Pison nudigenale Pulawski, 2018

Pulawski, Wojciech J., 2018, A Revision of the Wasp Genus Pison Jurine, 1808 of Australia and New Zealand, New Guinea, and the Pacific Islands (Hymenoptera: Crabronidae), Proceedings of the California Academy of Sciences 65, pp. 1-584 : 305-306

Pison nudigenale Pulawski , species nova

Figures 721 -723.

NAME DERIVATION.– Nudigenale is derived from the Latin adjectives: nudus meaning nude, bare, and genalis, meaning referring to the gena; with reference to the impunctate and glabrous lower gena (on each side of the oral cavity).

RECOGNITION.– The female of Pison nudigenale (the male is unknown) shares with P. gymnopareion a unique combination of the lower gena impunctate and glabrous on each side of the oral cavity, with the glabrous area bordered by a psammophore, and the presence of erect setae on tergum I. The females of the two species are quite similar morphologically, but they differ by the shape of the clypeal lamella: in P. nudigenale , it is obtusely angulate and relatively narrow (its corners are closer to each other than to the adjacent orbit); in P. gymnopareion , it is evenly arcuate and markedly broader (its corners are closer to the adjacent orbit than to each other).

DESCRIPTION.– Frons dull, finely punctate, punctures less than one diameter apart, middle supraantennal carina present, but covered by vestiture. Occipital carina slightly expanded ventrally, joining hypostomal carina. Mandible with abductor ridge. Gena narrow in dorsal view ( Fig. 722 ). Labrum not emarginate. Anteromedian pronotal pit transversely elongate, about as long as midocellar diameter. Propleuron sparsely punctate anteriorly. Scutum not foveate along flange, without longitudinal ridges adjacent to posterior margin; scutal punctures well defined, almost contiguous. Scutellum somewhat foveate along anterior margin. Tegula not enlarged. Mesopleural punctures well defined, contiguous. Postspiracular carina present, about 1.5 × as long as midocellar diameter. Metapleural sulcus costulate between dorsal and ventral metapleural pits. Propodeum with irregular longitudinal carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle; dorsum obliquely ridged; side ridged, punctate between ridges; posterior surface ridged, with several ridges radiating up from transverse carina just above gastropropodeal articulation. Posteroventral forefemoral surface with well-defined punctures less than one diameter apart. Punctures of tergum I about one diameter apart anterior to apical depression, uniform on anterior declivity. Punctures of sternum II several diameters apart (except laterally), sternum impunctate along midline; sterna III and IV with punctures several diameters apart, impunctate along midline.

Setae silvery, erect on postocellar area, thorax, forecoxal venter, femoral venters, and tergum I; completely concealing integument on clypeus (except for lamella); genal setae: see below. Apical depressions of terga with silvery, setal fasciae.

Body all black.

♀.– Upper interocular distance equal to 0.60 × lower interocular distance; ocellocular distance equal to 0.8 × hindocellar diameter, distance between hindocelli equal to 0.9 × hindocellar diameter; eye height equal to 0.90 × distance between eye notches. Free margin of clypeal lamella obtusely angulate ( Fig. 721 ). Dorsal length of flagellomere I 2.6 × apical width, of flagellomere IX 1.0 × apical width. Lower gena, mandibular posterior margin, propleural and forecoxal outer margins, and forefemoral venter with psammophores (longest setae of genal, mandibular, and forefemoral psammophores about 1.1 ×, 0.9 ×, and 0.9 ×, respectively, of greatest forefemoral width); lower gena impunctate and asetose between hypostomal carina and psammophore. Mandible:

trimmal carina with small incision at about two thirds of length. Length 6.5 mm; head width

2.4 mm.

♂.– Unknown.

GEOGRAPHIC DISTRIBUTION (Fig. 723).–

Known from one locality in southern Queensland.

RECORDS.– HOLOTYPE: ♀, AUSTRALIA:

Queensland: Dynevor Lakes at 28°05ʹS 144°12ʹE,

28 Sept, 1991, G. Daniels (QMB, registration number T228764).

PARATYPES: AUSTRALIA: Queensland: same FIGURE 723. Collecting locality of Pison nudigenale data as holotype (1 ♀, CAS). Pulawski, sp. nov .

Figures

FIGURES 721-722. Pison nudigenale Pulawski, sp. nov., female. (721) Clypeus and mandibles; (722) Head in dorsal view.