Pison deplanatum Pulawski, 2018

Pulawski, Wojciech J., 2018, A Revision of the Wasp Genus Pison Jurine, 1808 of Australia and New Zealand, New Guinea, and the Pacific Islands (Hymenoptera: Crabronidae), Proceedings of the California Academy of Sciences 65, pp. 1-584 : 150-152

Pison deplanatum Pulawski , species nova

Figures 302 -308.

NAME DERIVATION.– Deplanatum is a Latin neuter adjective meaning flattened; with reference to the shape of the thoracic dorsum of this species.

RECOGNITION.– Pison deplanatum has only two submarginal cells, the second being 1.9-2.2× as long posteriorly as high ( Fig. 306 ), the tegula impunctate and asetose in posterior half, and the propodeum without a longitudinal carina separating the side from the dorsum and the posterior surface. It is the only Pison in which the eye is covered with erect setae above the emargination only ( Fig. 303 ). In the female, the clypeal free margin ( Fig. 302 ) has no middle lobe (minimally prominent mesally, not concave laterally). Pison globosum is similar, but in the female of P. deplanatum the forefemur is conspicuously thickened ( Fig. 307 ) rather than insignificantly thickened, the frons is finely punctate, somewhat shiny between the punctures (rather than minutely reticulate, dull), the postocellar area has a transverse sulcus adjacent to the hindocelli (sulcus absent), the eye above the emargination ( Fig. 304 ) covered with erect setae (rather than asetose), the scutellum is flat, situat- ed in the same plane as the scutum and not foveate anteriorly (rather than slightly convex, slightly raising above the scutum level, and foveate anteriorly), the propodeal dorsum is finely obliquely ridged, punctate between ridges, only punctate along lateral margin (rather than irregularly, transversely ridged, impunctate), the propodeal side is unsculptured anteriorly or nearly so (rather than all sculptured), and the wing veins (all or most) are pale yellow brown in basal half (rather than all black). Like P. difficile and unlike the other species with two submarginal cells, the scutellum has no foveate sulcus along the anterior margin. The male is unknown.

DESCRIPTION.– Frons markedly convex, minutely punctate, punctures about one diameter apart; interspaces microsculptured but slightly shiny; middle supraantennal carina replaced by fine sulcus. Distance between antennal socket and orbit smaller than socket width. Midocellus smaller than hindocellus ( Fig. 303 ). Postocellar area with transverse sulcus adjacent to hindocelli. Gena narrow in dorsal view ( Fig. 304 ). Labrum minimally emarginate. Pronotal collar elongate ( Fig. 305 ), its horizontal part about 2.5 × as long as hindocellar diameter. Anteromedian pronotal pit absent. Propleuron sparsely punctate anterolaterally. Scutum not foveate along flange, without short longitudinal ridges adjacent to posterior margin; scutal punctures minute, averaging about one diameter apart; interspaces microsculptured. Tegula enlarged. Scutellum flattened, at same level as scutum, evenly sculptured, without foveate sulcus anteriorly between axillae. Mesopleural punctures fine, 1-2 diameters apart; interspaces unsculptured. Postspiracular carina absent. Metapleural sulcus not costulate between dorsal and ventral metapleural pits. Propodeum without longitudinal carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle; dorsum elongate, 2.0-2.5 × as long as scutellum, finely obliquely ridged, punctate between ridges, only punctate along lateral margin; side slightly concave, punctate except anteriorly (impunctate on most surface in some specimens), minutely ridged anteriorly in some specimens; posterior surface punctate, also with evanescent ridges in some specimens. Forewing with two submarginal cells ( Fig. 306 ); second submarginal cell 1.9-2.2 × as long posteriorly as high. Posteroventral forefemoral surface minutely punctate, punctures averaging about two diameters apart. Hindcoxal dorsum with outer margin blunt. Punctures of tergum I fine, about one diameter apart (some punctures about two diameters apart in some specimens). Sterna punctate throughout.

Setae silvery, appressed on frons, lower gena, thorax, forecoxal venter, femoral venters, and tergum I; partly concealing integument on clypeus. Eye above emargination covered with erect setae ( Fig. 303 ). Apical depressions of terga with ill-defined, silvery, setal fasciae.

Head, thorax, propodeum, and gaster black, mandible ferruginous (black basally, brown apically), flagellum black or yellowish brown ventrally. Wing veins (all or most) pale yellow brown in basal half. Femora black, foretibia and foretarsus ferruginous in most specimens, all tibiae and tarsi ferruginous in specimens from Mount Cook National Park and Shiptons Flats, Queensland. ♀.– Upper interocular distance equal to 1.00-1.05 × lower interocular distance; ocellocular distance equal to 1.0 × hindocellar diameter, distance between hindocelli equal to 1.4-1.7 × hindocellar diameter; eye height equal to 1.04-1.08 × distance between eye notches. Free margin of clypeal lamella without lobe, almost evenly arcuate except slightly prominent mesally, not concave laterally ( Fig. 302 ). Dorsal length of flagellomere I 1.3-1.4 × apical width, of flagellomere IX 0.9-1.1 × apical width. Mandible: trimmal carina with minute incision shortly before midlength. Forefemur markedly swollen ( Fig. 307 ). Tergum VI with median carina apically. Length 5.3-6.5 mm; head width 1.3-1.4 mm.

♂.– Unknown.

GEOGRAPHIC DISTRIBUTION (Fig. 308).–

Australian Capital Territory, eastern New

South Wales , eastern Queensland .

RECORDS.– HOLOTYPE: ♀, AUSTRALIA: Australian Capital Territory: Black Mountain, Dec

1982, I.D. Naumann and J.C. Cardale (ANIC).

PARATYPES: AUSTRALIA: Australian Capital

Territory: same data as holotype (1 ♀, ANIC) ; same data as holotype except Nov 1982 (1 ♀, ANIC). New

South Wales: Coolbaggie Forest Reserve 10 km

E Eumungerie at 31°58.5ʹS 148°40.5ʹE, 28 Dec

2011, V. Ahrens and W.J. Pulawski (1 ♀, CAS) ;

Shoalhaven River 30 km W Nowra, 25 Dec 1986,

G.A. Holloway (1 ♀, AMS). Queensland: Carnarvon National Park at 25°03.6ʹS 148°14.1ʹE, 1 Dec FIGURE 308. Collecting localities of Pison deplanatum Pulawski , sp. nov .

2012, V. Ahrens and W.J. Pulawski (1 ♀, CAS) ;

Crediton State Forest at 21°11.8ʹS 148°29.9ʹE, 2 Nov 2006, V. Ahrens and W.J. Pulawski (1 ♀, CAS) ; Eungella National Park at 21°10.5ʹS 148°30.3ʹE, 31 Oct 2006, V. Ahrens and W.J. Pulawski (2 ♀, CAS) ; George Creek 27.5 km W Bries Homestead at 19°32ʹ53ʺS 143°56ʹ33ʺE, 3-5 Nov 2001, D. Yeates, C. Lambkin, N. Stanick, and J. Hamilton (1 ♀, AMS) ; Mount Cook National Park at 15°29ʹS 145°16ʹE, 10-12 May 1981, I.D. Naumann (1 ♀, ANIC) ; Shiptons Flats at 15°47ʹS 145°14ʹE, 17-19 Oct, J.C. Cardale (1 ♀, ANIC) .

Figures

FIGURES 302–307. Pison deplanatum Pulawski, sp. nov., female. (302) Clypeus and mandibles; (303) Head in frontal view; (304) Head in dorsal view; (305) Pronotum; (306) Distal portion of forewing; (307) Forefemur.