than in P. protrudens (compare Figs. 26 and 861 ), and the setae of the propodeal dorsum extend beyond the lateral carina (they do not extend in protrudens ).

DESCRIPTION.– Frons dull, minutely punctate, punctures less than one diameter apart. Labrum not emarginate. Anteromedian pronotal pit transversely elongate, about as long as 1.5 × midocellar diameter. Scutum not foveate along flange, without longitudinal ridges adjacent to posterior margin; scutal punctures fine but well defined, less than one diameter apart. Tegula somewhat enlarged. Mesopleural punctures nearly compressed. Postspiracular carina absent. Metapleural sulcus costulate between dorsal and ventral metapleural pits. Propodeum with irregular longitudinal carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle; dorsum punctate (punctures less than one diameter apart), with short, oblique ridges on each side of median sulcus; side punctate (punctures less than one diameter apart, interspaces merging into fine ridges; posterior surface transversely ridged. Posteroventral forefemoral surface finely, closely punctate. Hindcoxal dorsum with outer margin obtusely carinate in anterior half. Punctures of tergum I less than one diameter apart on horizontal portion. Sterna punctate throughout.

Setae silvery, on scutum sparse and erect (about as long as 0.5 × midocellar diameter), appressed on tergum I, suberect to subappressed on lower gena and shorter than midocellar diameter (partly sinuous and about as long as midocellar diameter in single female from Wilpena, South Australia), oriented ventrally on frons; not concealing integument on clypeus. Apical depressions of terga with setal fasciae, fasciae silvery or with golden tinge.

Head, thorax, propodeum, and gaster black, mandible ferruginous mesally; apical depressions of terga yellowish brown. Forefemur varying from all black to all ferruginous, midfemur largely black to all ferruginous, hindfemur ferruginous; tibiae, and tarsi ferruginous.

♀.– Upper interocular distance equal to 0.84-0.96 × lower interocular distance; ocellocular distance equal to 1.1-1.3 × hindocellar diameter, distance between hindocelli equal to 1.2-1.5 × hindocellar diameter ( Fig. 27 ); eye height equal to 0.96-0.98 × distance between eye notches. Free margin of clypeal lamella prominently rounded ( Fig. 26 ). Dorsal length of flagellomere I 1.8-2.1 × apical width, of flagellomere IX 1.0 × apical width. Mandible: trimmal carina with broad incision at about three quarters of length, the proximal section of incision forming small preapical tooth. Tergum VI moderately broad ( Fig. 28 ). Length 7.9-8.6 mm; head width 2.4-2.6 mm.

♂.– Unknown.

GEOGRAPHIC DISTRIBUTION (Fig. 29).– New South Wales, South Australia, Western Australia.

RECORDS.– HOLOTYPE: ♀, AUSTRALIA: South Australia: Wilpena Pound Gap at 31°33ʹS 138°36ʹE, 5-6 Nov 1987, I.D. Naumann and J.C. Cardale ( ANIC).

PARATYPES: AUSTRALIA: New South Wales: Fowlers Gap Research Station at 31°05ʹS 141°42ʹE, 29 Nov – 2 Dec 1981, J.C. Cardale (1 ♀, ANIC) ; Menindee , 2 Dec 1971, N.W. Rodd (1 ♀, AMS; 1 ♀, CAS) . South Australia: Kings Mill Creek near Arkaroola Homestead, G.F. Gross , 20 Oct 1969 (1 ♀, CAS) and 29 Oct 1969 (1 ♀, SAM) ; Wilpena in Flinders Ranges National Park at 31°31.7ʹS 138°36.2ʹE, 21 Dec 2010, V. Ahrens and W.J. Pulawski (1 ♀, CAS) . Western Australia: Meekatharra-Billiluna Pool , Apr 1930 – Aug 1931, Canning Stock Route Expedition (1 ♀, SAM) .

Figures

FIGURES 26-28. Pison adnyamathanha Pulawski, sp. nov., female. (26) Clypeus and mandibles; (27) Head in dorsal view; (28) Apical terga in dorsal view. FIGURE 29. Collecting localities of Pison adnyamathanha Pulawski, sp. nov.

FIGURES 861-866. Pison protrudens Pulawski, sp. nov. (861) Female clypeus and mandibles; (862) Male clypeus and mandibles; (863) Female gena; (864) Female mesopleuron in slightly inclined view showing sparsely punctate anteroventral portion; (865) Propodeal dorsum of female; (866) Female gaster in dorsal view.